PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Glirulus (Japanese dormouse)

Mammalia - Rodentia - Gliridae

Taxonomy

Glirulus was named by Thomas (1906). It is extant. Its type is Glirulus japonicus.

It was assigned to Dryomyinae by Daams and de Bruijn (1995); to Glirinae by Kowalski (1997); and to Gliridae by McKenna and Bell (1997).

Species

Amphidyromys pusillus, G. conjunctus, G. diremptus, G. ekremi, G. gemmula, G. japonicus (syn. G. lasiotis, G. elegans) (type species), G. lissiensis, G. minor, G. miocaenicus, G. werenfelsi

Synonyms

Amphidyromys was named by Heller (1936). Its type is Amphidyromys pusillus.

It was synonymized subjectively with Glirulus by Daams and de Bruijn (1995).

Synonymy list

Year	Name and author
1906	Glirulus Thomas
1936	Amphidyromys Heller
1995	Glirulus Daams and de Bruijn p. 10
1997	Glirulus Kowalski
1997	Glirulus McKenna and Bell

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha

Rank	Name	Author
—	Mammaliaformes
class	Mammalia
—	Theriamorpha	(Rowe 1993)
—	Theriiformes	()
—	Trechnotheria
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Euarchontoglires	Murphy et al. 2001
—	Gliriformes	Wyss and Meng 1996
—	Glires	()
—	Simplicidentata	()
order	Rodentia	Bowdich 1821
superfamily	Gliroidea	Muirhead 1819
family	Gliridae	Muirhead 1819
subfamily	Glirinae	Thomas 1897
genus	Glirulus	Thomas 1906

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Glirulus Thomas 1906 [Japanese dormouse]

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†Glirulus conjunctus Mayr 1979

†Glirulus diremptus Mayr 1979

†Glirulus ekremi Unay 1994

†Glirulus gemmula Kretzoi 1962

Glirulus japonicus Schinz 1845

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Invalid names: Glirulus elegans Temminck 1845 [synonym], Glirulus lasiotis Thomas 1880 [synonym]

†Glirulus lissiensis Hugueney and Mein 1965

†Glirulus minor Wu 1993

†Glirulus miocaenicus Baudelot 1966

†Glirulus werenfelsi Engesser 1972

Invalid names: Amphidyromys Heller 1936 [synonym]

Diagnosis

Reference	Diagnosis
R. Daams and H. de Bruijn 1995	Small to medium-sized Dryomyinae with usually 9 transverse ridges and an endoloph in their upper molars. The labial ends of anteroloph and posteroloph are as high as paracone and metacone. The anterior centroloph is connected to the endoloph in all but the earliest species. An endolophid may be present in the lower molars.

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Life habit:

ground dwellingo

Diet:

herbivoreo

Reproduction:

viviparoussubc

Created:

2005-08-26 14:04:46

Modified:

2005-08-26 16:04:46

Source: o = order, subc = subclass, c = class, subp = subphylum

References: Lillegraven 1979, Carroll 1988, Ji et al. 2002, Nowak 1999, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Late/Upper Oligocene to the top of the Late/Upper Pleistocene or 28.40000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 23.03 Ma

Collections (37 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Oligocene	28.4 - 23.03	Turkey (Anatolia)	Glirulus sp. (182501)
MP 30	28.4 - 23.03	Turkey	Glirulus sp. (68157)
MP 30 - MN 1	28.4 - 22.4	Turkey (Anatolia)	Glirulus sp. (182505)
Early/Lower Miocene	23.03 - 15.97	Greece (Lesvos)	Glirulus diremptus (186558)
Burdigalian	20.44 - 15.97	Serbia and Montenegro (Serbia)	Glirulus ekremi (39714) Glirulus lissiensis (199782) Glirulus werenfelsi, Glirulus lissiensis, Glirulus ekremi (199780 199781)
Burdigalian	20.44 - 15.97	Greece	Glirulus diremptus (68032)
MN 4	16.9 - 15.97	Poland	Glirulus diremptus (31041)
MN 4	16.9 - 15.97	Greece (Thrace)	Glirulus diremptus (69435)
MN 4	16.9 - 15.97	Austria (Styria)	Glirulus sp., Glirulus lissiensis (74538 74539)
MN 5	15.97 - 13.65	Germany (Bayern)	Glirulus diremptus (49519)
MN 5 - MN 6	15.97 - 12.75	Poland	Glirulus werenfelsi, Glirulus lissiensis (31040)
MN 7 + 8	12.75 - 11.1	Switzerland (Thurgau)	Glirulus conjunctus (46277 53571)
MN 7 + 8	12.75 - 11.1	Romania (Bihor)	Glirulus lissiensis (200051)
Tortonian	11.62 - 7.246	Germany (Bavaria)	Glirulus conjunctus (206345)
Late/Upper Miocene	11.608 - 5.333	Greece (Crete)	Glirulus werenfelsi (183151)
MN 9	11.1 - 9.7	Hungary (Borsod-Abaúj-Zemplén)	Glirulus lissiensis (75481)
MN 9	11.1 - 9.7	Poland	Glirulus werenfelsi, Glirulus lissiensis (31039)
MN 11	8.7 - 7.75	Austria (Niederoesterreich)	Glirulus lissiensis (32649)
Messinian	7.246 - 5.333	Italy (Piedmont)	Glirulus lissiensis (172910 172915 172916 199859)
Ruscinian	5.333 - 3.2	Germany (Rheinland-Pfalz)	Glirulus pusillus (43633)
MN 14	4.9 - 4.2	Poland	Glirulus pusillus (32240)
Piacenzian	3.6 - 2.588	Germany (North Rhine-Westphalia)	Glirulus pusillus (199747 205172)
MN 16	3.2 - 2.588	Italy (Piedmont)	Glirulus pusillus (49360)
Villanyian	3.2 - 2.588	Hungary (Borsodabaujzemplen)	Glirulus pusillus (35358)
MN 17	2.588 - 1.806	Netherlands (Limburg)	Glirulus pusillus (68018)
Early/Lower Pleistocene	2.588 - 0.781	Romania	Glirulus pusillus (32192)
Pleistocene	2.588 - 0.0117	France (Cote d'Or)	Glirulus pusillus (49085)
Late/Upper Pleistocene	0.129 - 0.0117	Japan (Aomori)	Glirulus japonicus (101899)