PBDB Collection

Eckfeld Maar (NHMN 2000 collection) (Eocene of Germany)

Where: Eifel, Germany (50.1° N, 6.8° E: paleocoordinates 47.0° N, 2.9° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Eifel Formation, MP 13 (48.6 - 40.4 Ma)

• "MP 13... recently dated at 44.3 +/- 0.4 million years BP (Mertz et al. 2000)" (Franzen 2004). Insects come mostly from horizons near marker horizon HT & KaLH.

• bed-level stratigraphic resolution

Environment/lithology: crater lake; siltstone and shale

• "a small freshwater [crater] lake a few hundred meters in diameter" (Franzen 2004)

• ...laminitic "oilshales" and siltstones... with frequent turbiditic intercalations... plant macrofossils are ... distributed throughout the sequence under investigation. ... coarse and robust plant debris such as wood fragments and large fruits/seeds is concentrated in the turbiditic layers. (Wilde and Frankenhauser 1998).

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression, original carbon

Collected in 2000

Collection methods: quarrying, surface (in situ),

• Repository: Naturhistorisches Museum Mainz

Primary reference: G. M. Dlussky, T. Wappler, and S. Wedmann. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa 2031:1-20 [M. Clapham/M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 138912: authorized by Matthew Clapham, entered by Matthew Clapham on 01.02.2013, edited by Jered Karr and Jenna Sandlin

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Diptera - Limoniidae	Dicranoptycha cf. megaphallus⁵ Dicranoptycha cf. megaphallus⁵ Alexander 1926 crane fly NHMM PE 2000/843
Odonata - Megapodagrionidae	"Euarchistigma superstes n. sp." = Eckfeldia superstes⁶ "Euarchistigma superstes n. sp." = Eckfeldia superstes⁶ Wappler 2003 flatwing NHMM PE 2000/44a+b
Odonata -	Lestinoidea indet.⁶ Lestinoidea indet.⁶ Calvert 1901 damselfly NHMM PE2000/545a,b LS
Odonata - Priscalestidae	Priscalestes germanica n. gen. n. sp.¹¹ Priscalestes germanica n. gen. n. sp.¹¹ Wappler and Petrulevičius 2007 damselfly PE 2000/45a+b
Coleoptera - Scarabaeidae	Aphodiinae "indet. 1"⁶, Aphodiinae "indet. 2"⁶, Aphodiinae "indet. 3"⁶, Rutelinae indet.⁶ Aphodiinae "indet. 1"⁶ Leach 1815 scarab beetle NHMM PE2000/578 Aphodiinae "indet. 2"⁶ Leach 1815 scarab beetle NHMM PE2000/605 Aphodiinae "indet. 3"⁶ Leach 1815 scarab beetle NHMM PE2000/593, 592 Rutelinae indet.⁶ MacLeay 1819 scarab beetle NHMM PE2000/591, 590
Coleoptera - Lucanidae	Lucanidae indet.⁶ Lucanidae indet.⁶ Latreille 1804 stag beetle NHMM PE2000/1176, 1175
Coleoptera - Staphylinidae	Pselaphinae indet.⁶ Pselaphinae indet.⁶ Latreille 1802 rove beetle NHMM PE2000/987, 989, 988, 1984
Coleoptera - Dermestidae	Eckfeldattagenus eocenicus n. gen. n. sp.² Eckfeldattagenus eocenicus n. gen. n. sp.² Háva and Wappler 2014 skin beetle NHMM PE2000/995, 996, 997
Coleoptera - Throscidae	Throscidae indet.⁶ Throscidae indet.⁶ Laporte 1840 beetle NHMM PE2000/353
Coleoptera - Elateridae	? Lacon sp.⁶, Macropunctum densipunctum n. sp.⁶, Lanelater nicoleae n. sp.⁶, ? Athous sp.⁶ ? Lacon sp.⁶ Laporte 1838 click beetle NHMM PE2000/345 Macropunctum densipunctum n. sp.⁶ Wappler 2003 click beetle NHMM PE2000/955, 744 stratigraphischer Bereich ca. 30 cm unter Leithorizont KaLH. Lanelater nicoleae n. sp.⁶ Wappler 2003 click beetle NHMM PE2000/349 stratigraphischer Bereich 64 cm unter Leithorizont S8. ? Athous sp.⁶ Eschscholtz 1829 click beetle NHMM PE2000/2213, 347
Coleoptera - Buprestidae	? Agrilini "indet. 1"⁶, ? Agrilini "indet. 2"⁶, "Coroebini "indet. 2"" = Coraebini⁶, "Coroebini "indet. 3"" = Coraebini⁶, "Coroebini "indet. 5"" = Coraebini⁶, Buprestinae "indet. 1"⁶, Buprestinae "indet. 2"⁶, Buprestinae "indet. 3"⁶, Buprestinae "indet. 4"⁶, Buprestis sp.⁶, Julodinae indet.⁶ ? Agrilini "indet. 1"⁶ Laporte 1835 jewel beetle NHMM PE2000/668, 667 ? Agrilini "indet. 2"⁶ Laporte 1835 jewel beetle NHMM PE2000/666 "Coroebini "indet. 2"" = Coraebini⁶ Bedel 1921 jewel beetle NHMM PE2000/662, 661 "Coroebini "indet. 3"" = Coraebini⁶ Bedel 1921 jewel beetle NHMM PE2000/663 "Coroebini "indet. 5"" = Coraebini⁶ Bedel 1921 jewel beetle NHMM PE2000/664 Buprestinae "indet. 1"⁶ Leach 1815 jewel beetle NHMM PE2000/842 Buprestinae "indet. 2"⁶ Leach 1815 jewel beetle NHMM PE2000/708, 698 Buprestinae "indet. 3"⁶ Leach 1815 jewel beetle NHMM PE2000/692 a+b Buprestinae "indet. 4"⁶ Leach 1815 jewel beetle NHMM PE2000/665 Buprestis sp.⁶ Linnaeus 1758 jewel beetle NHMM PE2000/1224, 1225, 691, 690, 689 Julodinae indet.⁶ Lacordaire 1857 jewel beetle NHMM PE2000/839, 671
Coleoptera - Curculionidae	Curculionini indet.⁶ Curculionini indet.⁶ Latreille 1802 snout beetle NHMM PE2000/1493, 1492
Coleoptera - Brentidae	Brentinae "indet. 2"⁶, ? Trachelizini "indet. 1"⁶, Eckfelderolispa manderschieta n. sp.³ Brentinae "indet. 2"⁶ Billberg 1820 weevil NHMM PE2000/1248 ? Trachelizini "indet. 1"⁶ Lacordaire 1865 weevil NHMM PE2000/1279, 1247, 1246 Eckfelderolispa manderschieta n. sp.³ Legalov and Wappler 2019 weevil NHMM PE 2000/2166
Coleoptera - Cerambycidae	Cerambycidae "indet. 1"⁶ Cerambycidae "indet. 1"⁶ Latreille 1802 longhorned beetle NHMM PE2000/1186
Coleoptera - Chrysomelidae	Lamprosomatinae indet.⁶, Chrysomelinae indet.⁶, Cassidinae "indet. 2"⁶, Donaciinae indet.⁶ Lamprosomatinae indet.⁶ Lacordaire 1848 leaf beetle NHMM PE2000/474 Chrysomelinae indet.⁶ Latreille 1802 leaf beetle NHMM 2000 Cassidinae "indet. 2"⁶ Gyllenhal 1813 tortoise beetle NHMM 2000/476 Donaciinae indet.⁶ Kirby 1837 leaf beetle NHMM PE2000/477
Coleoptera - Cleridae	Cleridae indet.⁶ Cleridae indet.⁶ Latreille 1802 checkered beetle NHMM PE2000/991, 998
Coleoptera - Lophocateridae	Ancyrona eocenica n. sp.⁴ Ancyrona eocenica n. sp.⁴ Schmied et al. 2009 beetle NHMM 2000/475
Coleoptera - Tenebrionidae	Tenebrionidae indet.⁶ Tenebrionidae indet.⁶ Latreille 1802 darkling beetle NHMM PE2000/1179, 713
Coleoptera - Zopheridae	"Colydiidae indet." = Zopheridae⁶ "Colydiidae indet." = Zopheridae⁶ Solier 1834 ironclad beetle NHMM PE2000/992, 993, 994
Coleoptera - Cupedidae	Cupedidae "indet. 1"⁶, Cupedidae "indet. 2"⁶, "? Tenomerga eckfeldense" = Cupes eckfeldensis⁶ Cupedidae "indet. 1"⁶ Laporte 1836 reticulated beetle NHMM PE2000/1145 a+b Cupedidae "indet. 2"⁶ Laporte 1836 reticulated beetle NHMM PE2000/1146 a+b "? Tenomerga eckfeldense" = Cupes eckfeldensis⁶ Tröster 1993 reticulated beetle NHMM PE2000/1173, 1138, 1139, 1140, 1141, 1142, 1143, 1144
Coleoptera - Dytiscidae	? Dytiscidae indet.⁶ ? Dytiscidae indet.⁶ Leach 1815 predaceous diving beetle NHMM PE2000/990
Coleoptera - Carabidae	Carabidae "indet. 1"⁶, Carabidae "indet. 3"⁶, Pleuarthropterus (Blaticarthropterus) nonsucineus n. sp.⁶ Carabidae "indet. 1"⁶ Latreille 1802 ground beetle NHMM PE2000?1076 a+b Carabidae "indet. 3"⁶ Latreille 1802 ground beetle NHMM PE2000/1078 a+b Pleuarthropterus (Blaticarthropterus) nonsucineus n. sp.⁶ Wappler 2003 ant nest beetle NHMM PE2000/544; stratigraphischer Bereich ca. 24 cm unter Leithorizont S8.
Hymenoptera - Crabronidae	cf. Crabronidae indet.⁶ cf. Crabronidae indet.⁶ Latreille 1802 apoid wasp NHMM PE2000/865
Hymenoptera - Apidae	Protobombus "sp. 1"⁸, "Protobombus pristinus" = Protobombus (Thnetobombus) pristinus⁸, Electrapis "sp.1"⁸, Electrapis "sp.2"⁸, Electrapis "sp.3"⁸, "Electrapis micheneri" = Electrapis (Mesomelissa) micheneri⁸, "Electrapis electrapoides" = Electrapis (Eckfeldapis) electrapoides⁸, "Electrapis prolata" = Electrapis (Eckfeldapis) prolata⁸ Protobombus "sp. 1"⁸ Cockerell 1908 bee NHMM PE2000/851a,b LS "Protobombus pristinus" = Protobombus (Thnetobombus) pristinus⁸ Wappler and Engel 2003 bee NHMM PE2000/863a,b LS Electrapis "sp.1"⁸ Cockerell 1908 bee NHMM PE2000/853a,b LS Electrapis "sp.2"⁸ Cockerell 1908 bee NHMM PE2000/846a,b LS Electrapis "sp.3"⁸ Cockerell 1908 bee NHMM PE2000/849a,b LS "Electrapis micheneri" = Electrapis (Mesomelissa) micheneri⁸ Wappler and Engel 2003 bee NHMM PE2000/852a,b LS "Electrapis electrapoides" = Electrapis (Eckfeldapis) electrapoides⁸ Lutz 1993 bee PE2000/850 LS "Electrapis prolata" = Electrapis (Eckfeldapis) prolata⁸ Wappler and Engel 2003 bee NHMM PE2000/847 LS, PE2000/848a,b LS
Hymenoptera - Formicidae	Oecophylla eckfeldiana n. sp.¹, Gesomyrmex flavescens n. sp. Oecophylla eckfeldiana n. sp.¹ Dlussky et al. 2008 weaver ant NHMM PE-2000/8, 1948, 1947, 1951, 9, 11, 12 Gesomyrmex flavescens n. sp. Dlussky et al. 2009 ant PE2000/14
Dicondylia -	Blattaria indet.⁶ Blattaria indet.⁶ Latreille 1810 cockroach NHMM PE2000/1067 a+b LS, PE2000/1066 a+b LS
Blattodea - Blattidae	Blattidae indet.⁶ Blattidae indet.⁶ Latreille 1810 cockroach NHMM PE2000/1065 ab
Blattodea - Blattellidae	Blattellidae "indet. 1"⁶, Blattellidae "indet. 2"⁶ Blattellidae "indet. 1"⁶ Karny 1908 cockroach NHMM PE 2000/1070 Blattellidae "indet. 2"⁶ Karny 1908 cockroach NHMM PE 2000/1069, PE2000/1068
Dermaptera -	Neodermaptera "indet. 2"⁹ Neodermaptera "indet. 2"⁹ Engel 2003 earwig NHMM PE 2000/1062-LS
Dermaptera - Forficulidae	Forficula kotejai n. sp.⁹ Forficula kotejai n. sp.⁹ Wappler et al. 2005 common earwig NHMM PE2000/1061 LS, PE2000/1063 a+b LS.
Hemiptera -	Membracoidea indet.⁶, Cercopoidea "indet. 1"⁶, Cercopoidea "indet. 2"⁶, Cercopoidea "indet. 3"⁶ Membracoidea indet.⁶ Rafinesque 1815 true bug NHMM PE2000/836 Cercopoidea "indet. 1"⁶ Leach 1815 froghopper NHMM PE2000/824, 825, 1983 Cercopoidea "indet. 2"⁶ Leach 1815 froghopper NHMM PE2000/826, 827, 835, 1039, 1983 Cercopoidea "indet. 3"⁶ Leach 1815 froghopper NHMM PE2000/828 a+b
Hemiptera - Gerridae	Lutetiabates eckfeldensis⁶, Gerris sp.⁷ Lutetiabates eckfeldensis⁶ Wappler and Andersen 2004 water strider NHMM PE2000/1, 542, 543, 823 Gerris sp.⁷ Fabricius 1794 water strider NHMM PE2000/2-LS
Hemiptera - Aradidae	Mezira eckfeldensis¹⁰, Mezira crassifemur n. sp.¹⁰ Mezira eckfeldensis¹⁰ Wappler and Heiss 2006 flat bug NHMM PE2000/1049, 1064 Mezira crassifemur n. sp.¹⁰ Wappler and Heiss 2006 flat bug NHMM PE 2000/1048, 1050
Hemiptera - Lygaeidae	Lygaeidae "indet. 1"⁶, Lygaeidae "indet. 2"⁶ Lygaeidae "indet. 1"⁶ Schilling 1829 milkweed bug NHMM PE2000/1047 LS Lygaeidae "indet. 2"⁶ Schilling 1829 milkweed bug NHMM PE2000/1052 LS
Hemiptera - Pentatomidae	Pentatomidae "indet. 1"⁶ Pentatomidae "indet. 1"⁶ Leach 1815 stink bug NHMM PE2000/1046 a+b
Hemiptera - Acanthosomatidae	Acanthosomatidae indet.⁶ Acanthosomatidae indet.⁶ Signoret 1863 shield bug NHMM PE2000/1051 a+b
Hemiptera - Fulgoridae	Lycorma sp.⁶ Lycorma sp.⁶ Koch 1882 fulgorid planthopper NHMM PE2000/831 a+b LS
Hemiptera - Ricaniidae	Ricaniidae "indet. 1"⁶, Ricaniidae "indet. 2"⁶ Ricaniidae "indet. 1"⁶ Amyot and Audinet-Serville 1843 planthopper NHMM PE2000/832, 833 a+b LS Ricaniidae "indet. 2"⁶ Amyot and Audinet-Serville 1843 planthopper NHMM PE2000/829 a+b, PE2000/830 LS

¹Dlussky et al. 2008; ²Háva and Wappler 2014; ³Legalov and Wappler 2019; ⁴Schmied et al. 2009; ⁵Wappler 2002; ⁶Wappler 2003; ⁷Wappler and Andersen 2004; ⁸Wappler and Engel 2003; ⁹Wappler et al. 2005; ¹⁰Wappler and Heiss 2006; ¹¹Wappler and Petrulevičius 2007