PBDB Collection

Willis Creek near Steubenville (Carboniferous of the United States)

Also known as Richmond; Wills Creek

Where: Jefferson County, Ohio (40.4° N, 80.7° W: paleocoordinates 6.4° S, 17.2° W)

• coordinate based on political unit

• small collection-level geographic resolution

When: Conemaugh Formation, Stephanian (307.5 - 300.4 Ma)

• "Conemaugh formation; Shales above Arnes Limestone. Oberes Obercarbon"

•"Lower Barren Coal-measures of Ohio, less than 100 feet above the crinoidal limestone…"Scudder, 1895

Environment/lithology: terrestrial; lithified, gray, green shale

• Appalachian foreland basin

• were found in a medium-dark-gray to olive-gray calcareous shale that dominates the outcrop. This shale also contains disseminated plant debris, scattered root traces, and abundant bivalves. Coincident

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Collected by S. Huston; reposited in the USNM

Collection methods: quarrying,

Primary reference: S. H. Scudder. 1890. An interesting Paleozoic cockroach fauna at Richmond, Ohio. Proceedings of the Boston Society of Natural History 24:45-53 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 123812: authorized by Matthew Clapham, entered by Jered Karr on 31.01.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Paoliida - Blattinopsidae	"Oryctoblattina americana n. sp." = Blattinopsis americana¹, "Oryctoblattina latipennis n. sp." = Blattinopsis latipennis¹, Blattinopsis anthracina n. sp.¹ "Oryctoblattina americana n. sp." = Blattinopsis americana¹ Handlirsch 1906 winged insect USNM 38647 "Oryctoblattina latipennis n. sp." = Blattinopsis latipennis¹ Handlirsch 1906 winged insect USNM 38656 Blattinopsis anthracina n. sp.¹ Handlirsch 1906 winged insect USNM 38679
Dicondylia - Bradyblattidae	"Etoblattina fossa n. sp." = Metaxys fossa² "Etoblattina fossa n. sp." = Metaxys fossa² Scudder 1895 winged insect Coll Huston 182
Dicondylia - Schizoblattidae	Schizoblatta alutacea n. gen. n. sp.¹ Schizoblatta alutacea n. gen. n. sp.¹ Handlirsch 1906 winged insect USNM 38668
Dicondylia - Spiloblattinidae	"Etoblattina strigosa n. sp." = Ametroblatta strigosa, Syscioblatta steubenvilleana n. sp.¹, Syscioblatta obscura n. sp.¹, Syscioblatta misera n. sp.¹, Syscioblatta anomala n. sp.¹, Syscioblatta minor n. sp.¹, Doryblatta longipennis n. gen. n. sp.¹, Sysciophlebia nana n. sp.¹, Sysciophlebia obtusa n. sp.¹, Sysciophlebia acutipennis n. sp.¹, Sysciophlebia rotundata n. sp.¹, Sysciophlebia picta n. sp.¹, "Etoblattina benedicta n. sp." = Sysciophlebia benedicta², Sysciophlebia adumbrata n. sp.¹, Sysciophlebia schucherti n. sp.¹, Sysciophlebia affinis n. sp.¹, Sysciophlebia hybrida n. sp.¹, Sysciophlebia whitei n. sp.¹, Sysciophlebia scudderi n. sp.¹, "Etoblattina variegata n. sp." = Sysciophlebia variegata, Atactoblatta anomala n. gen. n. sp.¹, "Gerablattina apicalis n. sp." = Spiloblattina variegata², "Etoblattina tenuis n. sp." = Spiloblattina variegata, "Etoblattina hustoni n. sp." = Spiloblattina variegata, "Poroblattina longinqua n. sp." = Spiloblattina variegata², "Etoblattina marginata n. sp." = Spiloblattina variegata, "Etoblattina fasciata n. sp." = Spiloblattina variegata, "Etoblattina hastata n. sp." = Spiloblattina variegata², "Etoblattina exsensa n. sp." = Spiloblattina variegata², "Etoblattina gracilenta n. sp." = Spiloblattina variegata², "Etoblattina maledicta n. sp." = Spiloblattina variegata², "Etoblattina willsiana n. sp." = Spiloblattina variegata², "Etoblattina stipata n. sp." = Spiloblattina variegata, "Etoblattina funesta n. sp." = Spiloblattina variegata², "Etoblattina ramosa n. sp." = Spiloblattina variegata² "Etoblattina strigosa n. sp." = Ametroblatta strigosa Scudder 1890 winged insect coll. Henry Crewthen Syscioblatta steubenvilleana n. sp.¹ Handlirsch 1906 winged insect USNM 38671 Syscioblatta obscura n. sp.¹ Handlirsch 1906 winged insect E. maledicta in Scudder, 1895 Pl 6 fig 2 Syscioblatta misera n. sp.¹ Handlirsch 1906 winged insect USNM 38658 Syscioblatta anomala n. sp.¹ Handlirsch 1906 winged insect USNM 38653 Syscioblatta minor n. sp.¹ Handlirsch 1906 winged insect USNM 38665 Doryblatta longipennis n. gen. n. sp.¹ Handlirsch 1906 winged insect USNM 38662 Sysciophlebia nana n. sp.¹ Handlirsch 1906 winged insect USNM 38648 Sysciophlebia obtusa n. sp.¹ Handlirsch 1906 winged insect USNM 38660 Sysciophlebia acutipennis n. sp.¹ Handlirsch 1906 winged insect USNM 38639 Sysciophlebia rotundata n. sp.¹ Handlirsch 1906 winged insect USNM 38651 Sysciophlebia picta n. sp.¹ Handlirsch 1906 winged insect USNM 38673 "Etoblattina benedicta n. sp." = Sysciophlebia benedicta² Scudder 1895 winged insect Coll Huston 166(167), 168, 178, 179(191), 183(193), 199 Sysciophlebia adumbrata n. sp.¹ Handlirsch 1906 winged insect USNM 38640 Sysciophlebia schucherti n. sp.¹ Handlirsch 1906 winged insect USNM 38691 Sysciophlebia affinis n. sp.¹ Handlirsch 1906 winged insect E. benedicta in Scudder, 1895 pl 5 fig 15 Sysciophlebia hybrida n. sp.¹ Handlirsch 1906 winged insect E. maledicta in Scudder, 1895 fig 3 Sysciophlebia whitei n. sp.¹ Handlirsch 1906 winged insect USNM 38696 Sysciophlebia scudderi n. sp.¹ Handlirsch 1906 winged insect E. gracilenta in Scudder, 1895 fig 7 "Etoblattina variegata n. sp." = Sysciophlebia variegata Scudder 1890 winged insect Atactoblatta anomala n. gen. n. sp.¹ Handlirsch 1906 winged insect USNM 38698 "Gerablattina apicalis n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 112 "Etoblattina tenuis n. sp." = Spiloblattina variegata Scudder 1890 winged insect "Etoblattina hustoni n. sp." = Spiloblattina variegata Scudder 1890 winged insect "Poroblattina longinqua n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 111 "Etoblattina marginata n. sp." = Spiloblattina variegata Scudder 1890 winged insect "Etoblattina fasciata n. sp." = Spiloblattina variegata Scudder 1890 winged insect "Etoblattina hastata n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 106 "Etoblattina exsensa n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 161, 196 "Etoblattina gracilenta n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 114, 198, 105 "Etoblattina maledicta n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 163, 164, 165, 170, 174, 181, 192 "Etoblattina willsiana n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 176 "Etoblattina stipata n. sp." = Spiloblattina variegata Scudder 1890 winged insect "Etoblattina funesta n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 107(177), 180 USGS 2186 "Etoblattina ramosa n. sp." = Spiloblattina variegata² Scudder 1890 winged insect Coll Huston 108
Blattodea - Poroblattinidae	"Acmaeoblatta lanceolata n. gen. n. sp." = Poroblattina attenuata¹, Poroblattina brachyptera n. sp.¹, Poroblattina richmondiana n. sp.¹, Poroblattina ohioensis n. sp.², Poroblattina lata n. sp.¹, "Gerablattina minima n. sp." = Poroblattina rotundata² "Acmaeoblatta lanceolata n. gen. n. sp." = Poroblattina attenuata¹ Scudder 1885 cockroach USNM 38678 Poroblattina brachyptera n. sp.¹ Handlirsch 1906 cockroach USNM 38637 Poroblattina richmondiana n. sp.¹ Handlirsch 1906 cockroach USNM 38644 Poroblattina ohioensis n. sp.² Scudder 1895 cockroach Coll Huston 162 Poroblattina lata n. sp.¹ Handlirsch 1906 cockroach USNM 38696 "Gerablattina minima n. sp." = Poroblattina rotundata² Scudder 1885 cockroach USGS 2183(2184)
Blattodea - Archimylacridae	Stygetoblatta latipennis n. gen. n. sp.¹ Stygetoblatta latipennis n. gen. n. sp.¹ Handlirsch 1906 cockroach USNM 38642
Blattodea -	Gerablattina richmondiana n. sp.² Gerablattina richmondiana n. sp.² Scudder 1895 cockroach Coll Huston 169
Dicondylia - Phyloblattidae	"Etoblattina jeffersoniana n. sp." = Phyloblatta jeffersoniana² "Etoblattina jeffersoniana n. sp." = Phyloblatta jeffersoniana² Scudder 1895 winged insect Coll Huston 186(184)

¹Handlirsch 1906; ²Scudder 1895