Bore Lullingen 1, depth 48m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, sandy siltstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61906: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Gymnolaemata
 Cheilostomata - Membraniporidae
Reussirella haidingeri1
 Cheilostomata - Lunulitidae
Lunulites burdigalensis1
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Errantida indet. B and C
 Sabellida - Serpulidae
Spirorbis sp.1
Spirorbis sp.1 Daudin 1800
Scaphopoda
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, Fissidentalium twistringense1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Gastropoda
 Heterostropha - Pyramidellidae
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Syrnola neumayri3, Chrysallida (Pyrgulina) pygmaea3, Chrysallida (Parthenina) toftlundensis3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) tuexeni3, Kleinella (Kleinella) sp.3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
Kleinella (Kleinella) sp.3 Adams 1860 snail
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
 Heterostropha - Murchisonellidae
Ebala (Ebala) vandervlerki3
 Heterostropha - Architectonicidae
Architectonica (Pseudotorinia) sensu lato obtusa3, Architectonica (Architectonica) simplex3, "Architectonica (Pseudotorinia) planulata" = Pseudotorinia3
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
 Heterostropha - Acteonidae
Acteon (Acteon) arnumensis3, Acteon (Acteon) laevigatus3, Acteon (Acteon) semistriatus3, Acteon (Acteon) "sp. 3"3
Acteon (Acteon) arnumensis3 Sorgenfrei 1958 snail
spelled as 'Actaeon'
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 3"3 Montfort 1810 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Trochoidea - Skeneidae
Skenea minuta2, Solariorbis hosiusi2
 Thecosomata - Cavoliniidae
Vaginella austriaca3
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3, Retusa (Cylichnina) subangistoma3
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3
 Opisthobranchia -
Roxania utriculus3
 Cerithioidea - Cerithiidae
Bittium (Bittium) tenuispina2, "Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2, Turritella aff. triplicata2
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, Epitonium (Cinctoscala) linoe2, sensu lato Epitonium "sp. 1"2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
sensu lato Epitonium "sp. 1"2 Bolten 1798 wentletrap
 Sorbeoconcha - Naticidae
Naticidae indet.2, Neverita josephinia2, Polinices (Polinices) miocolligens2, Euspira staringi2, Euspira sp.2, Euspira helicina2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2, "Natica neglecta" = Cochlis neglecta2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
Euspira staringi2 moon snail
Euspira sp.2 Agassiz 1837 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
 Sorbeoconcha - Triphoridae
Biforina (Oriforina) aff. praeversa2
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Sorbeoconcha - Rissoidae
sensu lato Alvania holsatica2, Alvania (Alvania) partschi2, sensu lato Alvania antwerpiensis2, Ovirissoa westfalica2
 Sorbeoconcha - Vitrinellidae
Circulus subcirculus2, Circulus hennei2
Circulus hennei2 Glibert 1952 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2, Anachis (Costoanachis) "sp. 2"2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, "Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2, ? Exilifusus attenuatus2
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Nassariidae
Amyclina facki2, Amyclina banatica2, Amyclina badensis2, Amyclina "sp. 1"2, Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius woodwardi2, Nassarius karinae2
Amyclina "sp. 1"2 Iredale 1918 snail
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Cancellariidae
Narona (Calcarata) calcarata2, Cancellaria (Merica) contorta2, Cancellaria (Bivetiella) cancellata2, "Trigonostoma (Ventrilia) pouwi" = Ventrilia2, Sveltia varicosa2, Babylonella fusiformis2
Narona (Calcarata) calcarata2 Brochhi 1814 snail
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Cancellaria (Bivetiella) cancellata2 Linnaeus 1767 snail
C. cancellata praecedens
"Trigonostoma (Ventrilia) pouwi" = Ventrilia2 snail
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Mitridae
Mitra (Tiara) substriatula3
Mitra (Tiara) substriatula3 d'Orbigny 1852 mitre shell
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Mangeliidae
sensu lato Mangelia luellingensis n. sp.3, Benthomangelia aff. obtusangula3, Benthomangelia grippi3, Sorgenfreispira plicatelloides3, Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3
sensu lato Mangelia luellingensis n. sp.3 Wienrich and Janssen 2007 snail
Benthomangelia grippi3 Anderson 1964 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Clathurellidae
? Ceritoturris hannoverana3, Pleurotomoides luisae3
? Ceritoturris hannoverana3 snail
C. (?) hannoverana
 Neogastropoda - Terebridae
Strioterebrum fabritzi3, Strioterebrum hoernesi3, Strioterebrum ronaldi3, Strioterebrum basteroti3
Strioterebrum ronaldi3 auger snail
Strioterebrum basteroti3 Nyst 1843 auger snail
 Neogastropoda - Conidae
Conus (Chelyconus) cf. clavatulus3, "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Raphitomidae
Metuonella grippi3, Teretia anceps3, Raphitoma "sp. 3"3
Raphitoma "sp. 3"3 Monterosato 1884 snail
 Neogastropoda - Turridae
Perrona obliquiplicatula3, Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) schafferi3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) vermicularis3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3
sensu lato Unedogemmula boreoturricula3 turrid
incl. 3 specimens U. (s. lat.) boreoturricula x hanseata ?
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Borsoniidae
Asthenotoma pannus3, Asthenotoma festiva3, Bathytoma jugleri3, "Genota ramosa" = Genota valeriae3, Microdrillia aff. serratula3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3, "Vexillum (Uromitra) aciculum" = Uromitra3, "Vexillum (Uromitra) gliberti" = Uromitra3
"Vexillum (Uromitra) aciculum" = Uromitra3 ribbed mitre
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Muricidae
Typhis (Typhis) ? pungens2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2, Coralliophila (Pseudomurex) bracteata2, Lyrotyphis (Eotyphis) fistulosus2, Hadriania coelata2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
Hadriania coelata2 Dujardin 1837 murex snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Sorbeoconcha - Cassidae
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2, "Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Vanikoroidea - Eulimidae
Niso (Niso) terebellum2, Eulima (? Batheulima) sp.2, Eulima (Eulima) glabra2, Balcis (Balcis) alba2
Eulima (? Batheulima) sp.2 Risso 1826 snail
E. (? Batheulima) sp. nov.
Balcis (Balcis) alba2 snail
B. alba for. lactea
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculanida - Yoldiidae
"Portlandia (Yoldiella) pygmaea" = Yoldiella1, Yoldia (Yoldia) glaberrima1
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
 Nuculida - Nuculidae
Nucula (Nucula) nucleus1, Nuculoma haesendoncki1
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Mytilida - Mytilidae
Modiolula phaseolina1
Modiolula phaseolina1 Philippi 1844 mussel
 Pectinida - Pectinoidae
Aequipecten angelonii1, Aequipecten seniensis1, Korobkovia woodi1, Pseudamussium lilli1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Limidae
Limea (Limea) aff. strigilata1
Limea (Limea) aff. strigilata1 Brocchi 1814 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1, Pododesmus (Monia) anitae1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
Pododesmus (Monia) anitae1 Janse and Janssen 1983 scallop
 Pectinida - Propeamussiidae
Similipecten similis1
Similipecten similis1 Laskey 1811 mud scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Arcida - Arcidae
Bathyarca pectunculoides1, Scapharca (Scapharca) diluvii1
Bathyarca pectunculoides1 Scacchi 1834 clam
Scapharca (Scapharca) diluvii1 Lamarck 1805 clam
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium straeleni1, Parvicardium scabrum1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
Parvicardium scabrum1 Philippi 1844 cockle
 Cardiida - Semelidae
sensu lato Abra lehmanni1
 Cardiida - Tellinidae
"Macoma elliptica" = Laciolina magna1, Angulus (Laceolina) benedeni1
"Macoma elliptica" = Laciolina magna1 Spengler 1798 tellin clam
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Psammobiidae
Gari (Psammotaena) aff. angusta1
 Cardiida - Glossidae
Meiocardia harpa1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Hiatellida - Hiatellidae
"Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Poromyida - Verticordiidae
Haliris (Haliris) "sp. 1"1
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Lucinida - Thyasiridae
Thyasira (Thyasira) flexuosa1
Thyasira (Thyasira) flexuosa1 Montagu 1803 clam
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1, "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 35 specimens, Astarte radiata fa. convexior 40 specimens
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
Anthozoa
 Scleractinia - Caryophylliidae
Deltocyathus reussi
 Scleractinia - Turbinoliidae
"Cylindrophyllia duncani" = Peponocyathus duncani
"Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum tuberculatum
Flabellum sp. Lesson 1831 stony coral
Flabellum tuberculatum Keferstein 1859 stony coral
 Scleractinia - Dendrophylliidae
"Eupsammia calyculus" = Balanophyllia, "Eupsammia praelonga" = Balanophyllia praelonga
"Eupsammia calyculus" = Balanophyllia stony coral
"Eupsammia praelonga" = Balanophyllia praelonga Michelotti 1838 stony coral
Foraminifera
 Lituolida - Lituolidae
Textularia sp., Textularia gramen
Textularia sp. Defrance 1824
Textularia gramen dOrbigny 1846
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides dutemplei
Cibicides dutemplei dOrbigny 1846
 Rotaliida - Nonionidae
Nonion boueanum
 Rotaliida - Gavelinellidae
Gyroidina soldanii
Nodosariata
 Nodosariida - Nodosariidae
Vaginulina obtusicosta, Nodosaria bacillum, Dentalina bifurcata, Dentalina grossicosta, Dentalina konincki, Lenticulina (Robulus) clypeiformis
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Pseudopolymorphina subnodosa, Pseudopolymorphina incerta, Globulina gibba, Guttulina austriaca, Guttulina problema
Globulina gibba d'Orbigny 1832
Guttulina problema d'Orbigny 1826
 Vaginulinida - Vaginulinidae
Lenticulina (Robulus) orbicularis, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) grandis, Lenticulina (Robulus) ornata, Lenticulina (Planularia) grundensis, Lenticulina (Planularia) kubinyi, Lenticulina (Lenticulina) inornata, Lenticulina (Lenticulina) calcar
Tubothalamea
 Miliolida - Hauerinidae
Cycloforina badenensis
Foraminifera
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
  - Spiroplectamminidae
Spiroplectammina carinata
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007