Huangbanjigou, Shangyuan Town, NIGP collection (Cretaceous of China)

Where: Liaoning, China (41.6° N, 120.8° E: paleocoordinates 44.1° N, 122.9° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Yixian Formation (Jehol Group), Late/Upper Barremian (130.0 - 125.5 Ma)

• The Yixian Formation can be subdivided into five units, in ascending order, Lujiatun unit, Lower Lava, Jianshangou unit, Upper Lava, Dawangzhangzi unit, Jingangshan unit, and Huanghuashan unit. Most of the Jehol fossils from the Yixian Formation are preserved in four beds, that is, the Lujiatun bed (recalculated age is 124.0 ± 1.0/1.3 Ma, He et al., 2006), the Jianshangou bed (125.43 ± 0.50/0.88 Ma, Swisher et al., 1999), the Dawangzhangzi bed, and Jinggangshan bed (~122 Ma, Smith et al., 1995).U-Pb dating suggests that the Huangbanjigou section ranges from 124.9 ± 1.7 Ma to 122.8 ± 1.6 Ma (Yang et al., 2007). Chang et al. (2017) obtained Ar/Ar ages of 125.8 ± 1.0 Ma and 126.0 ± 0.8 Ma from the Lujiatun Unit, the lowest part of the Yixian Formation, below the lacustrine shales.

•He et al. (2004) reported a 40Ar/39Ar age of 120.3 ± 0.7 Ma for a tuff layer in the lower part of the Jiufotang Formation, and Chang et al. (2009) obtained a 40Ar/39Ar age of 122.1 ± 0.3 Ma for a tuff layer from the lowermost part of the Jiufotang Formation. Yu et al. (2021) dated an ash bed just above the Yixian/Jiufotang formation boundary to 122.0 ± 0.9 Ma and three tuff beds from the Xiaotaizi section to 118.9 ± 0.8 Ma, 118.8 ± 0.6 Ma, and 118.6 ± 1.1 Ma. These ages suggest that the Jiufotang Formation in Jianchang Basin was deposited at ca. 122.0–118.9 Ma.

•Olierrok et al. (2019) used a selection of U-Pb and 40Ar-39Ar geochronology to place the Barremian–Aptian boundary (the base of the Chron M0r) between 123.8 and 121.8 Ma. The GTS 2020 uses an age of 121.4 Ma for the base of the Aptian, and multiproxy orbital tuning gives an age of 120.2 Ma for the Barremian-Aptian boundary (Leandro et al., 2022). Therefore, the Yixian Formation should be entirely Late Barremian, with the Jiufotang Formation deposited in the Early Aptian (or latest Barremian at its base).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, tuffaceous mudstone

Size classes: macrofossils, mesofossils

Preservation: adpression

Reposited in the NIGPAS

Primary reference: J. Zhang. 1999. Aeschnidiid nymphs from the Jehol biota (latest Jurassic–Early Cretaceous), China, with a discussion of the family Aeschnidiidae (Insecta, Odonata). Cretaceous Research 20:813-827 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111725: authorized by Matthew Clapham, entered by Matthew Clapham on 28.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Orthoptera - Elcanidae
Jeholelcana yanensis n. gen. n. sp.11
Jeholelcana yanensis n. gen. n. sp.11 Fang et al. 2018 grasshopper
NIGP 159068
 Hemiptera - Palaeontinidae
Yanocossus cf. guoi21 Ren 1995 true bug
NIGP148388
 Hemiptera - Procercopidae
Sinocercopis cf. macula10 Hu et al. 2014 froghopper
NNH08016
Stellularis minutus10 Chen and Wang 2020 froghopper
NND08014, 08015
 Hymenoptera - Xyelidae
Heteroxyela ignota n. gen. n. sp.30, Isoxyela rudis n. gen. n. sp.30, Sinoxyela viriosa n. gen. n. sp.30, Liaoxyela antiqua n. gen. n. sp.30, Ceratoxyela decorosa n. gen. n. sp.30, Lethoxyela excurva n. gen. n. sp.30, Lethoxyela vulgata n. sp.30, Angaridyela robusta n. sp.30, Angaridyela endemica n. sp.30, Angaridyela suspecta n. sp.30, Angaridyela exculpta n. sp.30
Heteroxyela ignota n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131987 (LBSH98009)
Isoxyela rudis n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131989 (LBSH98011)
Sinoxyela viriosa n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131988 (LBSH98010)
Liaoxyela antiqua n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131986 (LBSH98008)
Ceratoxyela decorosa n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131985 (LBSH98007)
Lethoxyela excurva n. gen. n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131983 (LBSH98005)
Lethoxyela vulgata n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131984 (LBSH98006)
Angaridyela robusta n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131979 (LBSH98001)
Angaridyela endemica n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131982 (LBSH98004)
Angaridyela suspecta n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131981 (LBSH98003)
Angaridyela exculpta n. sp.30 Zhang and Zhang 2000 sawfly
NIGP 131980 (LBSH98002)
 Hymenoptera - Sepulcidae
"Trematothoracoides liaoningensis n. gen. n. sp." = Trematothorax liaoningensis33
"Trematothoracoides liaoningensis n. gen. n. sp." = Trematothorax liaoningensis33 Zhang et al. 2001 stem sawfly
LBSH98015/NIGP 132459
 Hymenoptera - Praeaulacidae
Sinowestratia communicata n. gen. n. sp.34
Sinowestratia communicata n. gen. n. sp.34 Zhang and Zheng 2000 wasp
LBSH98025/NIGP 132105
 Hymenoptera - Evaniidae
"Procretevania vesca n. sp." = Cretevania vesca25, "Procretevania exquisita n. sp." = Cretevania exquisita25
"Procretevania vesca n. sp." = Cretevania vesca25 Zhang et al. 2007 ensign wasp
LBSH02174/NIGP140527
"Procretevania exquisita n. sp." = Cretevania exquisita25 Zhang et al. 2007 ensign wasp
LBSH02175/NIGP140526
 Hymenoptera - Mesoserphidae
Beipiaoserphus elegans n. gen. n. sp.31
Beipiaoserphus elegans n. gen. n. sp.31 Zhang and Zhang 2000 wasp
NIGP 132048
 Hymenoptera - Pelecinidae
Eopelecinus similaris n. sp.28 Zhang et al. 2002 wasp
LBSH98046/NIGP 134163
Eopelecinus shangyuanensis n. sp.28 Zhang et al. 2002 wasp
LBSH98045/NIGP 134162
Eopelecinus vicinus n. gen. n. sp.28 Zhang et al. 2002 wasp
LBSH98042/NIGP 134161
Sinopelecinus epigaeus n. sp.28 Zhang et al. 2002 wasp
LBSH98043/NIGP 134158
Sinopelecinus magicus n. sp.28 Zhang et al. 2002 wasp
LBSH98044/NIGP 134159
Sinopelecinus viriosus n. sp.28 Zhang et al. 2002 wasp
LBSH98048/NIGP 134160
Sinopelecinus delicatus n. gen. n. sp.28 Zhang et al. 2002 wasp
LBSH98047/NIGP 134157
Scorpiopelecinus versatilis n. gen. n. sp.28 Zhang et al. 2002 wasp
LBSH98049/NIGP 134164
 Hymenoptera - Scoliidae
Protoscolia imperialis n. sp.29 Zhang et al. 2002 wasp
NIGP 133156 (LBSH98066)
Protoscolia normalis n. sp.29 Zhang et al. 2002 wasp
NIGP 133155 (LBSH98065)
Protoscolia sinensis n. gen. n. sp.29 Zhang et al. 2002 wasp
LBSH98064/NIGP 133154
 Hymenoptera - Angarosphecidae
Angarosphex saxosus n. sp.26 Zhang et al. 2018 apoid wasp
NIGP165267 (LBSH02212)
 Hymenoptera - Ephialtitidae
Tuphephialtites zherikhini n. gen. n. sp.27 Zhang et al. 2002 wasp
LBSH98019/NIGP134560
Crephanogaster rara n. sp.27 Zhang et al. 2002 wasp
LBSH98018/NIGP134559
 Hymenoptera - Ichneumonidae
Tanychorinae indet.15, ? Tanychorinae indet.15, Tanychorella dubia n. sp.23, Sinochora distorta n. gen. n. sp.15, Khasurtella zhangi n. sp.15, Amplicella sp.15, Amplicella townesi n. sp.15, Amplicella flagellata n. sp.15, "Tanychora spinata n. sp." = Amplicella spinata23, Amplicella exquisitissima n. sp.15, "Tanychora exquisita n. sp." = Amplicella exquisita23, "Tanychora beipiaoensis n. sp." = Amplicella beipiaoensis23, Tanychora liaoningensis n. sp.15, Tanychora rasnitsyni n. sp.15
Tanychorinae indet.15 Rasnitsyn 1980 ichneumon wasp
NIGP 159645 (LBSH02307), NIGP 159646 LBSH02363,2369)
? Tanychorinae indet.15 Rasnitsyn 1980 ichneumon wasp
NIGP 159647 (LBSH02339)
Tanychorella dubia n. sp.23 Zhang and Rasnitsyn 2003 ichneumon wasp
NIGP134823
Sinochora distorta n. gen. n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159643 (LBSH02188)
Khasurtella zhangi n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159642 (LBSH02184)
Amplicella sp.15 Kopylov 2010 ichneumon wasp
NIGP 159644 (LBSH02182)
Amplicella townesi n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159639 (LBSH02187)
Amplicella flagellata n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159637 (LBSH02181)
"Tanychora spinata n. sp." = Amplicella spinata23 Zhang and Rasnitsyn 2003 ichneumon wasp
NIGP134822
Amplicella exquisitissima n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159638 (LBSH02186)
"Tanychora exquisita n. sp." = Amplicella exquisita23 Zhang and Rasnitsyn 2003 ichneumon wasp
NIGP134820
"Tanychora beipiaoensis n. sp." = Amplicella beipiaoensis23 Zhang and Rasnitsyn 2003 ichneumon wasp
NIGP134821
Tanychora liaoningensis n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159640 (LBSH02185)
Tanychora rasnitsyni n. sp.15 Kopylov and Zhang 2015 ichneumon wasp
NIGP 159641 (LBSH02183)
 Hymenoptera - Anaxyelidae
Brachysyntexis robusta n. sp.24 Zhang and Rasnitsyn 2006 winged insect
LBSH98017/NIGP 135548
Syntexyela continentalis n. sp.24 Zhang and Rasnitsyn 2006 winged insect
LBSH98016/NIGP 135549
 Siphonaptera - Pseudopulicidae
Tyrannopsylla beipiaoensis n. gen. n. sp.13
Tyrannopsylla beipiaoensis n. gen. n. sp.13 Huang et al. 2013 flea
NIGP 154249, 154250
 Coleoptera -
Notocupes sp.17 Ponomarenko 1964 beetle
NIGP 175866
 Coleoptera - Hydroscaphidae
? Hydroscapha jeholensis n. sp.7 Cai et al. 2012 skiff beetle
NIGP 152505
 Coleoptera -
Mathesius liaoningensis n. gen. n. sp.14
Mathesius liaoningensis n. gen. n. sp.14 Kolibáč and Huang 2011 beetle
NIGP 152355
 Coleoptera - Staphylinidae
Hesterniasca lata n. sp.6, Ptomascopus sp.9, Cretoxyporus extraneus n. gen. n. sp.5, "Quedius cretaceus n. sp." = Cretoquedius infractus2, Megolisthaerus minor n. sp.1, Thayeralinus sp.9, Paroxyporus fossilis n. gen. n. sp.19, Mesocoprophilus clavatus n. gen. n. sp.3, Protodeleaster glaber n. gen. n. sp.8, Pseudanotylus archaicus4, Cretaceonanobius fossilis n. gen. n. sp.18
Hesterniasca lata n. sp.6 Cai et al. 2011 rove beetle
NIGP152502
Ptomascopus sp.9 Kraatz 1877 rove beetle
NIGP 156150a, 156161, 156153, 156154
Cretoxyporus extraneus n. gen. n. sp.5 Cai and Huang 2014 rove beetle
NIGP 153699
"Quedius cretaceus n. sp." = Cretoquedius infractus2 Solodovnikov and Yue 2013 rove beetle
NIGP 156220
Megolisthaerus minor n. sp.1 Cai and Huang 2013 rove beetle
NIGP 153697
Thayeralinus sp.9 Solodovnikov and Yue 2013 rove beetle
NIGP 156155a, 156156a, 156157, 157735
Paroxyporus fossilis n. gen. n. sp.19 Cai 2021 rove beetle
NIGP 173919
Mesocoprophilus clavatus n. gen. n. sp.3 Cai and Huang 2013 rove beetle
NIGP 153703
Protodeleaster glaber n. gen. n. sp.8 Cai et al. 2013 rove beetle
NIGP 152503, 152504
Pseudanotylus archaicus4 Yue et al. 2012 rove beetle
NIGP 153701, 153702
Cretaceonanobius fossilis n. gen. n. sp.18 Liu et al. 2022 rove beetle
NIGP 177043
 Coleoptera - Passalopalpidae
Mesoceratocanthus cf. tuberculifrons16 Nikolajev et al. 2010 scarab beetle
NIGP 177046
 Coleoptera - Lucanidae
Prolucanus beipiaoensis n. gen. n. sp.20
Prolucanus beipiaoensis n. gen. n. sp.20 Qi et al. 2022 stag beetle
NIGP 176633
 Coleoptera - Cretohisteridae
Cretohister sinensis n. gen. n. sp.37
Cretohister sinensis n. gen. n. sp.37 Zhou et al. 2018 beetle
NIGP166874
 Dicondylia - Tarsophlebiidae
Turanophlebia liaoningensis n. sp.12 Fang and Zheng 2022 winged insect
NIGP 163539
 Odonata - Aeschnidiidae
Stylaeschnidium rarum n. gen. n. sp.32, "Aeschnidium heishankowense" = Sinaeschnidia heishankowense, Dracontaeschnidium orientale n. gen. n. sp.32
Stylaeschnidium rarum n. gen. n. sp.32 Zhang and Zhang 2001 small darner
MI117
"Aeschnidium heishankowense" = Sinaeschnidia heishankowense Hong 1965 small darner
Adults B93001, B93002, B93003, B93004-2, B93005, B93006, B97601, B97602-1,2, B97603-1, 2; Nymphs B97611, B97613, B97614, B97615, B97619, B97618-1, 2, B97620, B97621-1, 2
Dracontaeschnidium orientale n. gen. n. sp.32 Zhang and Zhang 2001 small darner
B97602
 Odonata - Progobiaeshnidae
Paradecoraeshna liaoningensis n. gen. n. sp.35 Zheng et al. 2018 true dragonfly
NIGP 166151
 Odonata -
Pseudoliupanshania magnanicellula n. gen. n. sp.36 Zheng et al. 2022 true dragonfly
NIGP 174551
 Odonata - Liupanshaniidae
Protoliupanshania wangi36 Huang and Nel 2010 true dragonfly
NIGP 174549
 Odonata -
Parapetala yixianensis n. sp.36 Zheng et al. 2022 true dragonfly
NIGP 174550
 Diptera - Eremochaetidae
Alleremonomus xingi22 Ren and Guo 1995 fly
NIGP Y120604, Y120605
Eremomukha (Eremomukha) tenuissima n. sp.22 Zhang 2014 fly
NIGP Y120603
Eremomukha (Eremomukha) angusta n. sp.22 Zhang 2014 fly
NIGP Y120602
Lepteremochaetus elegans n. sp.22 Zhang 2014 fly
NIGP Y120601