Soyana (Iva Gora, PIN collection 94) (Permian of Russian Federation)

Where: Russian Federation (65.6° N, 43.6° E: paleocoordinates 33.2° N, 33.0° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size class: macrofossils

Collected by M. Edemskij in 1934

Primary reference: F. R. Schram. 1980. Miscellaneous late Paleozoic Malacostraca of the Soviet Union. Journal of Paleontology 54(3):542-547 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 96860: authorized by Matthew Clapham, entered by Matthew Clapham on 05.07.2010, edited by Jered Karr and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Bivalvia
 Pectinida - Pseudomonotidae
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria
"Pseudomonotis speluncaria" = Pseudomonotis (Pseudomonotis) speluncaria von Schlotheim 1816 scallop
 Pectinida - Streblochondriidae
"Pecten sericeum" = Streblochondria sericea
"Pecten sericeum" = Streblochondria sericea de Verneuil 1845 scallop
 Ostreida - Pinnidae
Aviculopinna ivanisciana
Lingulata
 Lingulida - Lingulidae
Lingula orientalis, Lingula credneri
Lingula credneri Geinitz 1848
Rhynchonellata
 Rhynchonellida - Rhynchoporidae
Rhynchopora geinitziana
Rhynchopora geinitziana de Verneuil 1845
Strophomenata
 Productida - Linoproductidae
"Productus cancrini" = Cancrinella cancrini
"Productus cancrini" = Cancrinella cancrini de Verneuil 1845
Malacostraca
 Isopoda - Palaeophreatoicidae
Palaeophreatoicus sojanensis n. gen. n. sp.
Palaeophreatoicus sojanensis n. gen. n. sp. Birshtein 1962 isopod
Insecta
 Eoblattida - Bardapteridae
Olgaephilus emendatus18
Olgaephilus emendatus18 Storozhenko 1990 insect
PIN 94/120(121), 133(134)
 Eoblattida - Megakhosaridae
"Megakhosarina minuscula" = Blattokhosara minuscula1
"Megakhosarina minuscula" = Blattokhosara minuscula1 Aristov 2008 insect
PIN 94/131, 338 (and/or 238)
 Reculida - Ivapteridae
Ivaptera sharovi2
Ivaptera sharovi2 Aristov 2009 winged insect
PIN 94/50,132, 580, 660, 748, 834,1044,1048
 Hypoperlida - Hypoperlidae
Boreopsocus danksae16
Boreopsocus danksae16 Shcherbakov 1994 winged insect
PIN 94/197, 1299, 662
 Hemiptera - Boreoscytidae
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7
"Archescytinopsis flexuosa n. sp." = Boreoscyta nefasta7 Becker-Migdisova 1949 true bug
PIN 94/1017, 775
 Hemiptera - Archescytinidae
Archescytina fusca8, "Sojanoscytina tetraclada" = Archescytina tetraclada8, Sojanoscytina ramosa n. sp.8, Sojanoscytina ivagorae n. sp.8
Archescytina fusca8 Becker-Migdisova 1960 true bug
PIN 94/474, 107
"Sojanoscytina tetraclada" = Archescytina tetraclada8 Martynov 1933 true bug
PIN 94/1185
Sojanoscytina ramosa n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/1182
Sojanoscytina ivagorae n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/887
 Hemiptera - Prosbolopseidae
Prosbolopsis ovalis8, Anaprosbole ivensis8, ? Scytoneura sp.8, Scytoneura elliptica8
Prosbolopsis ovalis8 Martynov 1935 true bug
PIN 94/420, 254, 251, 1184, 171
Anaprosbole ivensis8 Becker-Migdisova 1960 true bug
PIN 94/464
? Scytoneura sp.8 Martynov 1935 true bug
PIN 94/641
Scytoneura elliptica8 Martynov 1935 true bug
PIN 94/243, 450, 473, 480, 483, 577, 596, 1191, 1192, 1283
 Hemiptera - Prosbolidae
Prosbole sojanensis5, Prosbole longialata n. sp.5, Prosbole indistincta n. sp.5, Prosbole brevialata n. sp.5, Prosbole incerta n. sp.5, Prosbole ivagorae5, Prosbole permocicadoides5, Permocicada simulatrix5, Permocicada pusilla n. sp.5, Permocicada triramosa n. sp.5, Permocicada nigronervosa5, Permocicada idelensis5, Permocicada sublata n. sp.5, Permocicada integra n. sp.5, Sojanoneura kazanensis5, Sojanoneura triangularis5, Sojanoneura bimaculata5, Sojanoneura furcatula5, Permocicadopsis angustata5, Permocicadopsis aberrans5
Prosbole sojanensis5 Martynov 1935 true bug
PIN 94/1174(467)
Prosbole longialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1176
Prosbole indistincta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/942
Prosbole brevialata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/763(1190)
Prosbole incerta n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/902
Prosbole ivagorae5 Becker-Migdisova 1940 true bug
PIN 94/255(475)
Prosbole permocicadoides5 Becker-Migdisova 1940 true bug
PIN 94/943, 253
Permocicada simulatrix5 Martynov 1935 true bug
PIN 94/465, 1189
Permocicada pusilla n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/688
Permocicada triramosa n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1196
Permocicada nigronervosa5 Martynov 1928 true bug
PIN 94/249
Permocicada idelensis5 Zalessky 1929 true bug
PIN 94/250(451)
Permocicada sublata n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/1055(1043), 179
Permocicada integra n. sp.5 Becker-Migdisova 1940 true bug
PIN 94/906ab
Sojanoneura kazanensis5 Zalessky 1929 true bug
PIN 94/466(452), 933
Sojanoneura triangularis5 Martynov 1935 true bug
PIN 94/953, 1194, 640
Sojanoneura bimaculata5 Martynov 1935 true bug
PIN 94/458, 639ab, 921, 862
Sojanoneura furcatula5 Martynov 1935 true bug
PIN 94/624(586), 579, 810
Permocicadopsis angustata5 Martynov 1935 true bug
PIN 94/645, 964; 468, 1047, 389, 886(var. typica)
Permocicadopsis aberrans5 Martynov 1935 true bug
PIN 94/462, 8885
 Hemiptera - Coleoscytidae
Coleoscyta occallata n. sp.8
Coleoscyta occallata n. sp.8 Becker-Migdisova 1960 planthopper
PIN 94/1178(482)
 Hemiptera - Ingruidae
Ingruo lanceolata n. gen. n. sp.8
Ingruo lanceolata n. gen. n. sp.8 Becker-Migdisova 1960 true bug
PIN 94/455
 Hemiptera - Scytinopteridae
Scytinoptera kokeni6, Scytinoptera reducta6
Scytinoptera kokeni6 Handlirsch 1904 true bug
PIN 94/481, 875
Scytinoptera reducta6 Martynov 1928 true bug
PIN 94/1179, 665, 469, 975, 472, 461, 408, 493, 922, 638, 245, 1251, 395, 881, 257, 585, 431, 784, 731, 912
 Paoliida - Paoliidae
Sojanidelia striata20, Sojanidelia fluctuosa20, Sojanidelia nana n. sp.20, Sojanidelia makarkini20, Permotermopsis pectinata20, Mezenalicula connata n. gen. n. sp.3, Stenaropodites sojanensis20
Sojanidelia striata20 Storozhenko 1992 winged insect
PIN 94/559(814), 780(1132)
Sojanidelia fluctuosa20 Storozhenko 1992 winged insect
PIN 94/675(934)
Sojanidelia nana n. sp.20 Storozhenko 1992 winged insect
PIN 94/1122(1128), 1133
Sojanidelia makarkini20 Storozhenko 1992 winged insect
PIN 94/370, 377
Permotermopsis pectinata20 Martynov 1937 winged insect
PIN 94/115(6)
Mezenalicula connata n. gen. n. sp.3 Aristov and Storozhenko 2011 winged insect
PIN 94/883
Stenaropodites sojanensis20 Storozhenko 1992 winged insect
PIN 94/1126
 Cnemidolestodea - Sylvabestiidae
"Permula edemskii n. sp." = Sojanopermula edemskii3, Sojanopermula lucida n. gen. n. sp.19
"Permula edemskii n. sp." = Sojanopermula edemskii3 Aristov and Storozhenko 2011 winged insect
PIN 94/1027
Sojanopermula lucida n. gen. n. sp.19 Storozhenko 1992 winged insect
PIN 94/117
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis4
Camptoneurites soyanensis4 Aristov et al. 2010 ice crawler
PIN 94/29
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis11
Kamopanorpa rotundipennis11 Martynov 1932 winged insect
PIN 94/771
 Mecoptera - Permochoristidae
Petrochorista lepida11, Petrochorista sojanensis n. sp.11, Petromantis rossica11, "Permochorista maculipennis" = Petromantis maculipennis11, "Permochorista venosa" = Petromantis maculipennis11, "Petrochorista dubia" = Petromantis dubia11, Agetopanorpa sp.14, "Agetochorista tillyardiana" = Agetopanorpa tillyardiana11, "Agetochorista dubia" = Agetopanorpa dubia11, "Agetochorista letopalae" = Agetopanorpa letopalae11, "Agetochorista ornata" = Agetopanorpa ornata11, "Agetochorista umbrata" = Agetopanorpa ornata11, "Agetochorista splendida" = Agetopanorpa ornata11
Petrochorista lepida11 Martynova 1948 scorpionfly
PIN 94/754
Petrochorista sojanensis n. sp.11 Martynova 1948 scorpionfly
PIN 94/1151
Petromantis rossica11 Handlirsch 1904 scorpionfly
PIN 94/439, 501, 531, 746, 758, 861, 911; 94/1230 ("Soyana")
"Permochorista maculipennis" = Petromantis maculipennis11 Martynov 1932 scorpionfly
PIN 94/142, 167, 419, 592_589), 768, 1152, 1210
"Permochorista venosa" = Petromantis maculipennis11 Martynov 1932 scorpionfly
PIN 94/900, 1060, 1205
"Petrochorista dubia" = Petromantis dubia11 Martynov 1932 scorpionfly
PIN 94/681, 730, 978, 1198
Agetopanorpa sp.14 Carpenter 1930 scorpionfly
PIN 94/175
"Agetochorista tillyardiana" = Agetopanorpa tillyardiana11 Martynova 1948 scorpionfly
PIN 94/893, 909
"Agetochorista dubia" = Agetopanorpa dubia11 Martynov 1932 scorpionfly
PIN 94/179, 185, 540, 705 (2 wings), 827
"Agetochorista letopalae" = Agetopanorpa letopalae11 Martynova 1948 scorpionfly
PIN 94/1153
"Agetochorista ornata" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/149, 164, 168, 170, 174, 176, 516, 538, 778, 980, 1139, 1157, 1211, 1213, 1312
"Agetochorista umbrata" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/1204, 1213
"Agetochorista splendida" = Agetopanorpa ornata11 Martynov 1932 scorpionfly
PIN 94/165, 193, 595, 598, 671, 708, 709, 968, 1138; var. furcata 94/238
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata11, "Permopanorpa tenuis" = Permopanorpa martynovi11
"Permopanorpa angustata" = Martynopanorpa angustata11 Martynov 1932 scorpionfly
PIN 94/169, 515, 553, 620
"Permopanorpa tenuis" = Permopanorpa martynovi11 Willmann 1978 scorpionfly
PIN 94/544
 Raphidioptera - Parasialidae
Parasialis dissedens n. sp.15
Parasialis dissedens n. sp.15 Ponomarenko 1977 snakefly
PIN 94/1217, 820
 Dicondylia - Permithonidae
"Permithonopsis triramosa n. sp." = Permegalomus maculipennis12, "Permithonopsis proxima" = Permithonopsis ivensis12, Permithonopsis obscura n. sp.12
"Permithonopsis triramosa n. sp." = Permegalomus maculipennis12 Martynov 1932 winged insect
PIN 94/172
"Permithonopsis proxima" = Permithonopsis ivensis12 Martynov 1932 winged insect
PIN 94/152
Permithonopsis obscura n. sp.12 Martynova 1952 winged insect
PIN 94/173(726)
 Dicondylia - Permosialidae
"Permosialis cauleoides n. sp." = Permosialis paucinervis12, "Permosialis marmorata n. sp." = Permosialis paucinervis12, "Permosialis bifasciata" = Permosialis paucinervis12, Permosialis latiformis12
"Permosialis cauleoides n. sp." = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/798
"Permosialis marmorata n. sp." = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/186(192), 721(676)
"Permosialis bifasciata" = Permosialis paucinervis12 Martynov 1928 winged insect
PIN 94/497, 576, 1212
Permosialis latiformis12 Martynova 1952 winged insect
PIN 94/536
 Dicondylia - Permembiidae
Letopaloptera carpenteriana n. sp.13
Letopaloptera carpenteriana n. sp.13 Martynova 1961 winged insect
PIN 94/561
 Glosselytrodea - Jurinidae
Jurina reducta10, Jurina marginata10, Jurina sojanensis10
Jurina reducta10 Martynov 1938 winged insect
PIN 94/549
Jurina marginata10 Martynov 1938 winged insect
PIN 94/352, 369, 403, 409, 432, 549, 807, 937, 961
Jurina sojanensis10 Martynov 1938 winged insect
PIN 94/555, 865; 864, 868 form radialis
 Glosselytrodea - Glosselytridae
Glosselytron multivenosum10
Glosselytron multivenosum10 Martynov 1938 winged insect
PIN 94/276
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma17, Eukuloja cubitalis17
Eukuloja atrosumma17 Sinitshenkova 1981 winged insect
PIN 94/935, 1158
Eukuloja cubitalis17 Martynov 1932 winged insect
PIN 94/3, 7, 8, 10, 11, 13, 20, 710, 737, 1164
 Dicondylia - Kennedyidae
Kennedya ivensis9
Kennedya ivensis9 Nel et al. 2012 winged insect
PIN 94/2
1Aristov 2008; 2Aristov 2009; 3Aristov and Storozhenko 2011; 4Aristov et al. 2010; 5Becker-Migdisova 1940; 6Becker-Migdisova 1948; 7Becker-Migdisova 1949; 8Becker-Migdisova 1960; 9Felker 2023; 10Martynov 1938; 11Martynova 1948; 12Martynova 1952; 13Martynova 1961; 14Novokshonov 1994; 15Ponomarenko 1977; 16Shcherbakov 1994; 17Sinitshenkova 1981; 18Storozhenko 1990; 19Storozhenko 1992; 20Storozhenko 1992