PBDB Collection

Chapada do Araripe [Romualdo Fm.] (general) (Cretaceous of Brazil)

Also known as Araripe Plateau

Where: Ceará, Brazil (7.2° S, 39.3° W: paleocoordinates 8.7° S, 7.6° W)

• coordinate estimated from map

• basin-level geographic resolution

When: Romualdo Formation (Santana Group), Late/Upper Aptian (122.5 - 113.0 Ma)

• In recent stratigraphic schemes (e.g. Martill 2007) the Santana Formation and the "Romualdo Member" are essentially synonyms

•Santana Formation suggested to be early Albian in age by Heimhofer & Hochuli (2010) on the basis of palynomorphs

•Cerqueira et al. 2021:The Romualdo Formation was recently considered as Aptian in age, based on palynomorph, foraminifera and ostracod data (Arai and Assine 2020; Melo et al. 2020).

•From Melo et al. 2020: In summary, this integrated study of foraminifera, ostracodes and other microfossil data from the Romualdo Formation indicates that the local Alagoas Stage (Ostracoda Zone RT-011) can now be constrained to the Aptian. Arai and Assine 2020 also support that the Romualdo Formation and Santana Group is entirely Aptian in age.

• member-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; concretionary, calcareous shale

• Cerqueira et al. 2021: These layers probably were deposited follow- ing pulses of mass mortality, and the lithologies character- ize a coastal/marine environment (Melo et al. 2020).

• Limestone concretions in black shales

•Cerqueira et al. 2021: characterized by conglomeratic sandstone followed by a sequence of green and black shale (Assine 2007), stratified conglomerates, fine-to medium-grained sandstone, laminated limestone, marl, and coquina levels (Assine et al. 2014; Melo et al. 2020). The black shale is occasionally intercalated with sev- eral layers where carbonate concretions are abundant (Fara et al. 2005; Saraiva et al. 2007; Vila Nova et al. 2011; Kellner et al. 2013).

Size class: macrofossils

Preservation: concretion, soft parts

Reposited in the AMNH, BSP, SMNS

Collection methods: quarrying, chemical

Primary reference: L. I. Price. 1971. A presença de Pterosauria no Cretáceo Inferior da Chapada do Araripe, Brasil. Anais da Academia Brasileira de Ciências 43:451-461 [R. Butler/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 92290: authorized by Richard Butler, entered by Richard Butler on 05.11.2009, edited by Roger Benson, Philip Mannion, Mark Bell, Matthew Carrano, Grace Varnham and Jonathan Tennant

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

unclassified
-	Oviraptorosauridae indet.¹² Oviraptorosauridae indet.¹² SMNS 58023 (last 3 sacral vertebrae & ribs, 2 caudal vertebrae, & postacetabular process of R ilium)
Reptilia
Testudines -	Brasilemys josai n. gen. n. sp.²⁰ Brasilemys josai n. gen. n. sp.²⁰ Lapparent de Broin 2000 sideneck turtle Museu de Geologia de Barcelona, Catalunya, Spain, number MGB 37911. A carapace with a partial skull, two hyoid branches, left lower jaw; axis and 3d cervical vertebrae in the carapace
Theropoda - Spinosauridae	Spinosaurinae indet.³ Spinosaurinae indet.³ Sereno et al. 1998 tetanuran theropod LPP-PV-0042, a fragmentary tibia
Theropoda -	Megaraptora indet.² Megaraptora indet.² Benson et al. 2010 coelurosaur SMNS 58023 (last 3 sacral vertebrae & ribs, 2 caudal vertebrae, & postacetabular process of R ilium)
Pterosauria -	Pterodactyloidea indet.²³, Pterodactyloidea indet.¹⁹, Araripedactylus dehmi n. gen. n. sp.³⁰ Pterodactyloidea indet.²³ pterosaur DNPM D6M 488-LE, wing bones. AMNH 22569, pelvis. Geol. Inst. Univ. Amsterdam 4895, two cervical vertebrae. Gigase collection V-201, 10 articulated posterior cervical and anterior dorsal vertebrae. BSP 1987 I 65, right scapulocoracoid. SMNS 55408, humerus. SMNS 82001, ulna. SMNS 81976, humerus and ulna/radius. SMNS 55412, 55415, phalanges. SMNS 80437 Pterodactyloidea indet.¹⁹ pterosaur BSP 1982 I 91, 1982 I 93 Araripedactylus dehmi n. gen. n. sp.³⁰ Wellnhofer 1977 pterosaur BSPG 1975 I 166, proximal phalanx, right wing-finger
Pterosauria - Anhangueridae	"cf. Coloborhynchus spielbergi" = Maaradactylus spielbergi²⁶, "Coloborhynchus spielbergi n. sp." = Maaradactylus spielbergi²⁸, "cf. Criorhynchus mesembrinus" = Tropeognathus mesembrinus²⁶, "Tropeognathus mesembrinus n. gen. n. sp." = Tropeognathus mesembrinus³² "cf. Coloborhynchus spielbergi" = Maaradactylus spielbergi²⁶ Veldmeijer 2003 pterosaur SMNS 55413, ulna "Coloborhynchus spielbergi n. sp." = Maaradactylus spielbergi²⁸ Veldmeijer 2003 pterosaur RGM 401 880, skull and postcrania "cf. Criorhynchus mesembrinus" = Tropeognathus mesembrinus²⁶ Wellnhofer 1987 pterosaur SMNS 56994, mandible "Tropeognathus mesembrinus n. gen. n. sp." = Tropeognathus mesembrinus³² Wellnhofer 1987 pterosaur BSP 1987 I 46, skull and lower jaw
Pterosauria -	Cearadactylus atrox n. gen. n. sp.²¹, Brasileodactylus sp.²⁷, Brasileodactylus araripensis n. gen. n. sp.¹⁵, Anhanguera sp.²⁹, "Araripesaurus santanae n. sp." = Anhanguera santanae³¹, "? Cearadactylus ligabuei n. sp." = Anhanguera ligabuei⁹, "Tropeognathus robustus n. sp." = Anhanguera robustus³², "Santanadactylus araripensis n. sp." = Anhanguera araripensis³¹, "Santanadactylus cf. araripensis" = Anhanguera araripensis³¹, Anhanguera blittersdorffi n. gen. n. sp.⁷ Cearadactylus atrox n. gen. n. sp.²¹ Leonardi and Borgomanero 1985 pterosaur F-PV-93 (holotype), incomplete skull with mandibles Brasileodactylus sp.²⁷ Kellner 1984 pterosaur AMNH 24444, skull and partial left wing Brasileodactylus araripensis n. gen. n. sp.¹⁵ Kellner 1984 pterosaur MN 4804–V, anterior part of mandible Anhanguera sp.²⁹ Campos and Kellner 1985 pterosaur SAO 200602, mandible "Araripesaurus santanae n. sp." = Anhanguera santanae³¹ Wellnhofer 1985 pterosaur BSP 1982 I 90, partial skull and partial forelimb "? Cearadactylus ligabuei n. sp." = Anhanguera ligabuei⁹ Dalla Vecchia 1993 pterosaur CSRL 12692/12713 (holotype), anterior part of a skull "Tropeognathus robustus n. sp." = Anhanguera robustus³² Wellnhofer 1987 pterosaur BSP 1987 I 47, lower jaw "Santanadactylus araripensis n. sp." = Anhanguera araripensis³¹ Wellnhofer 1985 pterosaur BSP 1982 I 89, skull, lower jaws, partial forelimb. BSP 1987 I 66, right metacarpal. SMNS 55409, humerus. SAO 16494, skull "Santanadactylus cf. araripensis" = Anhanguera araripensis³¹ Wellnhofer 1985 pterosaur BSP 1982 I 92, forelimb material Anhanguera blittersdorffi n. gen. n. sp.⁷ Campos and Kellner 1985 pterosaur Blittersdorff collection, complete skull
Pterosauria - Pteranodontidae	Santanadactylus brasilensis n. gen. n. sp.⁶, Santanadactylus spixi n. sp.³¹ Santanadactylus brasilensis n. gen. n. sp.⁶ Buisonjé 1980 pterosaur Geol. Inst. Univ. Amsterdam 4894 (holotype), humerus/scapula/coracoid Santanadactylus spixi n. sp.³¹ Wellnhofer 1985 pterosaur BSP 1980 I 121, partial forelimb
Pterosauria -	Azhdarchoidea indet.¹¹ Azhdarchoidea indet.¹¹ Unwin 1992 pterosaur SMNK PAL 3985, MPSC R 859 (prob. different taxa)
Pterosauria - Thalassodromidae	Tupuxuara longicristatus n. gen. n. sp.¹⁸ Tupuxuara longicristatus n. gen. n. sp.¹⁸ Kellner and Campos 1988 pterosaur
Pterosauria - Tapejaridae	Thalassodrominae indet.¹, Kariridraco dianae n. gen. n. sp.⁸, Tapejara wellnhoferi n. gen. n. sp.¹⁷ Thalassodrominae indet.¹ Kellner and Campos 2007 pterosaur AMNH 22567, three posteriormost cervical vertebrae, first seven dorsals, (fused in to a notarium), right and left scapulocoracoids, a partial sternum, incomplete right humerus, part of the wing finger, incomplete right femur and parts of the right tibia/fibula Kariridraco dianae n. gen. n. sp.⁸ Cerqueira et al. 2021 pterosaur Holotype: MPSC R 1056, fairly complete skull, preserving most of its temporal, orbital, and preorbital portions, part of its premax- illary crest, an almost complete lower jaw and four cervical vertebrae Tapejara wellnhoferi n. gen. n. sp.¹⁷ Kellner 1989 pterosaur Blittersdorff collection, CD-R-080, partial skull
Pterosauria - Dsungaripteridae	Banguela oberlii n. gen. n. sp.¹³ Banguela oberlii n. gen. n. sp.¹³ Headden and Campos 2014 pterosaur SAO 251093, mandible
Pterosauria -	Araripesaurus castilhoi n. gen. n. sp. Araripesaurus castilhoi n. gen. n. sp. Price 1971 pterosaur 529-R, partial forelimb
Pterosauria - Ornithocheiridae	Barbosania gracilirostris n. gen. n. sp.¹⁰ Barbosania gracilirostris n. gen. n. sp.¹⁰ Elgin and Frey 2011 pterosaur MNHS/00/85, skull and partial postcranium
Pterosauria -	Santanadactylus pricei n. sp.³¹, cf. Santanadactylus pricei²⁶ Santanadactylus pricei n. sp.³¹ Wellnhofer 1985 pterosaur BSP 1980 I 122 (holotype), partial left forelimb. Paratypes: BSP 1980 I 43, 1980 I 120, MZ A/III 522, forelimb material. BSP 1987 I 1, partial left forelimb cf. Santanadactylus pricei²⁶ Wellnhofer 1985 pterosaur SMNS 55883, humerus, 55410, ulna, 55411, ulna and radius
Loricata - Peirosauridae	Caririsuchus camposi n. gen. n. sp.¹⁶ Caririsuchus camposi n. gen. n. sp.¹⁶ Kellner 1987 crocodilian CD-R-041 & DGM 1468 R - partial skeleton
Loricata -	Araripesuchus gomesii¹⁴ Araripesuchus gomesii¹⁴ Price 1959 crocodilian AMNH 24450 - complete articulated skeleton
Actinopteri
Lepisosteiformes -	Lepidotes wenzae⁵ Lepidotes wenzae⁵ Brito and Gallo 2003 gar Holotype: MNHN-BCE 387
Lepisosteiformes - Obaichthyidae	Obaichthys decoratus n. gen. n. sp.³³, Obaichthys ? laevis³³ Obaichthys decoratus n. gen. n. sp.³³ Wenz and Brito 1992 gar DGM 1336 Obaichthys ? laevis³³ Wenz and Brito 1992 gar DGM 1155
Pycnodontiformes - Pycnodontidae	Microdon penalvai n. sp.²⁵ Microdon penalvai n. sp.²⁵ Silva Santos 1970 Holotype: No. 918, DGM - DNPM
Osteichthyes
Halecomorphi - Ionoscopidae	Oshunia brevis³⁴ Oshunia brevis³⁴ Wenz and Kellner 1986 ray-finned fish CD-P-31
Chondrichthyes
Rajiformes - Rhinobatidae	Rhinobatos beurleni n. sp.²⁴ Rhinobatos beurleni n. sp.²⁴ Silva Santos 1968 guitarfish Holotype: No. 2510 D e V; Paratype: No. 917-P, D e V
Hybodontiformes - Hybodontidae	Tribodus limae n. gen. n. sp.⁴ Tribodus limae n. gen. n. sp.⁴ Brito and Ferreira 1989 elasmobranch Holotype: GP/2T-2 IG-USP
Santanaclupea silvasantosi
-	Santanaclupea silvasantosi n. gen. n. sp.²² Santanaclupea silvasantosi n. gen. n. sp.²² Maisey 1993 AMNH 12790, complete specimen; AMNH 12789

¹Aires et al. 2013; ²Aranciaga Rolando et al. 2018; ³Aureliano et al. 2018; ⁴Brito and Ferreira 1989; ⁵Brito and Gallo 2003; ⁶Buisonjé 1980; ⁷Campos and Kellner 1985; ⁸Cerqueira et al. 2021; ⁹Dalla Vecchia 1993; ¹⁰Elgin and Frey 2011; ¹¹Elgin and Hone 2013; ¹²Frey and Martill 1995; ¹³Headden and Campos 2014; ¹⁴Hecht 1991; ¹⁵Kellner 1984; ¹⁶Kellner 1987; ¹⁷Kellner 1989; ¹⁸Kellner and Campos 1988; ¹⁹Kellner and Tomida 2000; ²⁰Lapparent de Broin 2000; ²¹Leonardi and Borgomanero 1985; ²²Maisey 1993; ²³Martill and Unwin 1989; ²⁴Silva Santos 1968; ²⁵Silva Santos 1970; ²⁶Veldmeijer 2002; ²⁷Veldmeijer 2003; ²⁸Veldmeijer 2003; ²⁹Veldmeijer et al. 2005; ³⁰Wellnhofer 1977; ³¹Wellnhofer 1985; ³²Wellnhofer 1987; ³³Wenz and Brito 1992; ³⁴Wenz and Kellner 1986