PBDB Collection

Bore Kevelaer, depth 60m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown, gray, green sandstone

Size classes: macrofossils, mesofossils, microfossils

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62889: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Foraminifera
Rotaliacea - Rotaliidae	Hoeglundina elegans Hoeglundina elegans
Discorbacea - Bagginidae	Cancris auriculus Cancris auriculus Fichtel and Moll 1798
Buliminacea - Uvigerinidae	Uvigerina semiornata, Uvigerina acuminata Uvigerina semiornata U. semiornata for. semiornata Uvigerina acuminata
Nodosariacea - Glandulinidae	Glandulina ovula Glandulina ovula
Asterigerinacea - Asterigerinidae	Asterigerina guerichi Asterigerina guerichi A. gürichi staeschi
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
-	"Marginulina hirsuta" = Cristellaria (Marginulina) "Marginulina hirsuta" = Cristellaria (Marginulina)
Nodosariata
Nodosariida - Nodosariidae	Lenticulina (Robulus) vortex, Nodosaria bacillum, Dentalina intermittens, Dentalina konincki, Dentalina pauperata, Lagena globosa Lenticulina (Robulus) vortex Fichtel and Moll 1798 Nodosaria bacillum Dentalina intermittens Dentalina konincki Reuss 1861 Dentalina pauperata d'Orbigny 1846 Lagena globosa Montagu 1803
Polymorphinida - Polymorphinidae	Globulina gibba, Guttulina problema, Pseudopolymorphina subnodosa Globulina gibba d'Orbigny 1832 Guttulina problema d'Orbigny 1826 Pseudopolymorphina subnodosa incl. 1 specimen P. subnodosa for. rugosa
Vaginulinida - Vaginulinidae	Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) similis, Lenticulina (Robulus) ornata, Lenticulina (Lenticulina) calcar Lenticulina (Lenticulina) inornata d'Orbigny 1846 Lenticulina (Robulus) cultrata Lenticulina (Robulus) similis Lenticulina (Robulus) ornata Lenticulina (Lenticulina) calcar
Foraminifera
Lituolida - Lituolidae	Textularia gramen, Bigenerina nodosaria Textularia gramen dOrbigny 1846 Bigenerina nodosaria
Tubothalamea
Miliolida - Hauerinidae	Quinqueloculina akneriana Quinqueloculina akneriana
Miliolida - Miliolidae	Sigmoilina tenuis Sigmoilina tenuis
Foraminifera
Rotalioidea - Elphidiidae	Elphidium inflatum, Elphidium ungeri Elphidium inflatum Elphidium ungeri
Lituolacea - Ataxophragmiidae	Martinottiella communis Martinottiella communis Huang et al. 2019
Globothalamea
Rotaliida - Nonionidae	Cribrononion subnodosum, Nonion boueanum, Nonion affine, Nonion (Porosononion) soldanii Cribrononion subnodosum Nonion boueanum Nonion affine Nonion (Porosononion) soldanii
Rotaliida - Gavelinellidae	Gyroidina soldanii Gyroidina soldanii
Rotaliida - Cibicididae	Planulina austriaca, Cibicides lobatulus, Cibicides aknerianus, Cibicides ungerianus, Cibicides dutemplei, Cibicides pseudoungerianus Planulina austriaca Cibicides lobatulus incl. 1 specimen C. lobatulus var. ornatus Cibicides aknerianus Cibicides ungerianus Cibicides dutemplei dOrbigny 1846 Cibicides pseudoungerianus Cushman 1922
Foraminifera
- Chilostomellidae	Sphaeroidina bulloides Sphaeroidina bulloides d'Orbigny 1826
Anthozoa
Scleractinia - Flabellidae	Flabellum sp. Flabellum sp. Lesson 1831 stony coral
Bivalvia
Nuculanida - Yoldiidae	"Portlandia (Yoldiella) pygmaea" = Yoldiella¹, Yoldia (Yoldia) glaberrima¹ "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam Yoldia (Yoldia) glaberrima¹ von Münster 1837 clam
Nuculanida - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹, Nuculana (Jupiteria) curvirostris¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam Nuculana (Jupiteria) curvirostris¹ de Cserna 1961 pointed nut clam
Nuculida - Nuculidae	Nucula (Nucula) nucleus¹, Nucula (Lamellinucula) comta¹ Nucula (Nucula) nucleus¹ Linnaeus 1758 nut clam Nucula (Lamellinucula) comta¹ Goldfuss 1837 nut clam
Carditida - Astartidae	Astarte (Astarte) radiata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. radiata 1 specimen, Astarte radiata fa. convexior 7 specimens, Astarte radiata fa. goldfussi 1 specimen
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Veneridae	"Gouldia minima" = Gouldia (Gouldia) minima¹, Venus (Ventricoloidea) multilamella¹ "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella
Cardiida - Cardiidae	Parvicardium straeleni¹, "Habecardium subturgidum" = Laevicardium (Habecardium)¹ Parvicardium straeleni¹ cockle "Habecardium subturgidum" = Laevicardium (Habecardium)¹ cockle H. subturgidum subturgidum
Arcida - Limopsidae	"Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹, Limopsis (Limopsis) aurita¹ "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam Limopsis (Limopsis) aurita¹ Brocchi 1814 clam
Arcida - Arcidae	Scapharca (Scapharca) diluvii¹ Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Pectinida - Pectinoidae	Aequipecten angelonii¹ Aequipecten angelonii¹ scallop
Scaphopoda
Dentaliida - Dentaliidae	Lentigodentalium pseudomutabile¹, "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ Lentigodentalium pseudomutabile¹ tusk shell "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ tusk shell
Gastropoda
Opisthobranchia - Cylichnidae	Cylichna (Cylichna) pseudoconvoluta³, Scaphander grateloupi³ Cylichna (Cylichna) pseudoconvoluta³ snail Scaphander grateloupi³ snail
Opisthobranchia - Retusidae	Volvulella (Volvulella) acuminata³, Retusa (Cylichnina) elongata³ Volvulella (Volvulella) acuminata³ snail Retusa (Cylichnina) elongata³ snail
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Pseudomelanoidea - Pseudomelaniidae	sensu lato Inquisitor borealis³ sensu lato Inquisitor borealis³ snail
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Cerithioidea - Cerithiidae	"Bittium (Bittium) spina" = Bittium spina², Bittium (Bittium) tenuispina² "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail Bittium (Bittium) tenuispina² cerith snail
Cerithioidea - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Sorbeoconcha - Rissoidae	Alvania (Alvania) basisulcata² Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Triviidae	Trivia (Trivia) cf. westfalica² Trivia (Trivia) cf. westfalica² Schilder 1929 snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Neogastropoda - Muricidae	Lyrotyphis (Eotyphis) fistulosus² Lyrotyphis (Eotyphis) fistulosus² murex snail
Neogastropoda - Cancellariidae	Sveltia varicosa², Babylonella fusiformis², "Trigonostoma (Ventrilia) pouwi" = Ventrilia² Sveltia varicosa² snail S. varicosa paucicostata Babylonella fusiformis² snail "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Raphitomidae	Teretia anceps³, Favriella sinuosula³, Mioawateria andersoni³, Metuonella grippi³, Raphitoma aff. praehispida³ Teretia anceps³ snail Favriella sinuosula³ snail Mioawateria andersoni³ snail Metuonella grippi³ snail Raphitoma aff. praehispida³ snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³ Pleurotomoides luisae³ snail
Neogastropoda - Borsoniidae	Bathytoma jugleri³, "Genota ramosa" = Genota valeriae³, Asthenotoma pannoides³, Asthenotoma pannus³ Bathytoma jugleri³ snail "Genota ramosa" = Genota valeriae³ Hoernes and Auinger 1891 snail Asthenotoma pannoides³ snail Asthenotoma pannus³ snail
Neogastropoda - Mangeliidae	Sorgenfreispira tenella³, Benthomangelia aff. obtusangula³ Sorgenfreispira tenella³ Conrad 1848 snail Benthomangelia aff. obtusangula³ snail
Neogastropoda - Turridae	Boreodrillia hosiusi³, Nitidiclavus maitrejus³, Gemmula (Gemmula) zimmermanni³, Perrona obliquiplicatula³, Fusiturris aquensis³, Fusiturris duchastelii³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³ Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid Gemmula (Gemmula) zimmermanni³ turrid Perrona obliquiplicatula³ turrid Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid F. duchastelii flexiplicata sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Sorbeoconcha - Perissityidae	Mitrella (Macrurella) nassoides², Anachis (Costoanachis) hosiusi², ? Exilifusus attenuatus², Streptodictyon sexcostatus², Nassarius bocholtensis², Nassarius tenuistriatus² Mitrella (Macrurella) nassoides² true whelk Anachis (Costoanachis) hosiusi² true whelk ? Exilifusus attenuatus² true whelk Streptodictyon sexcostatus² true whelk Nassarius bocholtensis² true whelk Nassarius tenuistriatus² true whelk
Caenogastropoda - Fasciolariidae	Niso (Niso) terebellum², Norephora (Norephora) fritschi², Naticidae indet.², Euspira staringi², "Natica sensu lato tigrina" = Cochlis tigrina², Aquilofusus festivus² Niso (Niso) terebellum² snail Norephora (Norephora) fritschi² snail Naticidae indet.² Guilding 1834 moon snail Euspira staringi² moon snail "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Aquilofusus festivus² worm shell
Epitonioidea - Epitoniidae	"Amaea (Scalina) subreticula" = Scalina² "Amaea (Scalina) subreticula" = Scalina² wentletrap
Trochoidea - Trochidae	Calliostoma (Ampullotrochus) ? laureatum² Calliostoma (Ampullotrochus) ? laureatum² top snail
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Heterostropha - Acteonidae	Acteon (Acteon) "sp. 2"³ Acteon (Acteon) "sp. 2"³ Montfort 1810 snail spelled as 'Actaeon'
Heterostropha - Pyramidellidae	Pyramidella (Voluspa) elata³, Pyramidella (Voluspa) plicosa³, Chrysallida (Pyrgulina) pygmaea³, Chrysallida (Parthenina) cimbrica³, Turbonilla (Mormula) cf. miomutiniensis³, Turbonilla (Turbonilla) koeneniana³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, Syrnola hoernesi³, Odostomia (Megastomia) tuexeni³ Pyramidella (Voluspa) elata³ snail Pyramidella (Voluspa) plicosa³ snail Chrysallida (Pyrgulina) pygmaea³ Grateloup 1838 snail incl. 5 specimens Chrysallida pygmaea f. belgica Chrysallida (Parthenina) cimbrica³ snail Turbonilla (Mormula) cf. miomutiniensis³ snail sensu ANDERSON Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN Syrnola hoernesi³ snail Odostomia (Megastomia) tuexeni³ snail
Heterostropha - Architectonicidae	"Architectonica (Pseudotorinia) sensu lato obtusa" = Pseudotorinia³ "Architectonica (Pseudotorinia) sensu lato obtusa" = Pseudotorinia³ snail
Polychaeta
Sabellida - Serpulidae	Ditrupa cornea¹ Ditrupa cornea¹
Gymnolaemata
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Cheilostomata - Lunulitidae	Lunulites voigti¹, Lunulites burdigalensis¹ Lunulites voigti¹ Lunulites burdigalensis¹

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007