PBDB Collection

Bore Wetten, depth 28m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.9° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Miocene (23.0 - 5.3 Ma)

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1991

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62899: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Anthozoa
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral
Scleractinia - Turbinoliidae	"Cylindrophyllia duncani" = Peponocyathus duncani "Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
Scleractinia - Caryophylliidae	Deltocyathus reussi Deltocyathus reussi stony coral
Scleractinia - Flabellidae	Flabellum sp. Flabellum sp. Lesson 1831 stony coral
Gymnolaemata
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Cheilostomata - Lunulitidae	Lunulites voigti¹ Lunulites voigti¹
Scaphopoda
Dentaliida - Dentaliidae	Lentigodentalium pseudomutabile¹, "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹, Antalis dumasi¹ Lentigodentalium pseudomutabile¹ tusk shell "Pseudantalis aturensis" = Dentalium (Pseudantilis)¹ tusk shell Antalis dumasi¹ tusk shell Antalis dumasi subsp.
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Bivalvia
Arcida - Arcidae	Bathyarca pectunculoides¹, Scapharca (Scapharca) diluvii¹ Bathyarca pectunculoides¹ Scacchi 1834 clam Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Pectinida - Pectinoidae	Pseudamussium lilli¹, Korobkovia woodi¹, Aequipecten angelonii¹ Pseudamussium lilli¹ Pusch 1837 scallop Korobkovia woodi¹ scallop Aequipecten angelonii¹ scallop
Carditida - Astartidae	Astarte (Astarte) radiata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. convexior
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Lucinida - Lucinidae	"Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ "Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ Linnaeus 1767 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Cardiidae	"Habecardium subturgidum" = Laevicardium (Habecardium)¹, Parvicardium straeleni¹ "Habecardium subturgidum" = Laevicardium (Habecardium)¹ cockle H. subturgidum subturgidum Parvicardium straeleni¹ cockle
Cardiida - Veneridae	Venus (Ventricoloidea) multilamella¹, "Gouldia minima" = Gouldia (Gouldia) minima¹ Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam
Cardiida - Kelliellidae	Spaniodontella nitida¹ Spaniodontella nitida¹ clam
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Hiatellida - Hiatellidae	Panopea (Panopea) meynardi¹ Panopea (Panopea) meynardi¹ clam
Nuculanida - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Nuculanida - Yoldiidae	"Portlandia (Yoldiella) pygmaea" = Yoldiella¹ "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Gastropoda
Trochoidea - Trochidae	Calliostoma (Ampullotrochus) ? laureatum² Calliostoma (Ampullotrochus) ? laureatum² top snail
Opisthobranchia - Bullidae	"Tornatina bellardi" = Bulla (Tornatina)³ "Tornatina bellardi" = Bulla (Tornatina)³ snail
Opisthobranchia - Cylichnidae	Cylichna (Cylichna) pseudoconvoluta³, Bullichna paucistriata³ Cylichna (Cylichna) pseudoconvoluta³ snail Bullichna paucistriata³ snail
Opisthobranchia - Retusidae	Volvulella (Volvulella) acuminata³, Retusa (Cylichnina) elongata³ Volvulella (Volvulella) acuminata³ snail Retusa (Cylichnina) elongata³ snail
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Thecosomata - Limacinidae	Limacina miorostralis³, Limacina valvatina³ Limacina miorostralis³ snail Limacina valvatina³ Reuss 1867 snail
Thecosomata - Cavoliniidae	Vaginella austriaca³ Vaginella austriaca³ snail
Heterostropha - Architectonicidae	Architectonica (Pseudotorinia) sensu lato obtusa³ Architectonica (Pseudotorinia) sensu lato obtusa³ sundial
Heterostropha - Pyramidellidae	Kleinella (Kleinella) nordmanni³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros³, Eulimella (Eulimella) acicula³, Eulimella (Eulimella) scillae³, Pyramidella (Voluspa) elata³, Pyramidella (Voluspa) plicosa³, "Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla³, Odostomia (Megastomia) tuexeni³, Syrnola hoernesi³, Syrnola neumayri³, Turbonilla (Mormula) cf. miomutiniensis³, Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³ Kleinella (Kleinella) nordmanni³ snail "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) "sp. 6"" = Pyrgolampros³ Sacco 1892 snail Eulimella (Eulimella) acicula³ snail Eulimella (Eulimella) scillae³ snail Pyramidella (Voluspa) elata³ snail Pyramidella (Voluspa) plicosa³ snail "Turbonilla (Strioturbonilla) densecostata" = Strioturbonilla³ snail sensu JANSSEN Odostomia (Megastomia) tuexeni³ snail Syrnola hoernesi³ snail Syrnola neumayri³ snail Turbonilla (Mormula) cf. miomutiniensis³ snail sensu ANDERSON Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN
Heterostropha - Acteonidae	Acteon (Acteon) semistriatus³, Acteon (Acteon) "sp. 2"³ Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon' Acteon (Acteon) "sp. 2"³ Montfort 1810 snail spelled as 'Actaeon'
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Heterostropha - Mathildidae	sensu lato Mathilda concinna³ sensu lato Mathilda concinna³ snail
Epitonioidea - Epitoniidae	Opalia (Pliciscala) pertusa², "Amaea (Scalina) subreticula" = Scalina², Acirsa (Hemiacirsa) lanceolata², Acirsa (Hemiacirsa) aff. duvergieri², sensu lato Turriscala straeleni² Opalia (Pliciscala) pertusa² wentletrap "Amaea (Scalina) subreticula" = Scalina² wentletrap Acirsa (Hemiacirsa) lanceolata² wentletrap Acirsa (Hemiacirsa) aff. duvergieri² wentletrap sensu lato Turriscala straeleni² wentletrap T. (s. lat.) stareleni f. dingdensis
Vanikoroidea - Eulimidae	Niso (Niso) terebellum², Balcis (Polygyreulima) glabella², Balcis (Balcis) alba², Eulima (Eulima) bilineata², Eulima (Eulima) glabra² Niso (Niso) terebellum² snail Balcis (Polygyreulima) glabella² snail Balcis (Balcis) alba² snail Eulima (Eulima) bilineata² snail Eulima (Eulima) glabra² snail
Cerithioidea - Planaxidae	Couthouyia aff. roberti² Couthouyia aff. roberti² snail
Cerithioidea - Cerithiidae	"Bittium (Bittium) spina" = Bittium spina², Bittium (Bittium) tenuispina² "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail Bittium (Bittium) tenuispina² cerith snail
Cerithioidea - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Sorbeoconcha - Naticidae	Naticidae indet.², Natica koeneni², Tectonatica miopusilla², Euspira gottschei², Euspira nysti², Euspira staringi², Euspira helicina², Neverita josephinia², "Natica sensu lato tigrina" = Cochlis tigrina², "Natica neglecta" = Cochlis neglecta² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. Natica koeneni² moon snail Tectonatica miopusilla² moon snail Euspira gottschei² moon snail Euspira nysti² moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina Neverita josephinia² Risso 1826 moon snail N. josephinia olla "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail "Natica neglecta" = Cochlis neglecta² Mayer 1858 moon snail
Sorbeoconcha - Adeorbidae	Daronia dingdensis² Daronia dingdensis² snail
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus quadricarinatus², Circulus subcirculus², Circulus hennei² Circulus praecedens² snail C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti Circulus quadricarinatus² snail Circulus subcirculus² snail Circulus hennei² Glibert 1952 snail
Sorbeoconcha - Tornidae	Tornus (Tornus) trigonostoma² Tornus (Tornus) trigonostoma² snail
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Columbellidae	Anachis (Costoanachis) hosiusi², Anachis (Costoanachis) terebralis², Mitrella (Macrurella) nassoides² Anachis (Costoanachis) hosiusi² snail Anachis (Costoanachis) terebralis² snail Mitrella (Macrurella) nassoides² snail
Neogastropoda - Nassariidae	Amyclina facki², Nassarius bocholtensis², Nassarius tenuistriatus², Nassarius twistringensis², Nassarius karinae² Amyclina facki² snail Nassarius bocholtensis² snail Nassarius tenuistriatus² snail Nassarius twistringensis² snail Nassarius karinae² Wienrich 2001 snail
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus², Streptodictyon abruptus², ? Exilifusus attenuatus² Streptodictyon sexcostatus² snail Streptodictyon abruptus² snail incl. 1 specimen S. abruptus for. quinquecostatus ? Exilifusus attenuatus² snail
Neogastropoda - Cancellariidae	Sveltia varicosa², "Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta², Babylonella fusiformis² Sveltia varicosa² snail S. varicosa paucicostata "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana Babylonella fusiformis² snail
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Turridae	Fusiturris aquensis³, Fusiturris duchastelii³, Boreodrillia hosiusi³, Nitidiclavus maitrejus³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, Gemmula (Gemmula) vermicularis³, Gemmula (Gemmula) zimmermanni³ Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid F. duchastelii flexiplicata Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid Gemmula (Gemmula) vermicularis³ turrid Gemmula (Gemmula) zimmermanni³ turrid
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Cochlespiridae	Cochlespira corneti³ Cochlespira corneti³ snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³, Terebra (Myurellina) acuminata³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail Terebra (Myurellina) acuminata³ auger snail T. acuminata neglecta
Neogastropoda - Raphitomidae	Favriella sinuosula³, Mioawateria andersoni³, Metuonella grippi³, Raphitoma aff. praehispida³, Teretia anceps³, Teretia "sp. 1"³ Favriella sinuosula³ snail Mioawateria andersoni³ snail Metuonella grippi³ snail Raphitoma aff. praehispida³ snail Teretia anceps³ snail Teretia "sp. 1"³ Norman 1888 snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³, Pleurotomoides simplex³, ? Ceritoturris hannoverana³ Pleurotomoides luisae³ snail Pleurotomoides simplex³ snail ? Ceritoturris hannoverana³ snail
Neogastropoda - Mangeliidae	Sorgenfreispira sorgenfreii³, Sorgenfreispira tenella³, sensu lato Mangelia andersoni³, sensu lato Mangelia miorugulosa³, Benthomangelia aff. obtusangula³ Sorgenfreispira sorgenfreii³ snail Sorgenfreispira tenella³ Conrad 1848 snail sensu lato Mangelia andersoni³ Wienrich and Janssen 2007 snail sensu lato Mangelia miorugulosa³ Kautsky 1925 snail Benthomangelia aff. obtusangula³ snail
Neogastropoda - Borsoniidae	Bathytoma jugleri³, "Genota ramosa" = Genota valeriae³, Asthenotoma pannoides³, Microdrillia aff. serratula³ Bathytoma jugleri³ snail "Genota ramosa" = Genota valeriae³ Hoernes and Auinger 1891 snail Asthenotoma pannoides³ snail Microdrillia aff. serratula³ snail
Neogastropoda - Volutidae	Scaphella (Scaphella) bolli² Scaphella (Scaphella) bolli² volute
Neogastropoda - Muricidae	"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta², "Murex (Haustellum) inornatus" = Haustellum², Lyrotyphis (Eotyphis) fistulosus², Pterynotus (Alipurpura) parvifolius², Typhis (Typhis) ? pungens² "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta² Bronn 1831 murex snail "Murex (Haustellum) inornatus" = Haustellum² murex snail Lyrotyphis (Eotyphis) fistulosus² murex snail Pterynotus (Alipurpura) parvifolius² mustached bat Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Ranellidae	"Charonia (Sassia) tarbelliana" = Sassia² "Charonia (Sassia) tarbelliana" = Sassia² triton shell
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Pseudomelanoidea - Pseudomelaniidae	sensu lato Inquisitor borealis³ sensu lato Inquisitor borealis³ snail
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Nodosariata
Polymorphinida - Polymorphinidae	Guttulina problema, Globulina gibba, Pseudopolymorphina subnodosa Guttulina problema d'Orbigny 1826 Globulina gibba d'Orbigny 1832 Pseudopolymorphina subnodosa Pseudop. subn. for. rugosa
Vaginulinida - Lagenidae	Darbyella nitida Darbyella nitida
Vaginulinida - Vaginulinidae	Lenticulina (Vaginulinopsis) gerlachi, Lenticulina (Robulus) ariminensis, Lenticulina (Robulus) grandis, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) depauperata, Lenticulina (Robulus) ornata, Lenticulina (Lenticulina) inornata, Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) gibba Lenticulina (Vaginulinopsis) gerlachi Lenticulina (Robulus) ariminensis Lenticulina (Robulus) grandis Lenticulina (Robulus) cultrata Lenticulina (Robulus) depauperata L. (L.) depauperata for. costata Lenticulina (Robulus) ornata Lenticulina (Lenticulina) inornata d'Orbigny 1846 Lenticulina (Lenticulina) calcar Lenticulina (Lenticulina) gibba
Nodosariida - Nodosariidae	Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina urnula, Dentalina konincki, Nodosaria bacillum, Nodosaria rudis, Nodosaria grandis, Lenticulina (Robulus) simplex Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina urnula Dentalina konincki Reuss 1861 Nodosaria bacillum Nodosaria rudis Nodosaria grandis Lenticulina (Robulus) simplex d'Orbigny 1846
Foraminifera
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
Rotalioidea - Elphidiidae	Elphidium inflatum Elphidium inflatum
Nodosariacea - Glandulinidae	Glandulina ovula Glandulina ovula
-	"Marginulina cf. costata" = Cristellaria (Marginulina) "Marginulina cf. costata" = Cristellaria (Marginulina)
Globothalamea
Rotaliida - Cibicididae	Cibicides lobatulus, Cibicides dutemplei Cibicides lobatulus Cibicides dutemplei dOrbigny 1846

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007