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Dryosaurus

Reptilia - Dryosauridae

Taxonomy
Dryosaurus was named by Marsh (1894). It is the type genus of Dryosauridae, Dryosaurinae.

It was synonymized subjectively with Laosaurus by Romer (1956) and Romer (1966).

It was assigned to Ornithopoda by Marsh (1894); to Kalodontidae by Nopcsa (1902); to Nanosauridae by Huene (1909); to Orthopoda by Lull (1911); to Hypsilophontidae by von Zittel (1911); to Camptosauridae by Hay (1902), Lull (1911) and Lull (1924); to Camptosaurinae by Nopcsa (1928); to Laosauridae by Marsh (1895), Marsh (1896), Huene (1914), Osborn (1923), Gilmore (1925) and Tatarinov (1964); to Iguanodontoidea by Norman (1984); to Dryosaurinae by Cooper (1985); to Iguanodontia by Norman (1989); to Hypsilophodontidae by Kuhn (1964), Galton (1972), Galton and Jensen (1973), Galton and Jensen (1973), Galton (1973), Thulborn (1974), Galton (1974), Galton (1974), Thulborn (1975), Galton (1977), Jurcsak and Popa (1979), Sues (1980), Galton (1981), McIntosh (1981), Colbert (1981), Jurcsák (1982), Weishampel and Weishampel (1983), Galton (1983), Russell (1984), Martill (1988), Galton (1989), Miller et al. (1991) and Grigorescu (1992); to Mesoeucrocodylia by Brochu et al. (2002); to Dryomorpha by Sereno (1986), Weishampel and Heinrich (1992) and Novas et al. (2004); and to Dryosauridae by Stefano (1903), Milner and Norman (1984), Carroll (1988), Sues and Norman (1990), Chure et al. (1994), Ryan (1997), Sereno (1997), Sereno (1998), Norman (2004), Stanford et al. (2004), Jouve (2005), Jouve et al. (2005), Chure et al. (2006), Pereda Suberbiola et al. (2006), Sánchez-Hernández et al. (2007), Ruiz-Omeñaca et al. (2007), Galton (2007), Butler et al. (2008), McDonald et al. (2010), Mateus and Milàn (2010), Ruiz-Omeñaca et al. (2010) and Madzia et al. (2018).

Synonymy list
YearName and author
1894Dryosaurus Marsh
1895Dryosaurus Marsh
1896Dryosaurus Marsh pp. 243-244
1902Dryosaurus Hay p. 501
1902Dryosaurus Nopcsa p. 152
1903Dryosaurus Stefano p. 54
1909Dryosaurus Huene p. 16
1911Dryosaurus Lull p. 177
1911Dryosaurus von Zittel p. 289
1914Dryosaurus Huene p. 41
1923Dryosaurus Osborn
1924Dryosaurus Lull p. 239
1925Dryosaurus Gilmore p. 400
1928Dryosaurus Nopcsa p. 184
1964Dryosaurus Kuhn p. 7
1964Dryosaurus Tatarinov p. 555
1972Dryosaurus Galton p. 465
1973Dryosaurus Galton p. 17
1973Dryosaurus Galton and Jensen p. 129
1974Dryosaurus Galton p. 150
1974Dryosaurus Thulborn p. 171
1975Dryosaurus Thulborn p. 123
1977Dryosaurus Galton p. 230
1979Dryosaurus Jurcsak and Popa p. 40
1980Dryosaurus Sues p. 70
1981Dryosaurus Colbert p. 50
1981Dryosaurus Galton p. 273
1981Dryosaurus McIntosh p. 30
1982Dryosaurus Jurcsák p. 179
1983Dryosaurus Galton p. 232
1983Dryosaurus Weishampel and Weishampel p. 44
1984Dryosaurus Milner and Norman p. 146
1984Dryosaurus Norman pp. 342-344 fig. 7
1984Dryosaurus Russell p. 23
1985Dryosaurus Cooper p. 295
1986Dryosaurus Sereno
1988Dryosaurus Carroll
1988Dryosaurus Martill p. 183
1989Dryosaurus Galton p. 218
1989Dryosaurus Norman p. 61 fig. 1
1990Dryosaurus Sues and Norman p. 500
1991Dryosaurus Miller et al. p. 40
1992Dryosaurus Grigorescu p. 147
1992Dryosaurus Weishampel and Heinrich p. 166
1994Dryosaurus Chure et al. p. 304
1997Dryosaurus Ryan p. 197
1997Dryosaurus Sereno p. 449
1998Dryosaurus Sereno p. 61
2002Dryosaurus Brochu et al. p. 1060
2004Dryosaurus Norman p. 414
2004Dryosaurus Novas et al. p. 79 fig. 2C
2004Dryosaurus Stanford et al. p. 257
2005Dryosaurus Jouve p. 324
2005Dryosaurus Jouve et al.
2006Dryosaurus Chure et al. p. 237
2006Dryosaurus Pereda Suberbiola et al. p. 227
2007Dryosaurus Galton p. 38
2007Dryosaurus Ruiz-Omeñaca et al. p. 8
2007Dryosaurus Sánchez-Hernández et al. p. 203
2008Dryosaurus Butler et al. p. 8
2010Dryosaurus Mateus and Milàn p. 254
2010Dryosaurus McDonald et al. p. 33 fig. 39
2010Dryosaurus Ruiz-Omeñaca et al. p. 84
2018Dryosaurus Madzia et al. p. 973 fig. 4

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
RankNameAuthor
Diapsida()
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Neornithischia()
Ornithopoda()
Iguanodontia()
Dryomorpha
familyDryosauridae
genusDryosaurus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yesg
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Maximum body mass: 300 kgg
Minimum body mass: 100 kgg
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2005-08-25 05:58:40
Modified: 2005-08-25 07:58:40
Source: g = genus, subo = suborder, o = order
References: Kiessling 2004, Benton 1983, Peczkis 1995, Marsh 1875

Age range: base of the Late/Upper Kimmeridgian to the top of the Early/Lower Tithonian or 155.70000 to 145.00000 Ma

Collections (23 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Oxfordian - Late/Upper Kimmeridgian161.2 - 150.8Portugal (Estremdura) Dryosauridae indet. (38680)
Kimmeridgian157.3 - 152.1USA (Utah) D. sp. (57913)
Kimmeridgian157.3 - 152.1USA (Wyoming) D. sp. (46465 47065 49577)
Kimmeridgian157.3 - 152.1USA (Colorado) D. altus (53041)
Kimmeridgian - Tithonian157.3 - 145.0USA (Utah) D. sp. (22711)
Kimmeridgian - Tithonian157.3 - 145.0USA (Colorado) D. altus (25141) D. sp. (48161 53022)
Kimmeridgian - Tithonian157.3 - 145.0USA (Wyoming) D. altus (39253 56167 142334) D. sp. (49636 55487) Laosaurus altus (39344) Laosaurus sp. (22648 39340)
Early/Lower Kimmeridgian155.7 - 150.8France (Normandie) D. sp. (35968)
Early/Lower Kimmeridgian155.7 - 150.8USA (Colorado) D. sp. (48156)
Early/Lower Kimmeridgian155.7 - 150.8Germany (Nordrhein-Westfalen) D. sp. (182492)
Late/Upper Kimmeridgian - Early/Lower Tithonian155.7 - 145.0USA (Colorado) D. sp. (45975)
Early/Lower Tithonian150.8 - 145.0USA (Utah) D. altus, Laosaurus gracilis (21852)