Quarry 9, Como Bluff (YPM) (Jurassic to of the United States)

Also known as Mammal Quarry, WY-15

Where: Albany County, Wyoming (41.9° N, 106.0° W: paleocoordinates 37.5° N, 52.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Brushy Basin Member (Morrison Formation), Kimmeridgian to Kimmeridgian (157.3 - 145.0 Ma)

• middle upper part of the Morrison series

• bed-level stratigraphic resolution

Environment/lithology: fluvial-lacustrine; gray, green mudstone and coarse-grained sandstone

• Several hypotheses for the paleoenvironment (lacustrine, ephemeral ponds)
• Secondary lithology described as 'coarse sandstone'

Size classes: macrofossils, mesofossils

Collected by W. Reed; reposited in the USNM, YPM

Collection methods: bulk, quarrying, mechanical, sieve,

Primary reference: O. C. Marsh. 1881. Discovery of a fossil bird in the Jurassic of Wyoming. American Journal of Science 31:341-342 [M. Carrano/M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 12816: authorized by Emmanuel Fara, entered by Emmanuel Fara on 18.12.2001, edited by Robin Whatley, Matthew Carrano and Roger Benson

Creative Commons license: CC BY (attribution)

Taxonomic list

• Revision of previously proposed taxa. The fauna is said to also contain 'fish, small reptiles, and mammals'.
Bivalvia
 Unionida - Unionidae
Unio sp.25 Philipsson 1788 freshwater mussel
Gastropoda
 Basommatophora - Planorbidae
Planorbis sp.25 Muller 1774 pulmonate
 Valvatoidea - Valvatidae
Valvatidae indet.2 Gray 1840 snail
Gnathostomata
  -
Actinopterygii indet.2 ray-finned fish
Actinopteri
 Halecomorphi - Ophiopsiidae
Ophiopsis sp.19 Agassiz 1843
Osteichthyes
 Dipnoi - Ceratodontidae
Ceratodus sp.19 Agassiz 1838 lungfish
Ceratodus guentheri11 Marsh 1878 lungfish
 Salientia - Pelobatidae
Pelobatidae indet.3 Lataste 1879 spadefoot toad
Lissamphibia, Anura
 Salientia -
Eobatrachus agilis n. gen. n. sp.16, "Comobatrachus aenigmaticus n. gen. n. sp." = Comobatrachus aenigmatis10
Eobatrachus agilis n. gen. n. sp.16 Marsh 1887 frog
YPM 1862
"Comobatrachus aenigmaticus n. gen. n. sp." = Comobatrachus aenigmatis10 Hecht and Estes 1960 frog
YPM 1863
 Salientia - Discoglossidae
Enneabatrachus hechti n. gen. n. sp.3
Enneabatrachus hechti n. gen. n. sp.3 Evans and Milner 1993 painted frog
Lissamphibia, Anura, Discoglossidae
 Caudata -
Comonecturoides marshi n. gen. n. sp.10
Comonecturoides marshi n. gen. n. sp.10 Hecht and Estes 1960 salamander
YPM 3919
Reptilia
 Testudines - Pleurosternidae
Dinochelys whitei19, "Glyptops plicatulus" = Glyptops plicatulus4, "Glyptops ornatus n. gen. n. sp." = Glyptops plicatulus17
Dinochelys whitei19 Gaffney 1979 turtle
YPM 6057, right epiplastron, entoplastron, left hyoplastron, right and left hypoplastra, right and left xiphiplastra, and two left bridge peripherals. YPM 6078, right epiplastron; YPM 6068, costal fragment and two bridge peripherals; YPM 6089, right sixth costal; YPM 6069, left seventh costal and associated neural; various costal fragments: YPM 6056, 6079, 6081, 6084, 6085, 6090, 6092, 6093, 6096; various peripherals: 6070, 6071, 6072, 6073, 6074, 6075, 6080, 6103, 6104, 6106, 6113, 6119
"Glyptops plicatulus" = Glyptops plicatulus4 Cope 1877 turtle
Marsh's collections from Quarry Nine at Como Bluff, Wyoming, and now housed in the Yale Peabody Museum and the National Museum of Natural History contain many turtle shell fragments that can be identified on the basis of ornament as Glyptops plicatulus. This material is uncatalogued and not listed here
"Glyptops ornatus n. gen. n. sp." = Glyptops plicatulus17 Cope 1877 turtle
YPM 1784 (figs. 12-15; type of Glyptops ornatus Marsh, 1890, p. 177, pl. 7, fig. 1; Marsh, 1897, fig. 63; Hay, 1908a, pl. 5 fig. 2), a laterally compressed skull without lower jaws. Locality: Reed's YPM Quarry 9 (mammal quarry), T 22 N, R 77 W, sec. 12, Como Bluff, Wyoming (see Ostrom and McIntosh, 1966, for background, maps, and quarry list). Horizon: Morrison Formation, Late Jurassic. Collector: William H. Reed. YPM 1357 (figured by Marsh as Glyptops ornatus, 1890, pl. 7, fig. 2; Hay, 1908, fig. 18), carapace (not seen). YPM 4742, anterior end of right lower jaw ramus. YPM 4721, fragmentary left dentary. YPM 4717, braincase (fig. 19-22) consisting of basisphenoid, basioccipital, exoccipitals and prootics. YPM 5821, left lower jaw. YPM 2753, left lower jaw. YPM 6077, left epiplastron. NMNH 26370, basioccipital. NMNH 26371, basioccipital.
 Sauria -
? Sauria indet.10 diapsid
 Saurischia -
"Morosaurus sp." = Camarasaurus25
"Morosaurus sp." = Camarasaurus25 Cope 1877 sauropod
 Theropoda -
Theropoda indet.25 theropod
 Avetheropoda - Allosauridae
"Antrodemus valens" = Allosaurus fragilis8
"Antrodemus valens" = Allosaurus fragilis8 Marsh 1877 allosaurid
 Avetheropoda -
Coelurus fragilis7 Marsh 1879 coelurosaur
YPM 1996–7, 9163; USNM 5809–10, 6624–28
 Avetheropoda - Compsognathidae
? Ornitholestes hermanni2 Osborn 1903 compsognathid
 Avetheropoda -
? Stokesosaurus clevelandi2 Madsen 1974 coelurosaur
 Ornithischia -
Ankylosauria indet.6 ornithischian
YPM 7453
 Ornithischia - Stegosauridae
Stegosauridae indet.6 Marsh 1880 ornithischian
USNM 7474, YPM 1938, AMNH 11524
 Neornithischia -
"Laosaurus ? consors" = Othnielosaurus consors25, "Camptonotus sp." = Camptosaurus18, Drinker nisti12, Drinker nisti1, "Othnielia rex" = Nanosaurus rex2, "Laosaurus gracilis" = Nanosaurus rex7, Laosaurus sp.18, Laosaurus celer n. gen. n. sp.18
"Laosaurus ? consors" = Othnielosaurus consors25 Marsh 1894 ornithischian
AMNH 11526, 14328
"Camptonotus sp." = Camptosaurus18 Marsh 1885 bird foot dinosaur
Drinker nisti12 Bakker et al. 1990 bird foot dinosaur
USNM 5808, femur
Drinker nisti1 Bakker et al. 1990 bird foot dinosaur
teeth; YPM 9524
"Othnielia rex" = Nanosaurus rex2 Marsh 1877 bird foot dinosaur
"Laosaurus gracilis" = Nanosaurus rex7 Marsh 1877 bird foot dinosaur
YPM 9523-9525
Laosaurus sp.18 Marsh 1878 bird foot dinosaur
Laosaurus celer n. gen. n. sp.18 Marsh 1878 bird foot dinosaur
YPM 1874
 Pterosauria -
Laopteryx priscus n. gen. n. sp., Comodactylus ostromi n. gen. n. sp.5
Laopteryx priscus n. gen. n. sp. Marsh 1881 pterosaur
YPM 1800, partial crushed braincase
Comodactylus ostromi n. gen. n. sp.5 Galton 1981 pterosaur
YPM (holotype), right fourth metacarpal IV
 Loricata - Goniopholididae
Goniopholis sp.25 Owen 1842 crocodilian
 Loricata -
Macelognathus vagans n. gen. n. sp.15
Macelognathus vagans n. gen. n. sp.15 Marsh 1884 crocodilian
YPM 1415
 Squamata - Dorsetisauridae
Dorsetisaurus sp.20 Hoffstetter 1967 lizard
 Squamata - Paramacellodidae
Paramacellodus sp.20 Hoffstetter 1967 lizard
Paramacellodus sp.19 Hoffstetter 1967 lizard
 Rhynchocephalia - Sphenodontidae
Theretairus antiquus n. gen. n. sp.24
Theretairus antiquus n. gen. n. sp.24 Simpson 1926 rhynchocephalian
YPM 13764
 Rhynchocephalia -
Opisthias rarus n. sp.7 Gilmore 1909 rhynchocephalian
 Choristodera -
Cteniogenys antiquus n. gen. n. sp.9
Cteniogenys antiquus n. gen. n. sp.9 Gilmore 1928 choristodere
USNM 6134
Mammalia
 Mammalia -
Mammalia indet.19 mammal
AMNH 104926
 Eutriconodonta - Amphilestidae
Aploconodon comoensis n. gen. n. sp.22, "Phascolodon gidleyi n. gen. n. sp." = Comodon gidleyi22
Aploconodon comoensis n. gen. n. sp.22 Simpson 1925 mammal
USNM 2791 (partial right mandible)
"Phascolodon gidleyi n. gen. n. sp." = Comodon gidleyi22 Simpson 1925 mammal
USNM 2703 (holotype partial left mandible)
 Eutriconodonta - Triconodontidae
Priacodon lulli n. sp.22, "Tinodon robustus n. sp." = Priacodon robustus21, "Tinodon ferox n. sp." = Priacodon ferox16, Priacodon grandaevus n. sp.22, "Triconodon bisulcus n. sp." = Trioracodon bisulcus13
Priacodon lulli n. sp.22 Simpson 1925 mammal
YPM 13625 (holotype right maxilla) and YPM 10359 (right maxilla)
"Tinodon robustus n. sp." = Priacodon robustus21 Marsh 1879 mammal
YPM 11846 (holotype left mandible), 10343 (mandible)
"Tinodon ferox n. sp." = Priacodon ferox16 Marsh 1880 mammal
Priacodon grandaevus n. sp.22 Simpson 1925 mammal
YPM 10349 (holotype left maxilla) and USNM 2698
"Triconodon bisulcus n. sp." = Trioracodon bisulcus13 Marsh 1880 mammal
3 specimens originally listed, 4 by Simpson (1925): YPM 10345, 10340, 13636, and 11851 (holotype); YPM 10344 and USNM 2699 (left maxillae) added by Simpson (1925, part 2)
 Theriiformes - Dryolestidae
Dryolestidae indet.28, Laolestes sp.28, Laolestes eminens n. gen. n. sp.26, "Melanodon goodrichi n. sp." = Laolestes goodrichi28, "Melanodon oweni n. gen. n. sp." = Laolestes oweni26, "Malthacolestes osborni n. gen. n. sp." = Laolestes oweni28, "Dryolestes priscus n. sp." = Dryolestes priscus26, "Stylacodon validus n. sp." = Dryolestes priscus13, "Asthenodon segnis n. gen. n. sp." = Dryolestes priscus16, "Dryolestes arcuatus n. sp." = Dryolestes priscus26, "Laolestes grandis n. sp." = Dryolestes priscus28, "? Herpetairus humilis n. sp." = Dryolestes priscus28, Dryolestes vorax16, "Stylacodon gracilis n. gen. n. sp." = Amblotherium gracilis26, "Dryolestes gracilis n. sp." = Amblotherium gracilis14, "Laodon venustus n. gen. n. sp." = Amblotherium gracilis16, Miccylotyrans minimus n. gen. n. sp.26, Comotherium richi19
Dryolestidae indet.28 Marsh 1879 mammal
Laolestes sp.28 Simpson 1927 mammal
USNM 2862 (anterior part of left mandible with I1-4, C, and P2, figured as Asthenodon segnis by Marsh (1887, pl. 9, fig. 6)
Laolestes eminens n. gen. n. sp.26 Simpson 1927 mammal
YPM 13719 (holotype left mandible);YPM 10662, USNM 2727 etc. (listed by Simpson 1929) AMNH 104827
"Melanodon goodrichi n. sp." = Laolestes goodrichi28 Simpson 1929 mammal
YPM 13738 (holotype partial right maxilla)
"Melanodon oweni n. gen. n. sp." = Laolestes oweni26 Simpson 1927 mammal
YPM 10663 (holotype partial right maxilla)
"Malthacolestes osborni n. gen. n. sp." = Laolestes oweni28 Simpson 1927 mammal
YPM 13751 (holotype partial right maxilla)
"Dryolestes priscus n. sp." = Dryolestes priscus26 Marsh 1878 mammal
YPM 11820 (holotype partial right mandible); USNM 2772 (neotype right mandible)
"Stylacodon validus n. sp." = Dryolestes priscus13 Marsh 1878 mammal
YPM 11884 (holotype right mandible)
"Asthenodon segnis n. gen. n. sp." = Dryolestes priscus16 Marsh 1878 mammal
YPM 10646 (holotype right mandible
"Dryolestes arcuatus n. sp." = Dryolestes priscus26 Marsh 1878 mammal
Listed by Simpson (1929): YPM 11822 (holotype maxilla), YPM 13740 (neotype maxilla), YPM 13739, 13742, 13741, 13743, 13744, 13746, 13747, USNM 2724, 2762, 2800, 2818, 2845
"Laolestes grandis n. sp." = Dryolestes priscus28 Marsh 1878 mammal
YPM 13727 (holotype partial right mandible)
"? Herpetairus humilis n. sp." = Dryolestes priscus28 Marsh 1878 mammal
YPM 13745 (holotype left maxilla)
Dryolestes vorax16 Marsh 1879 mammal
"Stylacodon gracilis n. gen. n. sp." = Amblotherium gracilis26 Marsh 1879 mammal
Specimens listed by Simpson (1929): YPM 11883 (holotype left mandible), USNM 2693, YPM 13731-13733 (mandibles)
"Dryolestes gracilis n. sp." = Amblotherium gracilis14 Marsh 1879 mammal
AMNH 101134, and specimens listed by Simpson (1929): YPM 11821 (holotype left mandible), YPM 13728, 13730, 13734, USNM 2726 (mandibles)
"Laodon venustus n. gen. n. sp." = Amblotherium gracilis16 Marsh 1879 mammal
USNM 2142 (holotype left mandible)
Miccylotyrans minimus n. gen. n. sp.26 Simpson 1927 mammal
USNM 2754 (holotype left upper jaw)
Comotherium richi19 Prothero 1981 mammal
AMNH 104826
 Theriiformes - Paurodontidae
Archaeotrigon sp.28, Archaeotrigon brevimaxillus n. gen. n. sp.26, Paurodon valens n. gen. n. sp.16, Euthlastus cordiformis n. gen. n. sp.26, "Tanaodon agilis n. gen. n. sp." = Tathiodon agilis26, "Pelicopsis dubius n. gen. n. sp." = Tathiodon agilis26
Archaeotrigon sp.28 Simpson 1927 mammal
YPM 13642 (mandible with last M)
Archaeotrigon brevimaxillus n. gen. n. sp.26 Simpson 1927 mammal
USNM 2793
Paurodon valens n. gen. n. sp.16 Marsh 1887 mammal
USNM 2143 (holotype left mandible)
Euthlastus cordiformis n. gen. n. sp.26 Simpson 1927 mammal
YPM 13755 (holotype right maxilla)
"Tanaodon agilis n. gen. n. sp." = Tathiodon agilis26 Simpson 1927 mammal
YPM 13649 (holotype left mandible)
"Pelicopsis dubius n. gen. n. sp." = Tathiodon agilis26 Simpson 1927 mammal
YPM 13754 (holotype right maxilla)
 Docodonta - Docodontidae
"Diplocynodon victor n. gen. n. sp." = Docodon victor13, "Enneodon affinis n. sp." = Docodon affinis16, "Enneodon crassus n. gen. n. sp." = Docodon crassus16, Docodon superus n. sp.28, Docodon striatus n. gen. n. sp.14
"Diplocynodon victor n. gen. n. sp." = Docodon victor13 Marsh 1880 dodocont
YPM 11826 (holotype right mandible)
"Enneodon affinis n. sp." = Docodon affinis16 Marsh 1887 dodocont
USNM 2129 (holotype left mandible)
"Enneodon crassus n. gen. n. sp." = Docodon crassus16 Marsh 1887 dodocont
USNM 2130 (holotype right mandible)
Docodon superus n. sp.28 Simpson 1929 dodocont
YPM 10647 (holotype right maxilla)
Docodon striatus n. gen. n. sp.14 Marsh 1881 dodocont
Including YPM 11823 (holotype right mandible)
  - Amphidontidae
Amphidon superstes n. gen. n. sp.23
Amphidon superstes n. gen. n. sp.23 Simpson 1925 mammal
YPM 13638 (holotype partial right mandible)
 Multituberculata - Plagiaulacidae
Plagiaulacidae indet.19 Gill 1872 multituberculate
AMNH 104795
 Multituberculata - Allodontidae
"Allodon fortis n. sp." = Psalodon fortis16, "Ctenacodon potens n. sp." = Psalodon potens16, Psalodon marshi n. sp.28, "Allodon laticeps n. gen. n. sp." = Ctenacodon laticeps14, Ctenacodon scindens n. sp.28, "Ctenacodon serratus n. sp." = Ctenacodon serratus27, "Ctenacodon nanus n. sp." = Ctenacodon serratus14
"Allodon fortis n. sp." = Psalodon fortis16 Marsh 1887 multituberculate
YPM 11760 (holotype right premaxilla)
"Ctenacodon potens n. sp." = Psalodon potens16 Marsh 1887 multituberculate
YPM 11834 (holotype right upper jaw), YPM 10363, USNM 2680, AMNH 3003
Psalodon marshi n. sp.28 Simpson 1929 multituberculate
USNM 2684 (holotype left mandible), USNM 2687, 2690, 13669, 2679
"Allodon laticeps n. gen. n. sp." = Ctenacodon laticeps14 Marsh 1881 multituberculate
YPM 11761 (holotype left maxilla), USNM 2681, 2682 (partial left maxilla)
Ctenacodon scindens n. sp.28 Simpson 1928 multituberculate
YPM 10366 (holotype left mandible)
"Ctenacodon serratus n. sp." = Ctenacodon serratus27 Marsh 1879 multituberculate
YPM 11833 (holotype right mandible); YPM 13668, USNM 2688 (mandibles)
"Ctenacodon nanus n. sp." = Ctenacodon serratus14 Marsh 1879 multituberculate
YPM 11832 (holotype left mandible)
Osteichthyes
 Therapsida -
"Tinodon bellus n. sp." = Tinodon bellus16, "Amphidon aequicrurius n. sp." = Tinodon bellus23, "Tinodon lepidus n. sp." = Tinodon bellus28, "Menacodon rarus n. gen. n. sp." = Tinodon bellus16
"Tinodon bellus n. sp." = Tinodon bellus16 Marsh 1879 mammaliaform
YPM 11843 (holotype right mandible), 13644 (left mandible) listed by Simpson 1929
"Amphidon aequicrurius n. sp." = Tinodon bellus23 Marsh 1879 mammaliaform
YPM 13639 (holotype partial right maxilla)
"Tinodon lepidus n. sp." = Tinodon bellus28 Marsh 1879 mammaliaform
YPM 11845 (holotype left mandible)
"Menacodon rarus n. gen. n. sp." = Tinodon bellus16 Marsh 1879 mammaliaform
USNM 2131 (holotype left mandible)
unclassified
  -
Charophyta indet.2 Migula 1890