Abbey Wood (Eocene of the United Kingdom)

Also known as The Sandpit, Lessness Shell Bed, Blackheath Beds

Where: England, United Kingdom (51.4° N, 0.2° E: paleocoordinates 46.9° N, 2.3° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Abbey Wood Member (Blackheath Beds Formation), Ypresian (56.0 - 47.8 Ma)

• Blackheath Beds is an informal formation name. Lessness Shell Bed (Sparnacian) is within the lower sandy Abbey Wood Member of the Blackheath Beds.

•JA: assigned to the early Ypresian and Hooker zone PE III by Hooker 2007

• bed-level stratigraphic resolution

Environment/lithology: marine; sandstone

• from the lower sandy member

Size classes: macrofossils, mesofossils

Collection methods: bulk, chemical, sieve,

• Wrigley cites Mr. St. John Marriott (1925) as the first to publish on this locality in "British Woodlands as illustrated by Lessness Abbey Woods", which I couldn't get my hands on

Primary reference: A. Wrigley. 1931. The Lower Eocene Mollusca of Abbey Wood and of High Halstow. The Vertebrate Faunas of the English Eocene 1:110-112 [J. Alroy/E. Leckey/E. Leckey]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 38868: authorized by John Alroy, entered by Erin Leckey on 11.05.2004, edited by John Alroy, Matthew Carrano, Philip Mannion, Juan Benito Moreno and Mark Uhen

Creative Commons license: CC BY (attribution)

Taxonomic list

• Cooper doesn't specify the level at Abbey Wood from which his fossils were found. He calls the unit London Clay and gives the age as Sparnacian-Ypresian, based on the fossils.
Aves
 Procellariiformes - Marinavidae
Marinavis longirostris n. gen. n. sp.2
Marinavis longirostris n. gen. n. sp.2 Harrison and Walker 1977 bird
Reptilia
 Testudines - Trionychidae
Trionyx silvestris n. sp.11 Walker and Moody 1974 softshell turtle
almost complete skull
Mammalia
 Multituberculata - Neoplagiaulacidae
"Charlesmooria childei n. gen. n. sp." = Ectypodus childei8, Ectypodus cf. tardus6
"Charlesmooria childei n. gen. n. sp." = Ectypodus childei8 Kühne 1969 multituberculate
NHMUK M26617 - type; referred material: M15131
Ectypodus cf. tardus6 Jepsen 1940 multituberculate
NHMUK M60309
 Theriamorpha - Leptictidae
? Leptictidae indet.6 Gill 1872 eutherian
NHMUK M45247
Didelphodus sp.6 Cope 1882 eutherian
NHMUK M60071, 26140, 29733, 15124 and 44954
 Theriamorpha - Pseudorhyncocyonidae
Fordonia lawsoni7 Hooker 2013 eutherian
NHMUK.M45247 (LP3)
Leptictidium prouti n. sp.7 Hooker 2013 eutherian
NHMUK.M65382 - type (LM3)
 Cimolesta - Apatemyidae
Apatemys prouti n. sp.6 Hooker 2010 eutherian
NHMUK M51704 - type; paratypes: NHMUK M44880, 44948 and 13772
Eochiromys ? sp.4 Teilhard de Chardin 1927 eutherian
 Cimolesta - Esthonychidae
Plesiesthonyx minimus6 Baudry 1992 tillodont
NHMUK M34580, 29363 and 26787
 Chiroptera -
Chiroptera indet.6 Blumenbach 1779 bat
NHMUK M60045
 Chiroptera - Emballonuridae
"? Adapisorex anglicus n. sp." = Eppsinycteris anglica1, "Eppsinycteris anglica n. gen. n. sp." = Eppsinycteris anglica6
"? Adapisorex anglicus n. sp." = Eppsinycteris anglica1 Cooper 1932 sac-winged bat
n. sp., portion of right maxilla
"Eppsinycteris anglica n. gen. n. sp." = Eppsinycteris anglica6 Cooper 1932 sac-winged bat
NHMUK M13776 - holotye
 Insectivora -
Insectivora indet.1 Bowdich 1821 insectivoran
 Macroscelidea - Amphilemuridae
? Amphilemuridae indet.6 Hill 1953 elephant shrew
NHMUK M45500 and 45614
Macrocranion sp.6 Weitzel 1949 elephant shrew
NHMUK M44893, 44904, 51702, 45618, 44903, 44956 and 60058
Neomatronella gassoni n. sp.6 Hooker 2010 elephant shrew
NHMUK M60070 - type; paratypes: NHMUK M44953, 29365 and 51701; referred material: NHMUK M44952
 Condylarthra - Hyopsodontidae
Lessnessina packmani n. gen. n. sp.3, Hyopsodus wardi n. sp.3
Lessnessina packmani n. gen. n. sp.3 Hooker 1979 condylarth
left maxilla with P3-M3 and isolated teeth
Hyopsodus wardi n. sp.3 Hooker 1979 condylarth
jaw fragments and isolated teeth
 Acreodi - Mesonychidae
Pachyaena cf. ossifraga6 Cope 1874 condylarth
NHMUK M44984
 Placentalia -
Creodonta indet.1 Cope 1875 creodont
 Creodonta - Hyaenodontidae
Prototomus minimus10 Smith and Smith 2001 creodont
NHMUK M44936 and 45672
Prototomus girardoti10 Smith and Smith 2001 creodont
NHMUK M45651, 20212 and 63696
 Creodonta - Oxyaenidae
Oxyaena gulo6 Matthew and Granger 1915 creodont
NHMUK 13778, 15128 and 31877
Palaeonictis gigantea6 de Blainville 1842 creodont
NHMUK M44983, 15125, 45517, 63811, 60024, 15128, 60092, 29410, 60026, 29233, 29359
 Theriamorpha -
Miacis rundlei n. sp.6 Hooker 2010 placental
NHMUK M26248 - type; paratypes: NHMUK M44902, 29725, 60008, 29726, 44890, 44955, 44974, 26785, 60081, 44962 and 45245; referred material: NHMUK M45554
Uintacyon cf. rudis6 Rose 1981 placental
NHMUK M26246
 Theriamorpha - Viverravidae
Viverravus lawsoni n. sp.6 Hooker 2010 placental
NHMUK M51700 - type; paratypes: NHMUK 45487, 63196 and 44930
 Perissodactyla - Equidae
Pliolophus barnesi n. sp.6, "Hyracotherium aff. vulpiceps" = Pliolophus vulpiceps4
Pliolophus barnesi n. sp.6 Hooker 2010 horse
author notes that this is being described elsewhere
"Hyracotherium aff. vulpiceps" = Pliolophus vulpiceps4 Owen 1858 horse
 Perissodactyla - Lophiodontidae
Coryphodon eocaenicus1 odd-toed ungulate
NHMUK M13775, 83063, 29232, 44887, 29234, 83056, 34863, 21819, 44911, 20229, 29746, 26788, 29748, 63572, 15144, 29749 and 44981
 Theriamorpha - Phenacodontidae
Phenacodus lemoinei6 condylarth
NHMUK M44926, 45661, 29723 and 29724
 Artiodactyla - Diacodexeidae
Diacodexis morrisi n. sp.6 Hooker 2010 even-toed ungulate
NHMUK M83071 - type; paratypes: NHMUK M44943, 83065, 15141, 44919 and 60060; referred material: NHMUK M45485, 45647, 60794, 45515, 45619, 63702, 26897, 45638, 63697 and 63703
 Procreodi - Arctocyonidae
Arctocyonides jefferyi n. sp.6 Hooker 2010 condylarth
NHMUK M45650 - type; paratypes: NHMUK M31876, 29235 and 51526
 Theriamorpha - Nyctitheriidae
Leptacodon sp.6 Matthew and Granger 1921 placental
NHMUK M50384
? Wyonycteris sp.6 Gingerich 1987 placental
NHMUK M63198
Wyonycteris richardi6 Smith 1995 placental
NHMUK M44900, 44901 and 83068
? Remiculus sp.6 Russell 1964 placental
NHMUK M60003
 Theriamorpha - Pantolestoidae
Palaeosinopa cf. osborni6 placental
NHMUK M29761, 60665
 Placentalia -
Rodentia indet.1 Bowdich 1821 rodent
NHMUK M44899
 Rodentia -
Pseudoparamys sp.6 Michaux 1964 rodent
NHMUK M44950, 44938, 45494 and 45640
 Rodentia - Ischyromyidae
? Sparnacomys sp.6 Hartenberger 1971 rodent
NHMUK M44909
Sparnacomys georgei n. sp.6 Hooker 2010 rodent
NHMUK M29364 - type; paratype: NHMUK M26899; previously identified as Microparamys
Pantrogna cf. russelli6 Michaux 1964 rodent
NHMUK M26898 and 45643
Ailuravinae indet.6 Michaux 1968 rodent
NHMUK M44937
Ailuravus mitchelli n. sp.6 Hooker 2010 rodent
NHMUK M31909 - type; paratyps: NHMUK M45258, 63195, 45493, 44905, 45390 and 44892
cf. Meldimys sp.6 Michaux 1968 rodent
NHMUK M45391
 Primates - Notharctidae
"Protoadapis eppsi n. sp." = Cantius eppsi1
"Protoadapis eppsi n. sp." = Cantius eppsi1 Cooper 1932 primate
NHMUK M13773 - type; referred material: NHMUK M15145, 60055, 29641, 63691, 45637, 60007, 60022, 51750, 15147, 29639, 60795, 45616, 29753, 45491, 29638, 60799 and 45555
 Primates - Plesiadapidae
Platychoerops sp.6 Charlesworth 1855 primate
NHMUK M45623
 Primates - Omomyidae
Melaneremia bryanti n. gen. n. sp.5
Melaneremia bryanti n. gen. n. sp.5 Hooker 2007 tarsier
NHMUK M60001 - holotype; Teilhardina sp. of Hooker 1980, Nannopithex sp. of Collinson and Hooker 1987, and Omomyidae indet. of Hooker 1998
 Primates - Paromomyidae
Arcius hookeri n. sp.9 López-Torres and Silcox 2018 primate
NHMUK M44945
 Proteutheria - Palaeoryctidae
Didelphodontinae indet.4 Simpson 1927 eutherian
 Theriamorpha - Peradectidae
Peradectes sp.6 Matthew and Granger 1921 marsupial
NHMUK M44991
 Theriamorpha - Herpetotheriidae
Amphiperatherium "sp. 1"6 Filhol 1879 marsupial
NHMUK M60004, 63695, 60005, 44963; previously listed as Amphiperatherium brabantense
Amphiperatherium "sp. 2"6 Filhol 1879 marsupial
NHMUK M60002; previously listed as Amphiperatherium maximum
Osteichthyes
 Acipenseriformes - Acipenseridae
Acipenser sp.8 Linnaeus 1758 sturgeon
Actinopteri
 Teleostei -
Teleostei indet.8
2 species
Chondrichthyes
 Myliobatiformes -
Myliobatiformes indet.8 Compagno 1973 stingray
"rayid"
 Lamniformes - Lamnidae
Lamnidae indet.8 Müller and Henle 1838 mackerel shark
Gastropoda
 Heterostropha - Pyramidellidae
 Heterostropha - Planorbidae
 Heterostropha - Acteonidae
"Actaeon cf. granum" = Acteon, Tornatellaea sp.
"Actaeon cf. granum" = Acteon snail
one corroded specimen
Tornatellaea sp. Conrad 1860 snail
one imperfect example
 Heterostropha - Ringiculidae
 Neritoidea - Neritidae
Theodoxus globulus snail
common
Theodoxus subornatus snail
common
 Cerithioidea - Cerithiidae
"Tympanotonus funatus" = Cerithium (Tympanotonus)
"Tympanotonus funatus" = Cerithium (Tympanotonus) cerith snail
common
 Cerithioidea - Thiaridae
"Bayania" triticea snail
common
 Cerithioidea - Pachychilidae
 Cerithioidea - Melanopsidae
Melanopsis buccinoidea snail
abundant (in thousands)
 Neotaenioglossa - Calyptraeidae
Calyptraea sp. Lamarck 1799 slipper shell
 Epitonioidea - Epitoniidae
"Epitonium cf. stueri" = Cirsotrema (Gyroscala) stueri, Acirsa sp.
"Epitonium cf. stueri" = Cirsotrema (Gyroscala) stueri de Boury 1890 wentletrap
rare
Acirsa sp. Mörch 1857 wentletrap
rare and corroded
 Sorbeoconcha - Naticidae
"Natica glaucinoides" = Euspira glaucinoides, Natica abducta
"Natica glaucinoides" = Euspira glaucinoides Sowerby 1812 moon snail
common
Natica abducta moon snail
common
 Neogastropoda - Buccinidae
Pseudoliva fissurata true whelk
common
Tritonidea lata true whelk
common and very variable
Tritonidea sp. Sowerby 1842 true whelk
rare
Siphonalia mariae true whelk
 Neogastropoda - Terebridae
"Terebra plicatula" = Mirula plicatula
"Terebra plicatula" = Mirula plicatula Lamarck 1803 auger snail
common
 Neogastropoda - Turridae
"Hemipleurotoma spp." = Gemmula
"Hemipleurotoma spp." = Gemmula Weinkauff 1875 turrid
3 species, rare
 Neogastropoda - Muricidae
Murex abbatiae murex snail
rare
Bivalvia
 Nuculida - Nuculidae
"Nucula fragilis" = Nucula (Nucula) fragilis
"Nucula fragilis" = Nucula (Nucula) fragilis Deshayes 1829 nut clam
common
 Mytilida - Mytilidae
"Modiola sp." = Modiolus
"Modiola sp." = Modiolus Lamarck 1801 mussel
rare
 Ostreida - Ostreidae
Ostrea sp. Linnaeus 1758 oyster
fragmentary, "several indubitable pearls have been found. These probably originated in oysters rather than in Pinnae which are not known at this horizon.
 Arcida - Glycymerididae
 Cardiida - Cardiidae
"Protocardium plumsteadianum" = Nemocardium plumsteadianum
"Protocardium plumsteadianum" = Nemocardium plumsteadianum Sowerby 1813 cockle
 Cardiida - Veneridae
"Dosiniopsis bellovacensis" = Dosiniopsis bellovacina
"Dosiniopsis bellovacensis" = Dosiniopsis bellovacina Deshayes 1825 venus clam
rare
 Cardiida - Mactridae
"Mactra semisulcata" = Spisula semisulcata
"Mactra semisulcata" = Spisula semisulcata Lamarck 1805 clam
 Cardiida - Cyrenidae
Cyrena cuneiformis clam
abundant (in thousands)
Cyrena tellinella clam
abundant (in thousands)
 Pholadida - Corbulidae
 Pholadida - Pholadidae
Barnea levesquei clam
rare
 Pholadida - Teredinidae
Teredo sp. Linnaeus 1758 clam
 Hiatellida - Hiatellidae
Cyrtodaria rutupiensis Morris 1852 clam
rare