Villarroya: Pliocene, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Rodentia - Hystricidae
Hystrix cf. etrusca
1 specimen
    = Hystrix primigenia
Arribas Herrera and Bernad Garcia 1994
Mammalia - Perissodactyla - Rhinocerotidae
Dicerorhinus etruscus (Falconer 1868)
3 specimens
    = Stephanorhinus etruscus Falconer 1868
O'Regan 2003
Mammalia - Perissodactyla - Equidae
Hipparion crusafonti
    = Hipparion rocinantis Hernández Pacheco 1921
Aguirre and Morales 1990
    = Hipparion rocinantis Hernández Pacheco 1921
Arribas Herrera and Bernad Garcia 1994
Hipparion rocinantis crusafonti
Mammalia - Carnivora - Mustelidae
Lutra sinerizi n. sp.
1 individual
    = Enhydrictis ardea Bravard 1828
Arribas Herrera and Bernad Garcia 1994
Mammalia - Carnivora - Canidae
Canis donnezani
1 specimen
    = Canidae indet. Fischer 1817
Arribas Herrera and Bernad Garcia 1994
Vulpes (? Cynaloplex) alopecoides Forsyth Major 1877
3 specimens
    = Vulpes alopecoides Forsyth Major 1877
Arribas Herrera and Bernad Garcia 1994
Nyctereutes megamastoides Pomel 1842
12 specimens
Mammalia - Carnivora - Felidae
Megantereon megantereon Croizet and Jobert 1828
6 specimens
    = Megantereon cultridens Cuvier 1824
Arribas Herrera and Bernad Garcia 1994
Epimachairodus crenatidens
Villalta and Crusafont Pairó 1956
    = Homotherium crenatidens
O'Regan 2003
Sivapanthera arvernensis Croizet and Jobert 1828
3 specimens
    = Acinonyx pardinensis Croizet and Jobert 1828
Arribas Herrera and Bernad Garcia 1994
? Acinonyx pardinensis Croizet and Jobert 1828
Villalta and Crusafont Pairó 1956
    = Acinonyx pardinensis Croizet and Jobert 1828
Aguirre and Morales 1990
Mammalia - Carnivora - Hyaenidae
Hyaena marini n. sp.
7 specimens
    = Hyaena perrieri Croizet and Jobert 1828
Arribas Herrera and Bernad Garcia 1994
Euryboas lunensis (Del Campana 1914)
Villalta and Crusafont Pairó 1956
    = Chasmaporthetes lunensis Del Campana 1914
McKenna and Bell 1997
Mammalia - Proboscidea - Gomphotheriidae
? Mastodon sp. Cuvier 1817
3 specimens
    = Anancus arvernensis Croizet and Jobert 1828
Aguirre and Morales 1990
Mammalia - Ungulata - Bovidae
Hesperoceras merlae n. gen., n. sp. Villalta and Crusafont Pairó 1956
Villalta and Crusafont Pairó 1956 1 individual
Gazellospira torticornis
Aguirre and Morales 1990
Leptobos elatus
Aguirre and Morales 1990
Megalovis sp. Schaub 1924
Arribas Herrera and Bernad Garcia 1994
Gazella borbonica
Arribas Herrera and Bernad Garcia 1994
Mammalia - Ungulata - Cervidae
"Cervus" perrieri
Aguirre and Morales 1990
Croizetoceros ramosus (Croizet and Jobert 1828)
Aguirre and Morales 1990
Cervus cusanus
Aguirre and Morales 1990
    = Arvernoceros ardei
Arribas Herrera and Bernad Garcia 1994
see common names

Geography
Country:Spain State/province:La Rioja
Coordinates: 42.1° North, 2.1° West (view map)
Paleocoordinates:42.2° North, 2.4° West
Basis of coordinate:based on political unit
Altitude:851 meters
Geographic resolution:local area
Time
Period:Neogene Epoch:Pliocene
10 m.y. bin:Cenozoic 6
Key time interval:Pliocene
Age range of interval:5.33300 - 2.58800 m.y. ago
Stratigraphy
Stratigraphy comments: "Pliocene" according to Villalta (1952), "Villafranquiense" according to Villalta (1956)
Lithology and environment
Primary lithology:red sandy mudstone
Secondary lithology:red sandy mudstone
Includes fossils?Y
Includes fossils?Y
Lithology description: Upper: maroon sandy mudstone; Lower: light red sandy mudstone
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:some genera
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Metadata
Database number:38892
Authorizer:J. Alroy, A. Turner Enterer:J. Wertheim, H. O'Regan
Modifier:J. Alroy Research group:vertebrate
Created:2004-05-13 14:19:42 Last modified:2006-07-31 00:44:02
Access level:the public Released:2004-05-13 14:19:42
Creative Commons license:CC BY
Reference information

Primary reference:

10511. J. F. Villalta Comella. 1952. Contribución al conocimiento de la fauna de mamíferos fósiles del Plioceno de Villaroya (Logroño). Boletín del Instituto Geológico y Minero de España 64:1-201 [J. Alroy/J. Wertheim/J. Wertheim]

Secondary references:

10948PACED E. Aguirre and J. Morales. 1990. Villafranchian faunal record of Spain. Quartärpaläontologie 8:7-11 [A. Turner/H. O'Regan/T. Liebrecht]
13549PACED A. Arribas Herrera and J. Bernad Garcia. 1994. Catalogo de mamiferos pliocenos del yacimiento de Villarroya (La Rioja), en la coleccion del Museo Geominero. Boletin Geologico y Minero 105(3):236-248 [A. Turner/H. O'Regan/H. O'Regan]
7584 M. C. McKenna and S. K. Bell. 1997. Classification of Mammals Above the Species Level 1-640 [J. Alroy/J. Alroy/M. Uhen]
9580PACED H. J. O'Regan. 2003. Reidentifications and synonomies of European mammals. [A. Turner/H. O'Regan/M. Uhen]
105155% 44540J. F. Villalta and M. Crusafont Pairó. 1956. Un nuevo Ovicaprino en la fauna villafranquiense de Villaroya (Logroño). International Quaternary Association, 4th Congress 1:426-432 [J. Alroy/J. Wertheim/J. Wertheim]