PBDB Collection

Spring site, Middle Fork Flathead River (Kishenehn) (Eocene of the United States)

Where: Montana (48.4° N, 113.7° W: paleocoordinates 51.9° N, 100.4° W)

• coordinate stated in text

When: Coal Creek Member (Kishenehn Formation), Lutetian (47.8 - 41.3 Ma)

• Middle sequence of the Coal Creek member of the Kishenehn Formation, which has been estimated to be 46.2 +/- 0.4 myo (Lutetian) by 40Ar/39Ar analysis and 43.5 +/- 4.9 myo by fission-track analysis

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, carbonaceous shale

• Paper/oil shale

Size classes: macrofossils, mesofossils

Preservation: adpression

Collected by Dale Greenwalt; reposited in the USNM

Collection methods: mechanical,

Primary reference: J. T. Huber and D. Greenwalt. 2011. Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera. ZooKeys 130:473-494 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 117614: authorized by Matthew Clapham, entered by Matthew Clapham on 25.09.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Diptera - Scatopsidae	Psectrosciara crassieton n. sp.² Psectrosciara crassieton n. sp.² Amorim and Greenwalt 2022 minute black scavenger fly USNM 625035
Diptera - Lonchopteridae	Lonchoptera eocenica n. sp.⁴ Lonchoptera eocenica n. sp.⁴ Amorim and Brown 2019 fly USNM 625379
Diptera - Phoridae	Aenigmatias kishenehnensis n. sp.⁴ Aenigmatias kishenehnensis n. sp.⁴ Brown 2019 scuttle fly USNM 625132
Diptera - Mycetophilidae	Azana akarenos n. sp.² Azana akarenos n. sp.² Kerr and Greenwalt 2022 fungus gnat USNM 625099
Diptera - Bibionidae	Bibiodes kishenehnensis n. sp.⁴ Bibiodes kishenehnensis n. sp.⁴ Fitzgerald 2019 march fly USNM 625738
Diptera - Tipulidae	Tipula (Trichotipula) fji n. sp.⁴ Tipula (Trichotipula) fji n. sp.⁴ de Jong 2019 crane fly USNM 625687
Hemiptera - Corixidae	Corixidae indet.⁵ Corixidae indet.⁵ Leach 1815 water boatman USNM 595154 (exact collecting site not given)
Hymenoptera - Formicidae	Ponerites kishenehne n. sp.⁶, Protazteca eocenica n. sp.⁶, Ktunaxia jucunda n. gen. n. sp.⁶, Proiridomyrmex rotundatus n. sp.⁶, Dolichoderus dlusskyi n. sp.⁶, Camponotites spp.⁶, Lasius glom n. sp.⁶, Formica annosa n. sp.⁶, Pseudomyrmex saxulum n. sp.⁶, Solenopsites abdita n. sp.⁶, "Crematogaster aurora n. sp." = Incertogaster aurora⁶, Eoformica latimedia n. sp.⁶, Eoformica pinguis⁶, Eoformica brevipetiola n. sp.⁶ Ponerites kishenehne n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609604 Protazteca eocenica n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609590 Ktunaxia jucunda n. gen. n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609585 Proiridomyrmex rotundatus n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609586, 609587, 609588, 609589 Dolichoderus dlusskyi n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609584 Camponotites spp.⁶ Steinbach 1967 ant 9.2% of 152 specimens Lasius glom n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609593, 609594 Formica annosa n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609591, 609592 Pseudomyrmex saxulum n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609605 Solenopsites abdita n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609598, 609599-609603 "Crematogaster aurora n. sp." = Incertogaster aurora⁶ LaPolla and Greenwalt 2015 ant USNM 609595, 609596 Eoformica latimedia n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609612, 609613, 609614 Eoformica pinguis⁶ Scudder 1877 ant USNM 609606, 609607, 609608, 609609 Eoformica brevipetiola n. sp.⁶ LaPolla and Greenwalt 2015 ant USNM 609610, 609611
Hymenoptera - Ichneumonidae	Ichneumonidae indet.⁵ Ichneumonidae indet.⁵ Latreille 1802 ichneumon wasp USNM 595152 (exact collecting site not given)
Hymenoptera - Mymaridae	Eoeustochus sp., Eoeustochus kishenehn n. gen. n. sp., Eoeustochus borchersi n. sp., Eoanaphes stethynioides n. gen. n. sp., "Gonatocerus greenwalti n. sp." = Cosmocomoidea greenwalti, "Gonatocerus rasnitsyni n. sp." = Cosmocomoidea rasnitsyni, "Gonatocerus kootenai n. sp." = Cosmocomoidea kootenai Eoeustochus sp. Huber 2011 fairyfly Eoeustochus kishenehn n. gen. n. sp. Huber 2011 fairyfly Eoeustochus borchersi n. sp. Huber 2011 fairyfly Eoanaphes stethynioides n. gen. n. sp. Huber 2011 fairyfly "Gonatocerus greenwalti n. sp." = Cosmocomoidea greenwalti Huber 2011 fairyfly "Gonatocerus rasnitsyni n. sp." = Cosmocomoidea rasnitsyni Huber 2011 fairyfly "Gonatocerus kootenai n. sp." = Cosmocomoidea kootenai Huber 2011 fairyfly
Coleoptera - Mordellidae	Mordellidae indet.⁵ Mordellidae indet.⁵ Latreille 1802 tumbling flower beetle USNM 595151 (exact collecting site not given)
Coleoptera - Staphylinidae	Tympanophorus greenwalti n. sp.¹, Pinophilini indet.⁵ Tympanophorus greenwalti n. sp.¹ Chatzimanolis et al. 2017 rove beetle USNM 595140 Pinophilini indet.⁵ Nordmann 1837 rove beetle USNM 595153 (exact collecting site not given)
Coleoptera - Ptiliidae	Ptenidium kishenehnicum n. sp.⁷ Ptenidium kishenehnicum n. sp.⁷ Shockley and Greenwalt 2013 feather-winged beetle USNM 545816
Odonata - Eolestidae	Eolestes syntheticus³ Eolestes syntheticus³ Cockerell 1940 damselfly USNM 559049

¹Brunke et al. 2017; ²Greenwalt et al. 2022; ³Greenwalt and Bechly 2014; ⁴Greenwalt et al. 2019; ⁵Greenwalt et al. 2016; ⁶LaPolla and Greenwalt 2015; ⁷Shockley and Greenwalt 2013