Bembridge Marls (A'Court Smith collection) (Eocene of the United Kingdom)

Also known as ETE Locality 797

Where: England, United Kingdom (50.7° N, 1.4° W: paleocoordinates 49.2° N, 5.2° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Bembridge Marls Member (Bouldnor Formation), Priabonian (38.0 - 33.9 Ma)

• Upper part of 13r chron just above 1n interval

Environment/lithology: lagoonal or restricted shallow subtidal; lithified, argillaceous lime mudstone

• The Bembridge Marls were deposited in fresh-brackish water lakes and lagoons

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by E. J. A'Court Smith; reposited in the BMNH

• Specimens kept in British Natural History Museum

Primary reference: E. A. Jarzembowski. 1980. Fossil Insects from the Bembridge Marls, Palaeogene of the Isle of Wight, Southern England. Bulletin of the British Museum of Natural History (Geology) 33(4):237-293 [C. Labandeira/C. Labandeira/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 21976: authorized by Conrad Labandeira, entered by Conrad Labandeira on 08.06.1998, edited by Jenna Sandlin, Jered Karr and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Diptera - Psychodidae
Wightipsychoda leucospila12
Wightipsychoda leucospila12 Cockerell 1921 moth fly
NHMUK In.17204
 Diptera - Chironomidae
Chironomidae indet.12, Chironomini indet.12, Orthocladiinae indet.12
Chironomidae indet.12 Macquart 1838 non biting midge
NHMUK In.17470
Chironomini indet.12 Macquart 1838 non biting midge
NHMUK In.17388
Orthocladiinae indet.12 Lenz 1921 non biting midge
NHMUK In.17390(1)
 Diptera - Cylindrotomidae
Cyttaromyia rossi n. sp.12
Cyttaromyia rossi n. sp.12 Krzemiński 2019 long bodied crane fly
CAMSM X.50140.14
 Diptera - Limoniidae
Cheilotrichia (Empeda) szwedoi n. sp.12, "Gymnastes fasciatipennis" = Gymnastes (Paragymnastes) fasciatipennis12, "Gonomyia (Gonomyia) indecisa" = Gonomyia (Gonomyia) indecisa12, "Gonomyia lutescens" = Gonomyia (Gonomyia) indecisa4, Dicranomyia (Dicranomyia) brodiei12, Trentepohlia (Mongoma) pallescens12, "Mongoma cruciferella" = Trentepohlia (Mongoma) cruciferella4, "Limnobia spilota n. sp." = Limonia spilota4, Limnophila (Limnophila) deleta12, Limnophila (Limnophila) cyclospila12, Pseudolimnophila (Pseudolimnophila) disjunctula12
Cheilotrichia (Empeda) szwedoi n. sp.12 Krzemiński 2019 crane fly
NHMUK In.17082
"Gymnastes fasciatipennis" = Gymnastes (Paragymnastes) fasciatipennis12 Cockerell 1921 crane fly
NHMUK In.17494
"Gonomyia (Gonomyia) indecisa" = Gonomyia (Gonomyia) indecisa12 Cockerell and Haines 1921 crane fly
NHMUK In.17248
"Gonomyia lutescens" = Gonomyia (Gonomyia) indecisa4 Cockerell and Haines 1921 crane fly
BMNH In.17099
Dicranomyia (Dicranomyia) brodiei12 Cockerell 1917 crane fly
NHMUK In.17486
Trentepohlia (Mongoma) pallescens12 Cockerell 1921 crane fly
NHMUK In.17117/I.9051 (Brodie collection), 17123
"Mongoma cruciferella" = Trentepohlia (Mongoma) cruciferella4 Cockerell 1917 crane fly
BMNH I.17259; In.17116, 17130, 17264/I.9200 (Brodie collection) in Krzeminski et al. 2020
"Limnobia spilota n. sp." = Limonia spilota4 Cockerell 1921 crane fly
BMNH In.17114
Limnophila (Limnophila) deleta12 Cockerell 1921 crane fly
NHMUK In.17115, 17171, 24414, 17484/In.64127 (Hooley collection)
Limnophila (Limnophila) cyclospila12 Cockerell and Haines 1921 crane fly
NHMUK In.17491
Pseudolimnophila (Pseudolimnophila) disjunctula12 Cockerell and Haines 1921 crane fly
NHMUK In.17383
 Diptera - Tipulidae
Leptotarsus (Longurio) wegiereki n. sp.12, "Tipula callarche n. sp." = Leptotarsus (Macromastix) cladoptera4, Tipula gardneri12, "Megistocera gurnetensis n. sp." = Brachypremna gurnetensis4
Leptotarsus (Longurio) wegiereki n. sp.12 Krzemiński 2019 crane fly
NHMUK In.17493, CAMSM X.50140, X.50140.89a
"Tipula callarche n. sp." = Leptotarsus (Macromastix) cladoptera4 Cockerell and Haines 1921 crane fly
BMNH In. 17381
Tipula gardneri12 Cockerell 1917 crane fly
NHMUK In.17429
"Megistocera gurnetensis n. sp." = Brachypremna gurnetensis4 Cockerell 1921 crane fly
BMNH In.17090
 Diptera - Anisopodidae
"Rhyphus hooleyi" = Sylvicola hooleyi4
"Rhyphus hooleyi" = Sylvicola hooleyi4 Cockerell 1921 wood gnat
BMNH In.17136 (counterpart I.9063 in Brodie coll)
 Diptera - Bibionidae
Plecia acourti n. sp.4
Plecia acourti n. sp.4 Cockerell 1921 love bug
BMNH In.17098
 Diptera - Stratiomyidae
Odontomyia brodiei12
Odontomyia brodiei12 Cockerell 1915 soldier fly
NHMUK In.17221, 17253, 17356, 17478
 Diptera - Bombyliidae
Systropus acourti n. sp.4
Systropus acourti n. sp.4 Cockerell 1921 bee fly
BMNH I.17208/17293
 Diptera - Ceratopogonidae
Stilobezzia sp.12
Stilobezzia sp.12 Kieffer 1911 biting midge
NHMUK In.17390(2), 17471
 Diptera - Dixidae
Dixella priscula12
Dixella priscula12 Cockerell 1921 meniscus midge
NHMUK In.17402
 Diptera - Culicidae
"Aedes protolepis" = Ochlerotatus protolepis9, Culex protorhinus9
"Aedes protolepis" = Ochlerotatus protolepis9 Cockerell 1915 mosquito
BMNH In. 17069, 17147, 17148, 17164, 20561
Culex protorhinus9 Cockerell 1915 mosquito
BMNH In. 17138, 17146
 Blattodea - Rhinotermitidae
Reticulitermes sp.
Reticulitermes sp. Holmgren 1913 termite
 Blattodea - Mastotermitidae
"Mastotermes anglicus n. sp." = Mastotermes anglicus23, "Mastotermes batheri n. sp." = Mastotermes anglicus23
"Mastotermes anglicus n. sp." = Mastotermes anglicus23 von Rosen 1913 termite
BMNH I 15037 (21 in von Rosen, 1913)
"Mastotermes batheri n. sp." = Mastotermes anglicus23 von Rosen 1913 termite
 Blattodea - Ectobiidae
Ectobiidae "indet. 6"17, Phyllodromica protosardea n. sp.17
Ectobiidae "indet. 6"17 Brunner von Wattenwyl 1865 cockroach
NHMUK In.17337
Phyllodromica protosardea n. sp.17 Ross 2019 cockroach
NHMUK I.15039/In.17142(2)
 Mantodea -
Protohierodula crabbi n. gen. n. sp.17
Protohierodula crabbi n. gen. n. sp.17 Ross 2019 mantis
NHMUK In.17462
 Plecoptera - Leuctridae
"Nemoura priscula n. sp." = Leuctra priscula7
"Nemoura priscula n. sp." = Leuctra priscula7 Cockerell 1922 rolled-winged stonefly
NHML In.17498
 Orthoptera - Acrididae
Proschistocerca oligocaenica24
Proschistocerca oligocaenica24 Zeuner 1937 short-horned grasshopper
BMNH In.17339, 17342, 17406, 17407, 20527; 17346, 17350, 20533 in Gorochov 2019
 Orthoptera - Gryllidae
Ontogryllus rossi n. gen. n. sp.10
Ontogryllus rossi n. gen. n. sp.10 Gorochov 2019 field cricket
NHMUK In.25806
 Orthoptera - Gryllotalpidae
Pterotriamescaptor primus10
Pterotriamescaptor primus10 Cockerell 1921 mole cricket
NHMUK In.17460a,b, 17461
 Orthoptera - Tettigoniidae
Archepseudophylla melanospila15
Archepseudophylla melanospila15 Cockerell 1921 katydid
BMNH In.25803/In.17326
 Psocodea -
Wightipsocus intasi2, Awightipsocus megaloi n. sp.2
Wightipsocus intasi2 Azar 2014 bark louse
BMNH In.17088
Awightipsocus megaloi n. sp.2 Azar 2014 bark louse
BMNH In.17187
 Coleoptera - Dytiscidae
Hydroporinae indet.11
Hydroporinae indet.11 Aubé 1836 predaceous diving beetle
NHMUK In.17437 (?Oreodytes sp. or ?Hydrovatus sp.)
 Coleoptera - Ptilodactylidae
Paralichas striatopunctatus n. sp.11
Paralichas striatopunctatus n. sp.11 Kirejtshuk 2019 beetle
NHMUK In.17449
 Coleoptera - Scirtidae
Scirtinae indet.11, Scirtes calcariferens11, Contacyphon involutus11
Scirtinae indet.11 Fleming 1821 marsh beetle
NHMUK In.17432
Scirtes calcariferens11 Kirejtshuk and Ponomarenko 2019 marsh beetle
NHMUK In.17437(2)
Contacyphon involutus11 Kirejtshuk and Ponomarenko 2019 marsh beetle
NHMUK In.17435
 Coleoptera - Chrysomelidae
Plateumaris (Plateumaris) aff. wightensis11
Plateumaris (Plateumaris) aff. wightensis11 Kurochkin and Kirejtshuk 2019 reed leaf beetle
NHMUK In.2580, 20552
 Coleoptera - Nitidulidae
Prometopia (Prometopia) europaea n. sp.11
Prometopia (Prometopia) europaea n. sp.11 Kirejtshuk and Kurochkin 2019 sap beetle
NHMUK In.17438
 Coleoptera - Scarabaeidae
Aphodius vectis n. sp.11, Pentodon dorcus n. sp.11
Aphodius vectis n. sp.11 Krell 2019 scarab beetle
NHMUK In.17436
Pentodon dorcus n. sp.11 Krell 2019 rhinoceras beetle
NHMUK In.17431
 Coleoptera - Staphylinidae
Staphylinidae indet.11, Pselaphinae indet.11
Staphylinidae indet.11 Latreille 1802 rove beetle
NHMUK In.17384
Pselaphinae indet.11 Latreille 1802 rove beetle
CAMSM X.50140.206
 Coleoptera - Hydrophilidae
Hydrochara woodwardi n. sp.11, Berosus barclayi11
Hydrochara woodwardi n. sp.11 Ponomarenko and Soriano 2019 water scavenger beetle
NHMUK In.17424
Berosus barclayi11 Ponomarenko and Soriano 2019 water scavenger beetle
NHMUK In.17426
 Coleoptera - Eucinetidae
Eucinetus nikolaevae n. sp.11
Eucinetus nikolaevae n. sp.11 Kirejtshuk and Ponomarenko 2019 plate-thigh beetle
NHMUK In.17286
 Dicondylia -
Neuroptera indet.
Neuroptera indet. Linnaeus 1758 lacewing
NHML In.20557
 Neuroptera - Hemerobiidae
Hemerobiidae "sp.B", Hemerobiidae "sp.C", ? Neuronema "sp. A"
Hemerobiidae "sp.B" Latreille 1802 brown lacewing
NHML In.17173
Hemerobiidae "sp.C" Latreille 1802 brown lacewing
NHML In.17445
? Neuronema "sp. A" McLachlan 1869 brown lacewing
NHML In.17404
 Neuroptera - Sisyridae
Paleosisyra sp.16
Paleosisyra sp.16 Nel et al. 2003 spongillafly
NHML In.17287
 Trichoptera - Beraeidae
Beraeodes vectensis21
Beraeodes vectensis21 Cockerell 1921 caddisfly
BMNH In.17290, 17351, 17393, 17397, 17499
 Dicondylia -
Lepidoptera "sp.D"
Lepidoptera "sp.D" Linnaeus 1758 moth
NHML In.17392
 Lepidoptera - Micropterigidae
Micropterix anglica n. sp.
Micropterix anglica n. sp. Jarzembowski 1980 moth
NHML In.17411
 Lepidoptera -
Lithopsyche antiqua n. gen. n. sp.3
Lithopsyche antiqua n. gen. n. sp.3 Butler 1889 butterfly
NHML In.19984 I.10369
 Hymenoptera - Ichneumonidae
Ichneumonidae indet.1, Coleocentrus gurnetensis n. sp.5
Ichneumonidae indet.1 Latreille 1802 ichneumon wasp
BMNH In.17430
Coleocentrus gurnetensis n. sp.5 Cockerell 1921 ichneumon wasp
NHML In. 17079
 Hymenoptera - Braconidae
Dacnusites sepultus n. gen. n. sp.5, Semionis wightensis n. sp.1, Homolobus rasnitsyni n. sp.1, Ascogaster (Ascogaster) pygmaea n. sp.1, Eubazus flavistigma n. sp.1, Bassus magnareola n. sp.1, ? Hellenius kozlovi n. sp.1, Bembracon acourtsmithi n. gen. n. sp.1
Dacnusites sepultus n. gen. n. sp.5 Cockerell 1921 wasp
NHML In. 17176
Semionis wightensis n. sp.1 Belokobylskij 2014 wasp
BMNH In.17081
Homolobus rasnitsyni n. sp.1 Belokobylskij 2014 wasp
BMNH In.20545
Ascogaster (Ascogaster) pygmaea n. sp.1 Belokobylskij 2014 wasp
BMNH In.17379
Eubazus flavistigma n. sp.1 Belokobylskij 2014 wasp
BMNH In.17244
Bassus magnareola n. sp.1 Belokobylskij 2014 wasp
BMNH In.17127
? Hellenius kozlovi n. sp.1 Belokobylskij 2014 wasp
BMNH In.17298
Bembracon acourtsmithi n. gen. n. sp.1 Belokobylskij 2014 wasp
BMNH In.17224
 Hymenoptera - Apidae
Oligobombus cuspidatus n. gen. n. sp.1
Oligobombus cuspidatus n. gen. n. sp.1 Antropov 2014 bee
BMNH In.17349
 Hymenoptera - Sphecidae
"Sceliphron brevior n. sp." = Protosceliphron brevior5
"Sceliphron brevior n. sp." = Protosceliphron brevior5 Cockerell 1921 digger wasp
NHML In. 17472/24340 (Hooley collection)
 Hymenoptera - Tiphiidae
Dryophia oculata n. gen. n. sp.1
Dryophia oculata n. gen. n. sp.1 Antropov 2014 wasp
BMNH In.17086
 Hymenoptera - Vespidae
"Polybia oblita n. sp." = Protopolistes oblitus6
"Polybia oblita n. sp." = Protopolistes oblitus6 Cockerell 1921 wasp
BMNH In. 20530(17166)
 Hymenoptera - Formicidae
Emplastus hypolithus1, "Dolichoderus britannicus" = Emplastus britannicus8, "Dolichoderus ovigerus" = Emplastus britannicus8, Oecophylla megarche8, "Camponotus cockerelli" = Camponotites cockerelli1, Leucotaphus gurnetensis8
Emplastus hypolithus1 Cockerell 1915 ant
BMNH In.17311 (D. britannicus in Donisthorpe 1920)
"Dolichoderus britannicus" = Emplastus britannicus8 Cockerell 1915 ant
BMNH In.17315 (not 17311); In.17067/17074 in Antropov et al., 2014
"Dolichoderus ovigerus" = Emplastus britannicus8 Cockerell 1915 ant
BMNH In.17274
Oecophylla megarche8 Cockerell 1915 weaver ant
"Camponotus cockerelli" = Camponotites cockerelli1 Donisthorpe 1920 ant
BMNH In.17280, 17066
Leucotaphus gurnetensis8 Cockerell 1915 ant
not In.17066, 17213
 Hymenoptera -
Lithobelyta reducta n. gen. n. sp.5, "Macroteleia veterna n. sp." = Calotelea veterna5
Lithobelyta reducta n. gen. n. sp.5 Cockerell 1921 chalcid wasp
NHML In. 17091
"Macroteleia veterna n. sp." = Calotelea veterna5 Cockerell 1921 wasp
NHML In. 17262
 Hymenoptera - Vectevaniidae
Vectevania vetula n. gen. n. sp.7
Vectevania vetula n. gen. n. sp.7 Cockerell 1922 wasp
BMNH In.20535
 Hemiptera -
Aphidoidea indet.22
Aphidoidea indet.22 Latreille 1802 aphid
NHMUK In.17198, In.17210(2,3), In.17214
 Hemiptera - Aphididae
? Colopha incognita22
? Colopha incognita22 Wegierek 2019 aphid
NHMUK In.17178, 17198
 Hemiptera - Gerridae
Gerris sp.22
Gerris sp.22 Fabricius 1794 water strider
CAMSM X.50140.44 (TN94)
 Hemiptera - Cydnidae
Cydnidae indet.22
Cydnidae indet.22 Billberg 1820 burrowing bug
NHMUK In.17441b
 Hemiptera - Lygaeidae
Gurnardobaya rossi22
Gurnardobaya rossi22 Popov 2019 milkweed bug
CAMSM X.50140.71 (TN122)
 Hemiptera - Miridae
Gurnardinia herczeki n. gen. n. sp.22
Gurnardinia herczeki n. gen. n. sp.22 Popov 2019 plant bug
NHMUK In.17231/In.17232
 Hemiptera - Cicadellidae
Typhlocybinae indet.22, Teniwitta andrewrossi n. gen. n. sp.22
Typhlocybinae indet.22 Kirschbaum 1868 leafhopper
NHMUK In.17142(3)
Teniwitta andrewrossi n. gen. n. sp.22 Szwedo 2019 leafhopper
NHMUK In.20529
 Hemiptera - Aphrophoridae
Aphrophoridae "indet. 2"22, Aphrophoridae "indet. 3"22, Blenniphora skaka n. sp.22
Aphrophoridae "indet. 2"22 Amyot and Audinet-Serville 1843 froghopper
CAMSM X.50140.145 (TN203)
Aphrophoridae "indet. 3"22 Amyot and Audinet-Serville 1843 froghopper
NHMUK In.24560
Blenniphora skaka n. sp.22 Szwedo 2019 froghopper
CAMSM X.50140.90 (TN145)
 Hemiptera - Nogodinidae
Ambitaktoa stoumma n. gen. n. sp.22, Phariberea gurdonika n. gen. n. sp.22
Ambitaktoa stoumma n. gen. n. sp.22 Szwedo 2019 planthopper
NHMUK In.17282
Phariberea gurdonika n. gen. n. sp.22 Szwedo 2019 planthopper
NHMUK In.17096
 Hemiptera - Cixiidae
Margaxius sp.22
Margaxius sp.22 Szwedo 2019 cixiid planthopper
NHMUK In.17369
 Thysanoptera - Aeolothripidae
Aeolothrips jarzembowskii20, Sinaeolothrips brodiei20
Aeolothrips jarzembowskii20 Shmakov 2014 thrips
BMNH In.17210
Sinaeolothrips brodiei20 Cockerell 1917 thrips
BMNH In.17401
 Odonata - Aeshnidae
Anglogomphaeschna eocenica14, Aeschnophlebia andreasi14
Anglogomphaeschna eocenica14 Nel and Fleck 2014 darner dragonfly
BMNH In.17368, 17330
Aeschnophlebia andreasi14 Nel et al. 2005 darner dragonfly
BMNH In.17371, 17395
 Odonata - Gomphaeschnidae
? Oligoaeschna anglica14
? Oligoaeschna anglica14 Cockerell and Andrews 1916 true dragonfly
BMNH In.17353
 Odonata - Urothemistidae
Eomacrodiplax incompleta14
Eomacrodiplax incompleta14 Nel and Fleck 2014 true dragonfly
BMNH In.17333
 Odonata - Corduliidae
Corduliidae "indet. B"14, Corduliidae "indet. C"14
Corduliidae "indet. B"14 Selys-Longchamps 1850 green-eyed skimmer
BMNH In.17370/24763 (counterpart in Hooley coll.)
Corduliidae "indet. C"14 Selys-Longchamps 1850 green-eyed skimmer
BMNH In.17324/24771 (counterpart in Hooley coll.)
 Odonata - Hypolestidae
Anglohypolestes fasciata n. gen. n. sp.14
Anglohypolestes fasciata n. gen. n. sp.14 Nel and Fleck 2014 damselfly
BMNH In.17373
 Odonata - Protoneuridae
Angloprotoneura emilielacroixi n. gen. n. sp.14
Angloprotoneura emilielacroixi n. gen. n. sp.14 Nel and Fleck 2014 damselfly
BMNH In.17246
 Odonata - Lestidae
Lestes aff. regina14
Lestes aff. regina14 Theobald 1937 spread-winged damselfly
BMNH In.17408
Arachnida
 Araneae - Dictynidae
Vectaraneus yulei18
Vectaraneus yulei18 Selden 2001 spider
BMNH In. 17151
 Araneae - Araneidae
Araneidae indet.19
Araneidae indet.19 Simon 1895 orb-weaving spider
BMNH In.17177
 Araneae -
Eoatypus woodwardii n. sp.13
Eoatypus woodwardii n. sp.13 McCook 1888 spider
BMNH In. 61271 (collection not specified)
1Antropov et al. 2014; 2Azar 2014; 3Butler 1889; 4Cockerell 1921; 5Cockerell 1921; 6Cockerell 1921; 7Cockerell 1922; 8Donisthorpe 1920; 9Edwards 1923; 10Gorochov 2019; 11Kirejtshuk et al. 2019; 12Krzemiński et al. 2019; 13McCook 1888; 14Nel and Fleck 2014; 15Nel et al. 2008; 16Perkovsky and Makarkin 2015; 17Ross 2019; 18Selden 2001; 19Selden 2014; 20Shmakov 2014; 21Sukatsheva 2014; 22Szwedo et al. 2019; 23von Rosen 1913; 24Zeuner 1937