PBDB Collection

Cassville (Permian of the United States)

Where: Monongolia County, West Virginia (39.7° N, 80.1° W: paleocoordinates 5.1° S, 16.0° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Cassville Shales Formation, Asselian (298.9 - 295.5 Ma)

• "...roof shales of the Waynesburg coal, or the very highest of the beds of the upper productive coal series, in the nomenclature of the first Pennsylvania survey, or the beds termed Permo-carboniferous by Professor Fontaine." Scudder, 1879. Asselian (or possibly earliest Sakmarian) according to Schneider et al. (2013).

Environment/lithology: terrestrial; shale

Size class: macrofossils

Preservation: mold/impression

Collected by Mr. Fontaine, Mr. White & R.D. Lacoe

Primary reference: S. H. Scudder. 1879. Palaeozoic Cockroaches: A complete revision of the species of both worlds, with an essay toward their classification. Memoirs of the Boston Society of Natural History 3:23-134 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122604: authorized by Matthew Clapham, entered by Jered Karr on 03.01.2012, edited by Amy Delelli and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Blattodea - Archimylacridae	Gerablattina inculta n. sp.⁴, Etoblattina sp.⁴, "Gerablattina ovata n. sp." = Liparoblatta ovata⁴, "Gerablattina radiata n. sp." = Liparoblatta radiata⁴, "Etoblattina arcta n. sp." = Scudderula arcta⁴, Gerablattina eversa n. sp.⁴, "Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana⁴, "Gerablattina permacra n. sp." = Acosmoblatta permacra⁴ Gerablattina inculta n. sp.⁴ Scudder 1895 cockroach Coll Lacoe 2139a Etoblattina sp.⁴ Scudder 1879 cockroach Coll Lacoe 2084 ab; hindwing "Gerablattina ovata n. sp." = Liparoblatta ovata⁴ Scudder 1895 cockroach Coll Lacoe 2150 ab "Gerablattina radiata n. sp." = Liparoblatta radiata⁴ Scudder 1895 cockroach Coll Lacoe 2147a "Etoblattina arcta n. sp." = Scudderula arcta⁴ Scudder 1895 cockroach Coll Lacoe 2118a Gerablattina eversa n. sp.⁴ Scudder 1895 cockroach Coll Lacoe 2145a "Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana⁴ Scudder 1895 cockroach Coll Lacoe 2111 ab "Gerablattina permacra n. sp." = Acosmoblatta permacra⁴ Scudder 1895 cockroach USGS 2187(2188) collected by Scudder
Blattodea -	Poroblattina gratiosa n. sp.⁴ Poroblattina gratiosa n. sp.⁴ Scudder 1895 cockroach Coll Lacoe 2134 ab (USNM 38166)
Blattodea - Poroblattinidae	"Petrablattina hastata n. sp." = Poroblattina parvula⁴ "Petrablattina hastata n. sp." = Poroblattina parvula⁴ Goldenberg 1869 cockroach Coll Lacoe 2133 ab
Blattodea - Mesoblattinidae	"Etoblattina expuncta n. sp." = Pareinoblatta expuncta⁴ "Etoblattina expuncta n. sp." = Pareinoblatta expuncta⁴ Scudder 1895 cockroach Coll Lacoe 2109 ab
Blattodea -	Gerablattina perita n. sp.⁴, Etoblattina exigua n. sp.⁴ Gerablattina perita n. sp.⁴ Scudder 1895 cockroach Coll Lacoe 2140 ab Etoblattina exigua n. sp.⁴ Scudder 1895 cockroach Coll Lacoe 2107 ab
Dicondylia - Phyloblattidae	"Etoblattina communis n. sp." = Phyloblatta communis⁴, Phyloblatta abbreviata n. sp.¹, "Etoblattina praedulcis n. sp." = Phyloblatta praedulcis⁴, "Etoblattina rogi n. sp." = Phyloblatta rogi⁴, "Etoblattina imperfecta n. sp." = Phyloblatta imperfecta⁴, Phyloblatta elatior n. sp.¹, "Etoblattina exsecuta n. sp." = Phyloblatta exsecuta⁴, "Gerablattina deducta n. sp." = Phyloblatta deducta⁴, "Etoblattina macilenta n. sp." = Phyloblatta macilenta⁴, Phyloblatta debilis n. sp.¹, "Etoblattina residua n. sp." = Phyloblatta residua⁴, "Etoblattina lata n. sp." = Phyloblatta lata⁴, "Etoblattina mactata n. sp." = Phyloblatta mactata⁴, ? Phyloblatta dimidiata n. sp.¹, Phyloblatta mortua n. sp.¹, Phyloblatta macroptera n. sp.¹, "Etoblattina macerata n. sp." = Phyloblatta macerata⁴, "Etoblattina expugnata n. sp." = Phyloblatta expugnata⁴, Phyloblatta arcuata n. sp.¹, Phyloblatta fracta n. sp.¹, "Etoblattina angusta n. sp." = Phyloblatta angusta⁴, Phyloblatta scudderiana n. sp.¹, "Etoblattina gratiosa n. sp." = Phyloblatta gratiosa⁴, Phyloblatta vulgata n. sp.¹, Phyloblatta dichotoma n. sp.¹, "Etoblattina obatra n. sp." = Phyloblatta obatra⁴, Phyloblatta virginiana n. sp.¹, ? Phyloblatta rebaptizata n. sp.¹, "Etoblattina funeraria n. sp." = Phyloblatta funeraria⁴, "Etoblattina immolata n. sp." = Phyloblatta immolata⁴, "Gerablattina abdicata n. sp." = Phyloblatta abdicata⁴, Phyloblatta cassvilleana n. sp.¹, "Gerablattina concinna n. sp." = Phyloblatta concinna⁴, Phyloblatta regularis n. sp.¹, "Etoblattina expulsata n. sp." = Phyloblatta expulsata⁴, "Etoblattina secreta n. sp." = Phyloblatta secreta⁴, "Gerablattina uniformis n. sp." = Phyloblatta uniformis⁴, "Etoblattina accubita n. sp." = Phyloblatta accubita⁴ "Etoblattina communis n. sp." = Phyloblatta communis⁴ Scudder 1895 winged insect Coll Lacoe 2187 ab, cd, e, f, g, h, i, j, kl, mn (USNM 38188; Handlirsch assigns 6 specimens to new species) Phyloblatta abbreviata n. sp.¹ Handlirsch 1906 winged insect USNM 38588 "Etoblattina praedulcis n. sp." = Phyloblatta praedulcis⁴ Scudder 1895 winged insect Coll Lacoe 2119 ab (USNM 38165) "Etoblattina rogi n. sp." = Phyloblatta rogi⁴ Scudder 1895 winged insect Coll Lacoe 2122 a, bc (USNM 38088) "Etoblattina imperfecta n. sp." = Phyloblatta imperfecta⁴ Scudder 1895 winged insect Coll Lacoe 2124 ab (USNM 38083) Phyloblatta elatior n. sp.¹ Handlirsch 1906 winged insect USNM 38087 (specimen of E. communis in Scudder, 1895, pl 7, fig 14) "Etoblattina exsecuta n. sp." = Phyloblatta exsecuta⁴ Scudder 1895 winged insect Coll Lacoe 2120 ab, cd, e (USNM 38180; others assigned to new species by Handlirsch) "Gerablattina deducta n. sp." = Phyloblatta deducta⁴ Scudder 1895 winged insect Coll Lacoe 2146 a (USNM 38063) "Etoblattina macilenta n. sp." = Phyloblatta macilenta⁴ Scudder 1895 winged insect Coll Lacoe 2121 ab (USNM 38163) Phyloblatta debilis n. sp.¹ Handlirsch 1906 winged insect USNM 38094 (specimen of E. immolata in Scudder, 1895, pl. 7, fig 8) "Etoblattina residua n. sp." = Phyloblatta residua⁴ Scudder 1895 winged insect Coll Lacoe 2131a (USNM 38179) "Etoblattina lata n. sp." = Phyloblatta lata⁴ Scudder 1895 winged insect Coll Lacoe 2099a (USNM 38200) "Etoblattina mactata n. sp." = Phyloblatta mactata⁴ Scudder 1895 winged insect Coll Lacoe 2116a (USNM 38081) ? Phyloblatta dimidiata n. sp.¹ Handlirsch 1906 winged insect USNM 38905 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 11)) Phyloblatta mortua n. sp.¹ Handlirsch 1906 winged insect USNM 38899 (specimen of E. communis in Scudder, 1895, pl 7, fig 15) Phyloblatta macroptera n. sp.¹ Handlirsch 1906 winged insect USNM 38891 (specimen of E. communis in Scudder, 1895, pl 7, fig 17) "Etoblattina macerata n. sp." = Phyloblatta macerata⁴ Scudder 1895 winged insect Coll Lacoe 2081 ab, c (USNM 38183) "Etoblattina expugnata n. sp." = Phyloblatta expugnata⁴ Scudder 1895 winged insect Coll Lacoe 2085 ab (USNM 38193) Phyloblatta arcuata n. sp.¹ Handlirsch 1906 winged insect USNM 38898 (specimen of E. communis in Scudder, 1895, pl 7, fig 13) Phyloblatta fracta n. sp.¹ Handlirsch 1906 winged insect USNM 38897 (specimen of E. communis in Scudder, 1895, pl 7, fig 12) "Etoblattina angusta n. sp." = Phyloblatta angusta⁴ Scudder 1895 winged insect Coll Lacoe 2080 ab (USNM 38185) Phyloblatta scudderiana n. sp.¹ Handlirsch 1906 winged insect USNM 38904 (specimen of G. concinna in Scudder, 1895, pl 10, fig 5) "Etoblattina gratiosa n. sp." = Phyloblatta gratiosa⁴ Scudder 1895 winged insect Coll Lacoe 2114 ab Phyloblatta vulgata n. sp.¹ Handlirsch 1906 winged insect USNM 38901 (specimen of E. expulsata in Scudder, 1895, pl. 9, fig 4) Phyloblatta dichotoma n. sp.¹ Handlirsch 1906 winged insect USNM 38896 (specimen of E. communis in Scudder, 1895, pl 7, fig 11) "Etoblattina obatra n. sp." = Phyloblatta obatra⁴ Scudder 1895 winged insect Coll Lacoe 2123 ab (USNM 38087) Phyloblatta virginiana n. sp.¹ Handlirsch 1906 winged insect USNM 38902 (specimen of E. secreta in Scudder, 1895, pl. 9, fig 7) ? Phyloblatta rebaptizata n. sp.¹ Handlirsch 1906 winged insect USNM 38106 (specimen of P. gratiosa in Scudder, 1895, pl 11, fig 13) "Etoblattina funeraria n. sp." = Phyloblatta funeraria⁴ Scudder 1895 winged insect Coll Lacoe 2108 ab, c (USNM 38078) "Etoblattina immolata n. sp." = Phyloblatta immolata⁴ Scudder 1895 winged insect Coll Lacoe 2115 ab, cd (USNM 38079; other specimen assigned to P. debilis Handlirsch) "Gerablattina abdicata n. sp." = Phyloblatta abdicata⁴ Scudder 1895 winged insect Coll Lacoe 2143a (USNM 38065) Phyloblatta cassvilleana n. sp.¹ Handlirsch 1906 winged insect USNM 38892 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 10) "Gerablattina concinna n. sp." = Phyloblatta concinna⁴ Scudder 1895 winged insect Coll Lacoe 2077 ab, cd (USNM 38172; Handlirsch assigns 1 to P. scudderiana) Phyloblatta regularis n. sp.¹ Handlirsch 1906 winged insect USNM 38893 (specimen of G. uniformis in Scudder, 1895, pl. 10, fig 9) "Etoblattina expulsata n. sp." = Phyloblatta expulsata⁴ Scudder 1895 winged insect Coll Lacoe 2113 ab, cd, ef (USNM 38178; Handlirsch assigns 1 to P. vulgata) "Etoblattina secreta n. sp." = Phyloblatta secreta⁴ Scudder 1895 winged insect Coll Lacoe 2125 ab, cd, ef, g (USNM 38167; Handlirsch assigns 1 to P. virginiana) "Gerablattina uniformis n. sp." = Phyloblatta uniformis⁴ Scudder 1895 winged insect Coll Lacoe 2082 ab, cd, e, fg (USNM 38177; Handlirsch assigns 3 to other species) "Etoblattina accubita n. sp." = Phyloblatta accubita⁴ Scudder 1895 winged insect Coll Lacoe 2112a (USNM 38169)
Dicondylia - Spiloblattinidae	Arrhythmoblatta scudderiana n. sp.², "Etoblattina detecta n. sp." = Arrhythmoblatta detecta⁴, "Etoblattina defossa n. sp." = Dicladoblatta defossa⁴, "Etoblattina recidiva n. sp." = Sysciophlebia recidiva⁴, "Gerablattina balteata n. sp." = Sysciophlebia balteata, "Sysciophlebia balteata" = Sysciophlebia balteata³, "Gerablattina diversinervis n. sp." = Sysciophlebia balteata⁴, "Etoblattina occulta n. sp." = Sysciophlebia balteata⁴, "Gerablattina cassvici n. sp." = Sysciophlebia balteata⁴, "Etoblattina patiens n. sp." = Sysciophlebia patiens⁴, "Etoblattina invisa n. sp." = Sysciophlebia invisa⁴, Etoblattina aperta n. sp.⁴ Arrhythmoblatta scudderiana n. sp.² Handlirsch 1906 winged insect USNM 38894 (specimen of E. detecta in Scudder, 1895; pl 4, fig 13) "Etoblattina detecta n. sp." = Arrhythmoblatta detecta⁴ Scudder 1895 winged insect Coll Lacoe 2106 ab, c "Etoblattina defossa n. sp." = Dicladoblatta defossa⁴ Scudder 1895 winged insect Coll Lacoe 2128 ab "Etoblattina recidiva n. sp." = Sysciophlebia recidiva⁴ Scudder 1895 winged insect Coll Lacoe 2129 a "Gerablattina balteata n. sp." = Sysciophlebia balteata Scudder 1879 winged insect "Sysciophlebia balteata" = Sysciophlebia balteata³ Scudder 1879 winged insect "Gerablattina diversinervis n. sp." = Sysciophlebia balteata⁴ Scudder 1879 winged insect Coll Lacoe 2141 ab "Etoblattina occulta n. sp." = Sysciophlebia balteata⁴ Scudder 1879 winged insect Coll Lacoe 2127 ab "Gerablattina cassvici n. sp." = Sysciophlebia balteata⁴ Scudder 1879 winged insect Coll Lacoe 2142 ab, cd "Etoblattina patiens n. sp." = Sysciophlebia patiens⁴ Scudder 1895 winged insect Coll Lacoe 2104 ab "Etoblattina invisa n. sp." = Sysciophlebia invisa⁴ Scudder 1895 winged insect Coll Lacoe 2126 ab Etoblattina aperta n. sp.⁴ Scudder 1895 winged insect Coll Lacoe 2066 ab
Dicondylia - Petrablattinidae	"Etoblattina debilis n. sp." = Symphyoblatta debilis⁴, "Etoblattina ovata n. sp." = Petrablattina ovata⁴, "Gerablattina permanenta n. sp." = Amoeboblatta permanenta⁴, "Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis⁴, "Gerablattina rotundata n. sp." = Penetoblatta rotundata⁴, "Gerablattina lata n. sp." = Amblyblatta lata⁴ "Etoblattina debilis n. sp." = Symphyoblatta debilis⁴ Scudder 1895 winged insect Coll Lacoe 2103 ab, cd, e "Etoblattina ovata n. sp." = Petrablattina ovata⁴ Scudder 1895 winged insect Coll Lacoe 2102a (USNM 38201) "Gerablattina permanenta n. sp." = Amoeboblatta permanenta⁴ Scudder 1895 winged insect Coll Lacoe 2144 ab "Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis⁴ Scudder 1895 winged insect Coll Lacoe 2138 ab "Gerablattina rotundata n. sp." = Penetoblatta rotundata⁴ Scudder 1895 winged insect Coll Lacoe 2149a "Gerablattina lata n. sp." = Amblyblatta lata⁴ Scudder 1895 winged insect Coll Lacoe 2148a
Dicondylia - Schizoblattidae	"Poroblattina complexinervis n. sp." = Apempherus complexinervis⁴, "Poroblattina fossa n. sp." = Apempherus fossus⁴ "Poroblattina complexinervis n. sp." = Apempherus complexinervis⁴ Scudder 1895 winged insect Coll Lacoe 2135 ab "Poroblattina fossa n. sp." = Apempherus fossus⁴ Scudder 1895 winged insect Coll Lacoe 2083 ab, c
Dicondylia - Bradyblattidae	"Etoblattina mucronata n. sp." = Bradyblatta mucronata⁴, "Etoblattina sagittaria n. sp." = Bradyblatta sagittaria⁴, "Etoblattina mediana n. sp." = Bradyblatta sagittaria⁴ "Etoblattina mucronata n. sp." = Bradyblatta mucronata⁴ Scudder 1895 winged insect Coll lacoe 2105 ab (USNM 38199) "Etoblattina sagittaria n. sp." = Bradyblatta sagittaria⁴ Scudder 1895 winged insect Coll Lacoe 2100a, b, c, d "Etoblattina mediana n. sp." = Bradyblatta sagittaria⁴ Scudder 1895 winged insect Coll Lacoe 2101 ab (USNM 38198)

¹Handlirsch 1906; ²Rees and Evans 2002; ³Schneider 1978; ⁴Scudder 1895