Bore Lullingen 1, depth 54m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Tichelh. stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, sandy siltstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61898: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Polychaeta
 Sabellida - Serpulidae
Ditrupa cornea1
Rhynchonellata
 Terebratulida - Terebratulidae
Terebratula sp.1
Terebratula sp.1 Müller 1776
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
Gymnolaemata
 Cheilostomata - Bryocryptellidae
Porella ? pulchra1
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Celleporidae
Turbicellepora compressa1
 Cheilostomata - Membraniporidae
Reussirella haidingeri1, Biflustra savartii1
Biflustra savartii1 Audouin 1826
Gastropoda
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2, Turritella aff. triplicata2
 Sorbeoconcha - Naticidae
Naticidae indet.2, Euspira gottschei2, Euspira nysti2, Euspira staringi2, Euspira helicina2, Neverita josephinia2, "Natica sensu lato tigrina" = Cochlis tigrina2, Tectonatica miopusilla2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Euspira gottschei2 moon snail
Euspira nysti2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
 Sorbeoconcha - Cerithiopsidae
Cerithiopsis lottae n. sp.2, Cerithiopsis (Cerithiopsis) vandermarki2
Cerithiopsis lottae n. sp.2 snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Neogastropoda - Muricidae
Coralliophila (Pseudomurex) bracteata2, Typhis (Typhis) ? pungens2, Lyrotyphis (Eotyphis) fistulosus2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3
 Neogastropoda - Terebridae
Strioterebrum fabritzi3, Strioterebrum basteroti3
Strioterebrum basteroti3 Nyst 1843 auger snail
 Neogastropoda - Turridae
Fusiturris aquensis3, Fusiturris duchastelii3, Nitidiclavus maitrejus3, Boreodrillia hosiusi3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Perrona obliquiplicatula3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) zimmermanni3
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
sensu lato Unedogemmula boreoturricula3 turrid
incl. 66 specimens U. (s. lat.) boreoturricula x hanseata ?
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
 Neogastropoda - Raphitomidae
Raphitoma "sp. 3"3, Pleurotomella mioweberi3, Metuonella grippi3, Favriella sinuosula3, Teretia anceps3
Raphitoma "sp. 3"3 Monterosato 1884 snail
 Neogastropoda - Borsoniidae
Bathytoma jugleri3, Asthenotoma pannus3, Asthenotoma festiva3
 Neogastropoda - Mangeliidae
Benthomangelia aff. obtusangula3, Benthomangelia grippi3, Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3
Benthomangelia grippi3 Anderson 1964 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3, Clavatula sp.3
Clavatula sp.3 Lamarck 1801 snail
 Neogastropoda - Columbellidae
Mitrella (Macrurella) nassoides2, Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius cavatus2, Nassarius karinae n. sp.2, Amyclina facki2, Amyclina "sp. 2"2
Nassarius karinae n. sp.2 Wienrich 2001 snail
Amyclina "sp. 2"2 Iredale 1918 snail
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, "Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Mitridae
Mitra (Tiara) substriatula3
Mitra (Tiara) substriatula3 d'Orbigny 1852 mitre shell
 Neogastropoda - Cancellariidae
Cancellaria (Merica) contorta2, "Trigonostoma (Ventrilia) geslini" = Ventrilia2, "Trigonostoma (Ventrilia) pouwi" = Ventrilia2, Sveltia varicosa2, Babylonella fusiformis2
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
"Trigonostoma (Ventrilia) geslini" = Ventrilia2 snail
T. geslini josephinae
"Trigonostoma (Ventrilia) pouwi" = Ventrilia2 snail
Sveltia varicosa2 snail
S. varicosa paucicostata
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. gliberti
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Iravadiidae
Ceratia regiorivi2, Ceratia proxima2
Ceratia proxima2 Forbes and Hanley 1850 snail
 Sorbeoconcha - Rissoidae
Ovirissoa gottscheana2, Ovirissoa westfalica2
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
"Epitonium (Clathrus) clathrus" = Clathrus2, Epitonium (Spiniscala) frondiculum2, "Amaea (Scalina) subreticula" = Scalina2, Opalia (Nodiscala) rugatina2
"Epitonium (Clathrus) clathrus" = Clathrus2 wentletrap
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
 Vanikoroidea - Eulimidae
Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 18 specimens B. alba for. lactea
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3, Cylichna (Cylichna) subcylindrica3
 Opisthobranchia -
Roxania utriculus3
 Heterostropha - Acteonidae
Acteon (Acteon) laevigatus3, Acteon (Acteon) "sp. 2"3
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Heterostropha - Architectonicidae
Architectonica (Pseudotorinia) sensu lato obtusa3
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. pallidaeformis3, Odostomia (Megastomia) tuexeni3, Odostomia (Megastomia) "sp. 1"3, Odostomia (Megastomia) cf. conoidea3, Pyramidella (Voluspa) plicosa3, "Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla3, Eulimella (Eulimella) acicula3, Turbonilla (Turbonilla) gastaldi3, Chrysallida (Parthenina) cimbrica3, Chrysallida (Pyrgulina) pygmaea3, Kleinella (Kleinella) sp.3, Syrnola hoernesi3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla3 snail
sensu JANSSEN
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
Kleinella (Kleinella) sp.3 Adams 1860 snail
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Trochoidea - Skeneidae
Solariorbis hosiusi2
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
Bivalvia
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
incl. 3 specimens of Portlandia pygm. fa. alta
 Nuculanida - Nuculanidae
Nuculana (Jupiteria) curvirostris1, "Nuculana (Saccella) westendorpi" = Saccella westendorpi1
Nuculana (Jupiteria) curvirostris1 de Cserna 1961 pointed nut clam
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculida - Nuculidae
Nuculoma haesendoncki1, Nuculoma laevigata1, Nucula (Lamellinucula) jeffreysi1, Nucula (Nucula) nucleus1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Nucula (Lamellinucula) jeffreysi1 Bellardi 1875 nut clam
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
 Carditida - Astartidae
Goodallia (Goodallia) angulata1, Astarte (Astarte) radiata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Poromyida - Cuspidariidae
Cardiomya (Cardiomya) costellata1, Cuspidaria sp.1
Cuspidaria sp.1 Dall 1886 clam
 Cardiida - Lasaeidae
Scacchia (Scacchia) degrangei1
Scacchia (Scacchia) degrangei1 Cossmann and Peyrot 1911 clam
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Glossidae
Meiocardia harpa1, Glossus (Glossus) lunulatus1
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Veneridae
"Gouldia minima" = Gouldia (Gouldia) minima1, Venus (Ventricoloidea) multilamella1
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Semelidae
Abra (? Abra) sorgenfreii1
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, "Habecardium sp." = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium ovale1, Parvicardium straeleni1, Parvicardium scabrum1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
"Habecardium sp." = Laevicardium (Habecardium)1 Glibert and van de Poel 1970 cockle
Habecardium juv.
Habecardium dingdense1 Lehmann 1885 cockle
Parvicardium scabrum1 Philippi 1844 cockle
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Xylophagaidae
Xylophaga sp.1
Xylophaga sp.1 Turton 1822 clam
 Pholadida - Pholadidae
Pholadidae indet.1
Pholadidae indet.1 Lamarck 1809 clam
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Hiatellida - Hiatellidae
"Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Lucinida - Thyasiridae
Thyasira (Thyasira) flexuosa1
Thyasira (Thyasira) flexuosa1 Montagu 1803 clam
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
 Mytilida - Mytilidae
Modiolula phaseolina1
Modiolula phaseolina1 Philippi 1844 mussel
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Pectinida - Limidae
Limatula sulcata1, Limea (Limea) aff. strigilata1
Limatula sulcata1 file clam
Limea (Limea) aff. strigilata1 Brocchi 1814 file clam
 Pectinida - Pectinoidae
Korobkovia woodi1, Aequipecten seniensis1, Pseudamussium lilli1
Pseudamussium lilli1 Pusch 1837 scallop
 Arcida - Arcidae
Bathyarca pectunculoides1, Scapharca (Scapharca) diluvii1
Bathyarca pectunculoides1 Scacchi 1834 clam
Scapharca (Scapharca) diluvii1 Lamarck 1805 clam
 Arcida - Glycymerididae
Glycymeris (Glycymeris) obovata1
Glycymeris (Glycymeris) obovata1 Lamarck 1819 clam
G. obovata baldii
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
Scaphopoda
 Dentaliida - Laevidentaliidae
Laevidentalium sp.1
Laevidentalium sp.1 Cossmann 1888 tusk shell
Laevidentalium sp. juv.
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Pennatulacea -
Graphularia beyrichi
Globothalamea
 Rotaliida - Nonionidae
Nonion boueanum
 Rotaliida - Cibicididae
Cibicides aknerianus, Cibicides dutemplei
Cibicides dutemplei dOrbigny 1846
Foraminifera
 Globigerinoidea - Globigerinidae
Globigerinoides trilobus
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
Nodosariata
 Robertinida - Ceratobuliminidae
Ceratobulimina haueri
 Nodosariida - Nodosariidae
Dentalina guttifera, Dentalina obliquestriata, Dentalina inornata, Dentalina semiplicata, Dentalina konincki
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Guttulina austriaca, Guttulina problema, Globulina gibba, Pseudopolymorphina obscura, Sigmomorphina schwageri, Sigmomorphina sp.
Guttulina problema d'Orbigny 1826
Globulina gibba d'Orbigny 1832
incl. 3 specimens G. gibba for. tubulosa
Sigmomorphina sp. Cushman and Ozawa 1928
 Vaginulinida - Vaginulinidae
Lenticulina (Lenticulina) calcar, Lenticulina (Robulus) orbicularis
Foraminifera
  - Spiroplectamminidae
Spiroplectammina carinata
 Lituolida - Lituolidae
Textularia laevigata, Textularia gramen
Textularia gramen dOrbigny 1846
Tubothalamea
 Miliolida - Miliolidae
Miliolidae indet., Pyrgo simplex, Spiroloculina striatula
Miliolidae indet. Ehrenberg 1839
 Miliolida - Hauerinidae
Triloculina tricarinata, Triloculina inflata, Triloculina scapha, Quinqueloculina haidingeri, Quinqueloculina concinna, Quinqueloculina akneriana, Quinqueloculina buchiana, Quinqueloculina triangularis
Quinqueloculina buchiana d'Orbigny 1846
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007