Woodring Loc. 138 [lower Gatun Fm]: Middle Miocene, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Murex (Murex) recurvirostris (Broderip 1832)
Woodring 1959 R category
Subspecies: Murex (Murex) recurvirostris recurvirostris
    = Haustellum messorium Sowerby 1841
Hendy et al. 2008
Bivalvia - Arcida - Arcidae
Anadara (Rasia) sechurana (Olsson 1932)
Woodring 1973 A category
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 C category
Subspecies: Anadara (Rasia) dariensis dariensis
Anadara (Potiarca) chavezi
Woodring 1973 C category
Bivalvia - Pectinida - Pectinidae
Flabellipecten gatunensis (Toula 1909)
Woodring 1982 F category
recombined as Leopecten gatunensis
Subspecies: Flabellipecten gatunensis gatunensis
Aequipecten plurinominis (Pilsbry and Johnson 1917)
Woodring 1982 R category
    = Lindapecten buchivcanoanus
Waller 2011
SUBSPECIES: Lindapecten buchivcanoanus falconensis
Leptopecten ecnomius n. sp. Woodring 1982
Woodring 1982 A category
Bivalvia - Pectinida - Limidae
Limatula asymbleta n. sp. Woodring 1982
Woodring 1982 R category
Bivalvia - Ostreida - Pteriidae
Pteria inornata (Gabb 1873)
Woodring 1973 R category
Bivalvia - Carditida - Crassatellidae
Eucrassatella (Hybolophus) elassa Woodring 1982
Woodring 1982 A category
Crassinella martinicensis (d'Orbigny 1853)
Woodring 1982 F category
Bivalvia - Pholadida - Corbulidae
Bothrocorbula (Hexacorbula) gatunensis (Toula 1909)
Woodring 1982 A category
Caryocorbula oropendula (Olsson 1922)
Woodring 1982 C category
Subspecies: Caryocorbula oropendula stena
Varicorbula disparilis (d'Orbigny 1853)
Woodring 1982 R category
Bivalvia - Cardiida - Kelliellidae
Alveinus rotundus Dall 1900
Woodring 1982 A category
Bivalvia - Cardiida - Veneridae
Chione (Chionopsis) tegulum (Brown and Pilsbry 1911)
Woodring 1982 C category
Lirophora (Lirophora) falconensis (Hodson et al. 1927)
Woodring 1982 R category
Lirophora (Panchione) mactropsis (Conrad 1855)
Woodring 1982 R category
Agriopoma (Pitarella) gatunensis (Dall 1903)
Woodring 1982 R category
recombined as Callocardia (Agriopoma) gatunensis
Lamelliconcha cora (Brown and Pilsbry 1911)
Woodring 1982 F category
Macrocallista (Paradione) maculata (Linnaeus 1758)
Woodring 1982 C category
recombined as Macrocallista maculata
Bivalvia - Cardiida - Tellinidae
Psammacoma gatunensis (Toula 1909)
Woodring 1982 R category
Tellina (Eurytellina) angulosa (Gmelin 1791)
Woodring 1982 R category
Subspecies: Tellina (Eurytellina) angulosa costaricana
Tellina (Eurytellina) punicea (Born 1778)
Woodring 1982 R category
recombined as Eurytellina punicea
Bivalvia - Cardiida - Semelidae
Semele laevis (Sowerby 1833)
Woodring 1982 R category
Subspecies: Semele laevis costaricensis
Ervilia valhosierr Gardner 1928
Woodring 1982 A category
Bivalvia - Cardiida - Cardiidae
Trachycardium (Dallocardia) baiterum n. sp. Woodring 1982
Woodring 1982 R category
Laevicardium laevigatum (Linnaeus 1758)
Woodring 1982 R category
replaced by Laevicardium serratum
Trigoniocardia (Apiocardia) aminensis (Dall 1900)
Woodring 1982 A category
recombined as Trigoniocardia (Trigoniocardia) aminensis
Bivalvia - Lucinida - Lucinidae
Bellucina actinus (Dall 1903)
Woodring 1982 F category
Bivalvia - Protobranchia - Nuculanidae
Saccella balboae (Brown and Pilsbry 1911)
Woodring 1973 R category
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) gatunensis (Toula 1908)
Woodring 1964 C category
original and current combination Oliva gatunensis
Olivella (Niteoliva) terryi Olsson 1956
Woodring 1964 A category
Gastropoda - Neogastropoda - Olivellidae
Agaronia testacea (Lamarck 1811)
Woodring 1964 R category
Subspecies: Agaronia testacea mancinella
Gastropoda - Neogastropoda - Columbellidae
Strombina (Strombina) lessepsiana (Brown and Pilsbry 1911)
Jung 1989 8 specimens
common in Woodring (1957-1982)
Mitrella (Columbellopsis) aff. fenestrata (Adams 1850)
Woodring 1964 R category
synonym of Columbellopsis nycteis
Anachis (Costoanachis) mira (Dall 1896)
Woodring 1964 F category
Subspecies: Anachis (Costanachis) mira fugax
Gastropoda - Neogastropoda - Buccinidae
Strombinophos mimicus n. sp. Woodring 1964
Woodring 1964 F category
Cymatophos veatchi (Olsson 1922)
Woodring 1964 A category
Subspecies: Cymatophos veatchi veatchi
Antillophos (Antillophos) candei (d'Orbigny 1842)
Woodring 1964 R category
recombined as Antillophos candeanus
Subspecies: Antillophos (Antillophos) candei gatunensis
Hanetia dalli (Brown and Pilsbry 1911)
Woodring 1964 C category
Subspecies: Hanetia dalli protera
Gastropoda - Neogastropoda - Nassariidae
Calophos ectyphus Woodring 1964
Woodring 1964 C category
Nassarius (? Uzita) praeambiguus (Brown and Pilsbry 1913)
Woodring 1964 C category
Gastropoda - Neogastropoda - Melongenidae
Melongena melongena (Linnaeus 1758)
Woodring 1964 R category
Subspecies: Melongena melongena consors
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Euclia) dinota n. sp. Woodring 1970
Woodring 1970 R category
recombined as Euclia dinota
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 F category
Subspecies: Cancellaria (Cancellaria) epsitomitera dariena
    = Bivetiella dariena Toula 1908
Landau et al. 2012
Cancellaria (Pyruclia) cibarcola (Anderson 1929)
Woodring 1970 R category
Subspecies: Cancellaria (Pyruclia) cibarcola cibarcola
Cancellaria (Narona) barystoma n. sp. Woodring 1970
Woodring 1970 R category
recombined as Narona barystoma
Gastropoda - Neogastropoda - Muricidae
Trachypollia aneureta n. sp. Woodring 1964
Woodring 1964 R category
Murex (? Murex) polynematicus (Brown and Pilsbry 1911)
Woodring 1959 C category
recombined as Vokesimurex polynematicus
Gastropoda - Neogastropoda - Turridae
Pleurofusia (Cruziturricula) fusinus (Brown and Pilsbry 1911)
Woodring 1970 C category
recombined as Cruziturricula fusinus
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 R category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Gastropoda - Neogastropoda - Conidae
Conus aemulator Brown and Pilsbry 1911
Woodring 1970 R category
Subspecies: Conus aemulator aemulator
Gastropoda - Eulimidae
Balcis (Balcis) lipara n. sp. Woodring 1970
Woodring 1970 F category
Balcis (Balcis) jacululum (Maury 1917)
Woodring 1970 R category
Balcis (Balcis) aulaca n. sp. Woodring 1970
Woodring 1970 R category
Eulima sarissiformis (Pilsbry and Johnson 1917)
Woodring 1970 F category
Eulima nobilis Guppy 1896
Woodring 1970 R category
Gastropoda - Naticidae
Natica (? Natica) bolus Brown and Pilsbry 1913
Stigmaulax guppiana (Toula 1909)
Natica (Naticarius) stenopa n. sp.
original and current combination Naticarius
Polinices stanislasmeunieri Maury 1917
Gastropoda - Cypraeidae
Cypraea (Muracypraea) henekeni (Sowerby 1850)
Woodring 1959 F category
    = Muracypraea woodringi Landau and Groves 2011
Landau and Groves 2011
Gastropoda - Calyptraeidae
Crucibulum (Dispotaea) springvaleense (Rutsch 1942)
Calyptraea centralis (Conrad 1841)
Crepidula plana Say 1822
Crepidula cf. maculosa Conrad 1846
Gastropoda - Personidae
Distorsio (Rhysema) decussata (Valenciennes 1832)
Woodring 1959 R category
recombined as Distorsio decussata
Subspecies: Distorsio (Rhysema) decussata gatunensis
Gastropoda - Tonnidae
Malea camura Guppy 1866
Woodring 1959 R category
    = Malea ringens Swainson 1822
Beu 2010
Gastropoda - Bursidae
Bursa (Colubrellina) caelata (Broderip 1833)
Woodring 1959 F category
Subspecies: Bursa (Colubrellina) caelata amphitrites
    = Bursa rugosa Sowerby 1835
Landau and Marques da Silva 2010
Gastropoda - Cassidae
Semicassis (Tylocassis) reclusa (Guppy 1873)
Woodring 1959 R category
recombined as Semicassis reclusa
Gastropoda - Vitrinellidae
Episcynia megalia n. sp.
Cyclostremiscus (Ponocyclus) pentagonus (Gabb 1873)
Anticlimax (Anticlimax) gatunensis
Gastropoda - Tornidae
Teinostoma (Pseudorotella) pycnum Woodring 1957
Gastropoda - Sorbeoconcha - Turritellidae
Turritella (Torcula) alitilira
original and current combination Torcula
Subspecies: Turritella (Torcula) alitilira praecellens
Turritella gatunensis Conrad 1857
Subspecies: Turritella gatunensis gatunensis
Gastropoda - Sorbeoconcha - Cerithiidae
Bittium nugatorium n. sp. Brown and Pilsbry 1911
Woodring 1959 R category
Alabina asperoides
Woodring 1959 R category
Subspecies: Alabina asperoides asperoides
Gastropoda - Architectonicidae
Heliacus (Astronacus) stonemanae (Maury 1917)
Woodring 1959 R category
Architectonica (Architectonica) nobilis (Röding 1798)
Woodring 1959 C category
original and current combination Architectonica nobilis
Subspecies: Architectonica (Architectonica) nobilis nobilis
Gastropoda - Turbinidae
Solariorbis (Solariorbis) strongylus n. sp.
Scaphopoda - Gadilida - Gadilidae
Cadulus (Platyschides) epetrion n. sp.
Woodring 1973 A category
see common names

Geography
Country:Panama
Coordinates: 9.3° North, 79.8° West (view map)
Paleocoordinates:8.9° North, 78.0° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 5
Key time interval:Middle Miocene
Age range of interval:15.97000 - 11.60800 m.y. ago
Age estimate:11.3 Ma (other)
Stratigraphy
Formation:Gatun Member:lower Gatun
Stratigraphic resolution:bed
Stratigraphy comments: Same stratigraphic position as following collection (52174). Lower part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 11.3 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology:fine poorly lithified silty sandstone
Includes fossils?Y
Lithology description: SPECIFIC: Soft, silty fine-grained sandstone. GENERAL: Massive medium-to very fine-grained sandstone and siltstone. Somewhat calcareous, or marly, somewhat tuffaceous, and with a clay-like matrix. The lower part of the formation consists principally of medium-very fine-grained sandstone.
Environment:marine indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:selective quarrying,field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collection method comments: Collected by Woodring (1947). Material is also held (but not specified for any specific collection) at Stanford University, Acad. Nat. Sci. Phil., Cornell University, PRI, U. Cal. and Cal. Acad. Sci.
Taxonomic list comments:Exhaustive for gastropoda and bivalvia.
Metadata
Also known as:USGS Loc. 6909
Database number:52173
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2005-07-28 14:28:06 Last modified:2013-11-25 13:37:23
Access level:group members Released:2006-07-28 14:28:06
Creative Commons license:CC BY
Reference information

Primary reference:

11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

30037 A. G. Beu. 2010. Neogene Tonnoidean Gastropods of Tropical and South America; contributions to the Dominican Republic and Panama Paleontology Projects and Uplift of the Central American Isthmus. Bulletins of American Paleontology (377-378)1-550 [A. Hendy/A. Hendy/A. Hendy]
19909 A. J. W. Hendy, D. P. Buick, K. V. Bulinski, C. A. Ferguson, and A. I. Miller. 2008. Unpublished census data from Atlantic coastal plain and circum-Caribbean Neogene assemblages and taxonomic opinions. [A. Hendy/A. Hendy/A. Hendy]
26828 P. Jung. 1989. Revision of the Strombina-group (Gastropoda; Columbellidae), fossil and living. Schweierische Paläontologische Abhandlungen 111:1-298 [A. Hendy/A. Hendy/A. Hendy]
36423 B. Landau and C. Marques da Silva. 2010. Early Pliocene gastropods of Cubagua, Venezuela: Taxonomy, palaeobiogeography and ecostratigraphy. Palaeontos 19:1-221 [A. Hendy/A. Hendy]
43491 B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy]
43452 B. M. Landau and L. T. Groves. 2011. Cypraeidae (Mollusca: Gastropoda) from the early Miocene Cantaure Formation of northern Venezuela. Novaplex 12(1-2):1-38 [A. Hendy/A. Hendy]
48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
37262 T. R. Waller. 2011. Neogene Paleontology of the Northern Dominican Republic. 24. Propeamussidae and Pectinidae (Mollusca: Bivalvia: Pectinoidea) of the Cibao Valley. Bulletins of American Paleontology (381)1-198 [A. Hendy/A. Hendy]
11372 W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]