PBDB Collection

Bore Wetten, depth 28m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.9° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Miocene (23.0 - 5.3 Ma)

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1991

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62899: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Foraminifera
Rotalioidea - Elphidiidae	Elphidium inflatum Elphidium inflatum
Nodosariacea - Glandulinidae	Glandulina ovula Glandulina ovula
Globothalamea
Rotaliida - Cibicididae	Cibicides lobatulus, Cibicides dutemplei Cibicides lobatulus Cibicides dutemplei dOrbigny 1846
Foraminifera
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
-	"Marginulina cf. costata" = Cristellaria (Marginulina) "Marginulina cf. costata" = Cristellaria (Marginulina)
Nodosariata
Vaginulinida - Vaginulinidae	Lenticulina (Vaginulinopsis) gerlachi, Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) gibba, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) ariminensis, Lenticulina (Robulus) grandis, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) depauperata, Lenticulina (Robulus) ornata Lenticulina (Vaginulinopsis) gerlachi Lenticulina (Lenticulina) calcar Lenticulina (Lenticulina) gibba Lenticulina (Lenticulina) inornata d'Orbigny 1846 Lenticulina (Robulus) ariminensis Lenticulina (Robulus) grandis Lenticulina (Robulus) cultrata Lenticulina (Robulus) depauperata L. (L.) depauperata for. costata Lenticulina (Robulus) ornata
Vaginulinida - Lagenidae	Darbyella nitida Darbyella nitida
Nodosariida - Nodosariidae	Lenticulina (Robulus) simplex, Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina urnula, Dentalina konincki, Nodosaria bacillum, Nodosaria rudis, Nodosaria grandis Lenticulina (Robulus) simplex d'Orbigny 1846 Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina urnula Dentalina konincki Reuss 1861 Nodosaria bacillum Nodosaria rudis Nodosaria grandis
Polymorphinida - Polymorphinidae	Globulina gibba, Guttulina problema, Pseudopolymorphina subnodosa Globulina gibba d'Orbigny 1832 Guttulina problema d'Orbigny 1826 Pseudopolymorphina subnodosa Pseudop. subn. for. rugosa
Scaphopoda
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Dentaliida - Dentaliidae	"Pseudantalis aturensis" = Dentalium (Pseudantilis)¹, Antalis dumasi¹, Lentigodentalium pseudomutabile¹ "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ tusk shell Antalis dumasi¹ tusk shell Antalis dumasi subsp. Lentigodentalium pseudomutabile¹ tusk shell
Gastropoda
Epitonioidea - Epitoniidae	Opalia (Pliciscala) pertusa², "Amaea (Scalina) subreticula" = Scalina², sensu lato Turriscala straeleni², Acirsa (Hemiacirsa) lanceolata², Acirsa (Hemiacirsa) aff. duvergieri² Opalia (Pliciscala) pertusa² wentletrap "Amaea (Scalina) subreticula" = Scalina² wentletrap sensu lato Turriscala straeleni² wentletrap T. (s. lat.) stareleni f. dingdensis Acirsa (Hemiacirsa) lanceolata² wentletrap Acirsa (Hemiacirsa) aff. duvergieri² wentletrap
Vanikoroidea - Eulimidae	Niso (Niso) terebellum², Balcis (Polygyreulima) glabella², Balcis (Balcis) alba², Eulima (Eulima) bilineata², Eulima (Eulima) glabra² Niso (Niso) terebellum² snail Balcis (Polygyreulima) glabella² snail Balcis (Balcis) alba² snail Eulima (Eulima) bilineata² snail Eulima (Eulima) glabra² snail
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Cerithioidea - Planaxidae	Couthouyia aff. roberti² Couthouyia aff. roberti² snail
Cerithioidea - Cerithiidae	Bittium (Bittium) tenuispina², "Bittium (Bittium) spina" = Bittium spina² Bittium (Bittium) tenuispina² cerith snail "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail
Cerithioidea - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Pseudomelanoidea - Pseudomelaniidae	sensu lato Inquisitor borealis³ sensu lato Inquisitor borealis³ snail
Sorbeoconcha - Naticidae	Naticidae indet.², "Natica neglecta" = Cochlis neglecta², "Natica sensu lato tigrina" = Cochlis tigrina², Euspira gottschei², Euspira nysti², Euspira staringi², Euspira helicina², Neverita josephinia², Natica koeneni², Tectonatica miopusilla² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. "Natica neglecta" = Cochlis neglecta² Mayer 1858 moon snail "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Euspira gottschei² moon snail Euspira nysti² moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina Neverita josephinia² Risso 1826 moon snail N. josephinia olla Natica koeneni² moon snail Tectonatica miopusilla² moon snail
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Adeorbidae	Daronia dingdensis² Daronia dingdensis² snail
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Ranellidae	"Charonia (Sassia) tarbelliana" = Sassia² "Charonia (Sassia) tarbelliana" = Sassia² triton shell
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Cancellariidae	Babylonella fusiformis², Sveltia varicosa², "Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta² Babylonella fusiformis² snail Sveltia varicosa² snail S. varicosa paucicostata "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus², Streptodictyon abruptus², ? Exilifusus attenuatus² Streptodictyon sexcostatus² snail Streptodictyon abruptus² snail incl. 1 specimen S. abruptus for. quinquecostatus ? Exilifusus attenuatus² snail
Neogastropoda - Nassariidae	Amyclina facki², Nassarius bocholtensis², Nassarius tenuistriatus², Nassarius twistringensis², Nassarius karinae² Amyclina facki² snail Nassarius bocholtensis² snail Nassarius tenuistriatus² snail Nassarius twistringensis² snail Nassarius karinae² Wienrich 2001 snail
Neogastropoda - Columbellidae	Mitrella (Macrurella) nassoides², Anachis (Costoanachis) hosiusi², Anachis (Costoanachis) terebralis² Mitrella (Macrurella) nassoides² snail Anachis (Costoanachis) hosiusi² snail Anachis (Costoanachis) terebralis² snail
Neogastropoda - Volutidae	Scaphella (Scaphella) bolli² Scaphella (Scaphella) bolli² volute
Neogastropoda - Muricidae	"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta², "Murex (Haustellum) inornatus" = Haustellum², Pterynotus (Alipurpura) parvifolius², Lyrotyphis (Eotyphis) fistulosus², Typhis (Typhis) ? pungens² "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta² Bronn 1831 murex snail "Murex (Haustellum) inornatus" = Haustellum² murex snail Pterynotus (Alipurpura) parvifolius² mustached bat Lyrotyphis (Eotyphis) fistulosus² murex snail Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³, Terebra (Myurellina) acuminata³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail Terebra (Myurellina) acuminata³ auger snail T. acuminata neglecta
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Cochlespiridae	Cochlespira corneti³ Cochlespira corneti³ snail
Neogastropoda - Mangeliidae	sensu lato Mangelia andersoni³, sensu lato Mangelia miorugulosa³, Benthomangelia aff. obtusangula³, Sorgenfreispira sorgenfreii³, Sorgenfreispira tenella³ sensu lato Mangelia andersoni³ Wienrich and Janssen 2007 snail sensu lato Mangelia miorugulosa³ Kautsky 1925 snail Benthomangelia aff. obtusangula³ snail Sorgenfreispira sorgenfreii³ snail Sorgenfreispira tenella³ Conrad 1848 snail
Neogastropoda - Raphitomidae	Teretia anceps³, Teretia "sp. 1"³, Mioawateria andersoni³, Favriella sinuosula³, Metuonella grippi³, Raphitoma aff. praehispida³ Teretia anceps³ snail Teretia "sp. 1"³ Norman 1888 snail Mioawateria andersoni³ snail Favriella sinuosula³ snail Metuonella grippi³ snail Raphitoma aff. praehispida³ snail
Neogastropoda - Turridae	Boreodrillia hosiusi³, Nitidiclavus maitrejus³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, Gemmula (Gemmula) vermicularis³, Gemmula (Gemmula) zimmermanni³, Fusiturris aquensis³, Fusiturris duchastelii³ Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid Gemmula (Gemmula) vermicularis³ turrid Gemmula (Gemmula) zimmermanni³ turrid Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid F. duchastelii flexiplicata
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³, Pleurotomoides simplex³, ? Ceritoturris hannoverana³ Pleurotomoides luisae³ snail Pleurotomoides simplex³ snail ? Ceritoturris hannoverana³ snail
Neogastropoda - Borsoniidae	"Genota ramosa" = Genota valeriae³, Microdrillia aff. serratula³, Asthenotoma pannoides³, Bathytoma jugleri³ "Genota ramosa" = Genota valeriae³ Hoernes and Auinger 1891 snail Microdrillia aff. serratula³ snail Asthenotoma pannoides³ snail Bathytoma jugleri³ snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus quadricarinatus², Circulus subcirculus², Circulus hennei² Circulus praecedens² snail C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti Circulus quadricarinatus² snail Circulus subcirculus² snail Circulus hennei² Glibert 1952 snail
Sorbeoconcha - Tornidae	Tornus (Tornus) trigonostoma² Tornus (Tornus) trigonostoma² snail
Heterostropha - Mathildidae	sensu lato Mathilda concinna³ sensu lato Mathilda concinna³ snail
Heterostropha - Pyramidellidae	"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros³, Turbonilla (Mormula) cf. miomutiniensis³, Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³, Kleinella (Kleinella) nordmanni³, Pyramidella (Voluspa) elata³, Pyramidella (Voluspa) plicosa³, Eulimella (Eulimella) acicula³, Eulimella (Eulimella) scillae³, Odostomia (Megastomia) tuexeni³, "Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla³, Syrnola hoernesi³, Syrnola neumayri³ "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros³ Sacco 1892 snail Turbonilla (Mormula) cf. miomutiniensis³ snail sensu ANDERSON Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN Kleinella (Kleinella) nordmanni³ snail Pyramidella (Voluspa) elata³ snail Pyramidella (Voluspa) plicosa³ snail Eulimella (Eulimella) acicula³ snail Eulimella (Eulimella) scillae³ snail Odostomia (Megastomia) tuexeni³ snail "Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla³ snail sensu JANSSEN Syrnola hoernesi³ snail Syrnola neumayri³ snail
Heterostropha - Architectonicidae	Architectonica (Pseudotorinia) sensu lato obtusa³ Architectonica (Pseudotorinia) sensu lato obtusa³ sundial
Heterostropha - Acteonidae	Acteon (Acteon) semistriatus³, Acteon (Acteon) "sp. 2"³ Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon' Acteon (Acteon) "sp. 2"³ Montfort 1810 snail spelled as 'Actaeon'
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Trochoidea - Trochidae	Calliostoma (Ampullotrochus) ? laureatum² Calliostoma (Ampullotrochus) ? laureatum² top snail
Opisthobranchia - Retusidae	Retusa (Cylichnina) elongata³, Volvulella (Volvulella) acuminata³ Retusa (Cylichnina) elongata³ snail Volvulella (Volvulella) acuminata³ snail
Opisthobranchia - Cylichnidae	Bullichna paucistriata³, Cylichna (Cylichna) pseudoconvoluta³ Bullichna paucistriata³ snail Cylichna (Cylichna) pseudoconvoluta³ snail
Opisthobranchia - Bullidae	"Tornatina bellardi" = Bulla (Tornatina)³ "Tornatina bellardi" = Bulla (Tornatina)³ snail
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Thecosomata - Cavoliniidae	Vaginella austriaca³ Vaginella austriaca³ snail
Thecosomata - Limacinidae	Limacina miorostralis³, Limacina valvatina³ Limacina miorostralis³ snail Limacina valvatina³ Reuss 1867 snail
Bivalvia
Nuculanida - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Nuculanida - Yoldiidae	"Portlandia (Yoldiella) pygmaea" = Yoldiella¹ "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Hiatellida - Hiatellidae	Panopea (Panopea) meynardi¹ Panopea (Panopea) meynardi¹ clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Cardiida - Kelliellidae	Spaniodontella nitida¹ Spaniodontella nitida¹ clam
Cardiida - Veneridae	Venus (Ventricoloidea) multilamella¹, "Gouldia minima" = Gouldia (Gouldia) minima¹ Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam
Cardiida - Cardiidae	"Habecardium subturgidum" = Laevicardium (Habecardium)¹, Parvicardium straeleni¹ "Habecardium subturgidum" = Laevicardium (Habecardium)¹ cockle H. subturgidum subturgidum Parvicardium straeleni¹ cockle
Lucinida - Lucinidae	"Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ "Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ Linnaeus 1767 clam
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Carditida - Astartidae	Astarte (Astarte) radiata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. convexior
Pectinida - Pectinoidae	Korobkovia woodi¹, Pseudamussium lilli¹, Aequipecten angelonii¹ Korobkovia woodi¹ scallop Pseudamussium lilli¹ Pusch 1837 scallop Aequipecten angelonii¹ scallop
Arcida - Arcidae	Bathyarca pectunculoides¹, Scapharca (Scapharca) diluvii¹ Bathyarca pectunculoides¹ Scacchi 1834 clam Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Gymnolaemata
Cheilostomata - Lunulitidae	Lunulites voigti¹ Lunulites voigti¹
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Anthozoa
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral
Scleractinia - Caryophylliidae	Deltocyathus reussi Deltocyathus reussi stony coral
Scleractinia - Turbinoliidae	"Cylindrophyllia duncani" = Peponocyathus duncani "Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
Scleractinia - Flabellidae	Flabellum sp. Flabellum sp. Lesson 1831 stony coral

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007