PBDB Collection

Keeline ( of the United States)

Also known as Quarry Z

Where: Niobrara County, Wyoming (43.0° N, 104.5° W: paleocoordinates 43.9° N, 99.4° W)

• coordinate based on political unit

• small collection-level geographic resolution

When: Harrison Formation, Harrisonian (24.8 - 18.5 Ma)

• said to be late Arikareean

Environment/lithology: terrestrial

Primary reference: J. H. Wahlert and R. A. Souza. 1988. Skull morphology of Gregorymys and relationships of the Entoptychinae (Rodentia, Geomyidae). American Museum Novitates 2922:1-13 [J. Alroy/J. Alroy/J. Marcot]more details

PaleoDB collection 17794: authorized by John Alroy, entered by John Alroy on 07.06.1994, edited by Jonathan Marcot

Creative Commons license: CC BY (attribution)

Taxonomic list

• on the Pine Ridge Escarpment according to Schultz and Falkenbach 1950
virtually all Keeline records appear to be from a single F:AM "North of Keeline" locality

Show authors, comments, and common names

Mammalia
Rodentia - Entoptychidae	Gregorymys formosus Gregorymys formosus Matthew 1907 rodent
Carnivora - Canidae	Enhydrocyon crassidens¹⁵, Paraenhydrocyon wallovianus¹⁵, Paraenhydrocyon josephi¹⁵, Leptocyon gregorii¹⁴, Cormocyon haydeni¹⁶ Enhydrocyon crassidens¹⁵ Matthew 1907 canid North of Keeline Paraenhydrocyon wallovianus¹⁵ Cope 1881 canid North of Keeline, "Lower or Upper Arikaree Group" Paraenhydrocyon josephi¹⁵ Cope 1881 canid North of Keeline Leptocyon gregorii¹⁴ Matthew 1907 canine north of Keeline, middle brown sand Cormocyon haydeni¹⁶ Wang et al. 1999 bone-crushing dog "north of Keeline"
Carnivora - Mustelidae	Zodiolestes sp.⁶, Megalictis ferox⁶ Zodiolestes sp.⁶ Riggs 1942 mustelid carnivore Megalictis ferox⁶ Matthew 1907 mustelid carnivore "North of Keeline (Quarry Z)... Harrison"
Carnivora - Amphicyonidae	Mammacyon ferocior n. sp.⁵ Mammacyon ferocior n. sp.⁵ Hunt 2011 bear-dog "Arikaree Formation, N of Keeline"
Perissodactyla - Equidae	Kalobatippus avus⁷ Kalobatippus avus⁷ Schlaikjer 1935 anchitheriine horse
Perissodactyla - Rhinocerotidae	Diceratherium armatum², Diceratherium annectens⁹ Diceratherium armatum² Marsh 1875 rhinoceros "north of Keeline, Niobrara County, Wyoming" Diceratherium annectens⁹ Marsh 1873 rhinoceros
Perissodactyla - Tapiridae	Nexuotapirus marslandensis¹ Nexuotapirus marslandensis¹ Schoch 1984 tapir "Lusk area, north of Keeline, Niobrara County"
Artiodactyla - Merycoidodontidae	Phenacocoelus typus¹², Desmatochoerus curvidens¹³, "Promerycochoerus carrikeri" = Promerycochoerus carrikeri¹¹, "Promerycochoerus (Parapromerycochoerus) barbouri n. sp." = Promerycochoerus carrikeri¹¹, "Paramerychyus harrisonensis" = Paramerychyus harrisonensis¹⁰, "Merychyus siouxensis" = Paramerychyus harrisonensis¹⁰, "Merychyus crabilli" = Merychyus calaminthus¹⁰ Phenacocoelus typus¹² Peterson 1907 oreodont Harrison Fm. N of Keeline Desmatochoerus curvidens¹³ Thorpe 1921 oreodont as ssp. Desmatochoerus curvidens gregoryi; N of Keeline, Niobrara Co. "Promerycochoerus carrikeri" = Promerycochoerus carrikeri¹¹ Peterson 1907 oreodont "N. of Keeline" and "Quarry 'Z,' N. of Keeline" "Promerycochoerus (Parapromerycochoerus) barbouri n. sp." = Promerycochoerus carrikeri¹¹ Peterson 1907 oreodont Harrison Fm. N of Keeline "Paramerychyus harrisonensis" = Paramerychyus harrisonensis¹⁰ Peterson 1907 oreodont "Z Quarry" N of Keeline "Merychyus siouxensis" = Paramerychyus harrisonensis¹⁰ Peterson 1907 oreodont "N. of Keeline" and "8 mi. S.W. of Keeline" in Niobrara Co. "Merychyus crabilli" = Merychyus calaminthus¹⁰ Jahns 1940 oreodont "N. of Keeline" in Niobrara Co.
Artiodactyla - Hypertragulidae	Nanotragulus ordinatus³ Nanotragulus ordinatus³ Matthew 1907 ruminant "Keeline, Wyoming"
Artiodactyla - Protoceratidae	Protoceras neatodelpha n. sp.⁸ Protoceras neatodelpha n. sp.⁸ Patton and Taylor 1973 protoceratid N of Keeline, Niobrara Co., in Harrison Fm. "equivalent"
Artiodactyla - Camelidae	Stenomylus keelinensis⁴ Stenomylus keelinensis⁴ Frick and Taylor 1968 camel N of Keeline; S. gracilis of Frick 1937

¹Albright 1998; ²Albright 1999; ³Frick 1937; ⁴Frick and Taylor 1968; ⁵Hunt 2011; ⁶Hunt and Skolnick 1996; ⁷MacFadden 2001; ⁸Patton and Taylor 1973; ⁹Prothero 2005; ¹⁰Schultz and Falkenbach 1947; ¹¹Schultz and Falkenbach 1949; ¹²Schultz and Falkenbach 1950; ¹³Schultz and Falkenbach 1954; ¹⁴Tedford et al. 2009; ¹⁵Wang 1994; ¹⁶Wang et al. 1999