PBDB Taxon

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Ellipsocephalidae

Trilobita - Redlichiida - Ellipsocephalidae

Taxonomy

Ellipsocephalidae was named by Matthew (1887). It is not extant. It is the type family of Ellipsocephaloidea.

It was assigned to Ptychopariida by Whittington et al. (1997) and Jell and Adrain (2002); and to Ellipsocephaloidea by Landing et al. (2008) and Adrain (2011).

Subtaxa

Acadolenus, Alueva, Antatlasia, Argunaspis, Asiatella, Bergeroniaspis, Bergeroniellus, Bhargavia, Blayacina (syn. Sardaspis), Brevitermierella (syn. Paratermierella), Cambrunicornia, Catadoxides, Comluella, Culmenaspis, Ellipsocephalinae, Ellipsostrenua, Geyerorodes (syn. Orodes), Glabrella, Hamatolenus, Hupeolenus, Issafeniella, Kadyella, Kameschkoviella, Kijanella, Kingaspidoides (syn. Elatius), Kingaspis (syn. Mesetaia), Krolina, Kymataspis, Latikingaspis, Latoucheia, Latuzella, Lermontovia, Limataceps, Lusatiops, Myopsolenus (syn. Collyrolenus), Nelegeria, Olekmaspis, Ourikaia, Paramicmacca, Paraprotolenella, Planolimbus, Protagraulos, Protolenella, Protolenidae, Protolenus (syn. Bergeronia), Pruvostinoides, Pseudokadyella, Pseudolenus, Pseudoprotolenella, Ptychoparopsis (syn. Berabichia), Rinconia, Sailycaspis, Sectigena, Strenuella, Tadakoustia, Termieraspis, Termierella, Triangulaspis (syn. Angustaeva, Acutaspis, Triangulina), Yeshanaspis

Sister taxa

Protolenidae

Synonyms

Protolenidae was named by Richter and Richter.

It was considered an invalid subgroup of Ellipsocephalidae by Adrain (2011).

It was assigned to Redlichiida by Palmer and Gatehouse (1972).

Synonymy list

p. 15

Year	Name and author
1887	Ellipsocephalidae Matthew
1972	Protolenidae Palmer and Gatehouse p. 15
1997	Ellipsocephalidae Whittington et al. p. 302
2002	Ellipsocephalidae Jell and Adrain p. 470
2008	Ellipsocephalidae Landing et al. p. 893
2011	Ellipsocephalidae Adrain p. 104

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829

Rank	Name	Author
subphylum	Artiopoda	(Hou and Bergstrom)
class	Trilobita	Walch 1771
order	Redlichiida	Richter 1932
suborder	Redlichiina	Richter 1932
superfamily	Ellipsocephaloidea	Lin 1990
family	Ellipsocephalidae	Matthew 1887
family	Ellipsocephalidae	Matthew 1887

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Ellipsocephalidae Matthew 1887

show all | hide all

G. †Acadolenus Sdzuy 1967

G. †Alueva Lotze 1958

G. †Antatlasia Hupe 1952

G. †Argunaspis Repina 1966

G. †Asiatella Repina 1964

G. †Bergeroniaspis Lermontova 1951

G. †Bergeroniellus Lermontova 1940

G. †Bhargavia Peng et al. 2009

†Bhargavia prakritika Peng et al. 2009

G. †Blayacina Cobbold 1932

Invalid names: Sardaspis Brasier 1976 [synonym]

G. †Brevitermierella Hupé 1953

Invalid names: Paratermierella Hupe 1952 [synonym]

G. †Cambrunicornia Geyer 1990

G. †Catadoxides Matthew 1899

G. †Comluella Hupe 1953

G. †Culmenaspis Repina 1972

Subfm. †Ellipsocephalinae Mathew 1887

G. †Ellipsocephalus Zenker 1833

†Ellipsocephalus gripi Kautsky 1945

Invalid names: Germaropyge Snajdr 1957 [synonym]

G. †Ornamentaspis Geyer 1990

†Ornamentaspis angustigena Geyer 2014

†Ornamentaspis crassilimbata Geyer 1990

G. †Strenuaeva Richter and Richter 1940

†Strenuaeva inflata Ahlberg and Bergström 1978

Invalid names: Hindermeyeria Hupe 1953 [synonym]

G. †Ellipsostrenua Kautsky 1945

G. †Geyerorodes Özdikmen 2009

†Geyerorodes howleyi Walcott 1890

†Geyerorodes schmitti Geyer 1990

Invalid names: Orodes Geyer 1990 [replaced]

G. †Glabrella Lermontova 1940

†Glabrella pitans Palmer and Gatehouse 1972

†Glabrella ventrosa Lermontova 1940

G. †Hamatolenus Hupe 1952

Subg. †Hamatolenus (Hamatolenus) Hupé 1952

†Hamatolenus (Hamatolenus) vincenti Geyer and Landing 2004

G. †Hupeolenus Geyer 1990

G. †Issafeniella Geyer 1990

G. †Kadyella Pokrovskaya 1959

G. †Kameschkoviella Repina 1959

G. †Kijanella Repina 1958

G. †Kingaspidoides Hupe 1953

†Kingaspidoides alberti Geyer 2017

†Kingaspidoides frankenwaldensis Wurm 1925

†Kingaspidoides meieri Geyer 2017

†Kingaspidoides usitata Geyer 1990

Invalid names: Elatius Orlowski 1985 [synonym]

G. †Kingaspis Kobayashi 1935

Subg. †Kingaspis (Kingaspidoides) Hupe 1953

†Kingaspis (Kingaspidoides) armatus Liñán et al. 2003

Invalid names: Kingaspis (Kingaspis) brevifrons Hupé 1953 [synonym]

†Kingaspis (Kingaspidoides) velatus Lotze 1958

Subg. †Kingaspis (Kingaspis) Hupé 1954

†Kingaspis (Kingaspis) campbelli King 1923

†Kingaspis avalonensis Geyer and Landing 2001

†Kingaspis convexus Palmer and Rowell 1995

Invalid names: Mesetaia Hupe 1953 [synonym]

G. †Krolina Repina 1960

G. †Kymataspis Geyer 1990

G. †Latikingaspis Geyer 1990

†Latikingaspis alatus Hupé 1954

G. †Latoucheia Hupe 1953

G. †Latuzella Repina et al. 1999

G. †Lermontovia Suvorova 1956

G. †Limataceps Repina 1969

G. †Lusatiops Richter 1941

†Lusatiops harveyi Walcott 1890

G. †Myopsolenus Hupe 1952

Invalid names: Collyrolenus Hupe 1952 [synonym]

G. †Nelegeria Korobov 1966

G. †Olekmaspis Suvorova 1956

G. †Ourikaia Geyer 1990

G. †Paramicmacca Lermontova 1951

G. †Paraprotolenella Zhu and Jiang 1981

†Paraprotolenella subcylindrica Zhu and Jiang 1981

G. †Planolimbus Geyer 1990

G. †Protagraulos Matthew 1895

G. †Protolenella Qian and Yao 1974

†Protolenella conica Qian and Yao 1974

†Protolenella lata Zhou 1974

G. †Protolenus Matthew 1892

Subg. †Protolenus (Protolenus) Matthews 1892

†Protolenus (Protolenus) densigranulatus Geyer 1990

†Protolenus pustulatus Cobbold 1910

Invalid names: Bergeronia Matthew 1895 [objective synonym]

G. †Pruvostinoides Hupe 1952

G. †Pseudokadyella Repina 1966

G. †Pseudolenus Hupe 1953

G. †Pseudoprotolenella Zhu and Lin 1983

†Pseudoprotolenella subcylindrica Zhu and Lin 1983

†Pseudoprotolenella xinjiangensis Zhu and Lin 1983

G. †Ptychoparopsis Hupe 1953

Invalid names: Berabichia Geyer 1990 [synonym]

G. †Rinconia Hupe 1953

G. †Sailycaspis Repina 1965

G. †Sectigena Geyer 1990

G. †Strenuella Matthew 1887

†Strenuella calceola Cobbold 1931

†Strenuella cobboldi Richter 1919

†Strenuella platycephala Cobbold 1910

†Strenuella strenua Billings 1874

†Strenuella strenua nasutus Walcott 1891

G. †Tadakoustia Geyer 1990

G. †Termieraspis Repina 1960

G. †Termierella Hupe 1952

G. †Triangulaspis Lermontova 1940

Invalid names: Acutaspis Repina 1976 [synonym], Angustaeva Hupé 1953 [synonym], Triangulina Repina 1965 [synonym]

G. †Yeshanaspis Zhu and Jiang 1981

†Yeshanaspis subconicus Zhu and Jiang 1981

Invalid names: Protolenidae Richter and Richter [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

low Mg calcitec

Composition 2:

chitinc

Ontogeny:

molting, addition of partsc

Environment:

marinec

Locomotion:

fast-movingo

Life habit:

low-level epifaunalo

Diet:

deposit feedero

Vision:

well-developedf

Created:

2009-12-19 04:00:08

Modified:

2009-12-19 06:00:08

Source: f = family, o = order, c = class

References: Hendy et al. 2009, Whittington et al. 1997, Aberhan et al. 2004

Age range: base of the Branchian to the top of the Undillan or 517.10000 to 500.50000 Ma

Collections (161 total)

Time interval	Ma	Country or state	Original ID and collection number
Caerfai	538.8 - 509.0	Canada (New Brunswick)	Berabichia milleri (34579 34580) Berabichia milleri, Kingaspidoides obliquoculatus (34575 34576 34577 34578) Ellipsocephalus galeatus (34572 34597) Ellipsocephalus galeatus, Protolenus elegans (34571) Kingaspidoides obliquoculatus (34573) Protolenus articephalus, Protolenus elegans (34567 34751) Protolenus elegans (34569 34570 34595 34596 34752)
Early/Lower Cambrian	538.8 - 509.0	Antarctica	Glabrella pitans, Chorbusulina wilkesi, Chorbusulina subdita (154030)
Early/Lower Cambrian	538.8 - 509.0	Sweden (Lappland)	Ellipsocephalus gripi (190924)
Early/Lower Cambrian	538.8 - 509.0	Norway (Troms)	Ellipsocephalus sp. (190927) Strenuaeva inflata (190925 190926)
Early/Lower Cambrian	538.8 - 509.0	Spain (Aragon)	Protolenus jilocanus, Protolenus pisidianus, Alueva undulata, Hamatolenus ibericus, Kingaspis campbelli (120550 120552)
Cambrian	538.8 - 485.4	USA (Wisconsin)	Ellipsocephalus sp. (250)
Atdabanian	521.0 - 516.0	Poland	Ellipsocephalus sanctacrucensis (7905) Ellipsocephalus sanctacrucensis, Protolenus bodzanti, Protolenus medius, Strenuaeva primaeva, Strenuaeva orlowinensis, Strenuella polonica, Strenuella lakei, Termierella sandomirensis, Protolenus czarnockii (72261) Protolenus sp. (72268)
Atdabanian	521.0 - 516.0	Morocco	Antatlasia hollardi (225084) Antatlasia sp. indet. (225083) Pruvostinoides sp. (80900)
Atdabanian	521.0 - 516.0	Spain	Lusatiops sp., Protolenus schneideri, Strenueva annio, Strenueva vigilans (225087)
Atdabanian	521.0 - 516.0	Russian Federation (Siberia)	Triangulaspis lermontovae, Triangulaspis annio (96839) Triangulaspis sp. (56855) Triangulaspis sp., Protolenidae indet. (232892)
Atdabanian	521.0 - 516.0	Russian Federation (Khakasiya)	Asiatella elegans, Kijanella sp., Kijanella diffusica, Kijanella batenica (81327) Kijanella sp. (81312) Limataceps bajangolica (81314) Strenuella paraspinosa (81328)
Atdabanian	521.0 - 516.0	Antarctica	Berabichia subditus (56123)
Atdabanian	521.0 - 516.0	Sweden	Ellipsocephalus nordenskioeldi (87212)
Atdabanian	521.0 - 516.0	United Kingdom (Wales)	Hamatolenus douglasi (41750) Myopsolenus sp. (41747)
Atdabanian	521.0 - 516.0	Morocco (Ant-Atlas)	Antatlasia hollardi (232863)
Atdabanian	521.0 - 516.0	Russian Federation (Kemorovo)	Strenuella sp. (57019)
Atdabanian	521.0 - 516.0	Russian Federation (Tuva)	Bergeroniaspis vladimirskii (81465)
Nangaoian	521.0 - 514.0	China (Jiangsu)	Yeshanaspis subconicus, Paraprotolenella subcylindrica (127483)
Nangaoian	521.0 - 514.0	China (Xinjiang)	Pseudoprotolenella subcylindrica, Pseudoprotolenella xinjiangensis (127612)
Nangaoian	521.0 - 514.0	China (Guizhou)	Protolenella conica, Protolenella lata (107130)
Atdabanian - Lenian	521.0 - 510.2	Germany (Saxony)	Comluella sp., Lusatiops sp. (6899)
Branchian	517.1 - 509.0	Canada (Newfoundland and Labrador)	Solenopleura harveyi, Solenopleura howleyi, Agraulos nasutus (99495)
Branchian	517.1 - 509.0	USA (Massachusetts)	Strenuella strenua (56838 56854)
Branchian	517.1 - 509.0	United Kingdom (England)	Protolenus indet., Strenuella coboldi, Strenuella calceola, Strenuella platycephala, Protolenus pustulatus (79928)
Botomian	516.0 - 512.9	Russian Federation (Tuva)	Bergeroniaspis divergens, Bergeroniaspis dualis, Nelegeria lata (9978) Ellipsocephalus sp. (81480) Kadyella ubsanurica (81484) Strenuaeva repentina (81472)
Botomian	516.0 - 512.9	Spain (Andalucia)	Strenueva sampelayoi, Strenueva melendezi (8093)
Botomian	516.0 - 512.9	Norway (Mjosen)	Ellipsocephalus nordenskjoldi, Strenuella primaeva, Strenuella linnarssoni (42220)
Botomian	516.0 - 512.9	Russian Federation (Kemorovo)	Latuzella poletaevae (57020)
Botomian	516.0 - 512.9	Russian Federation (Yakutia)	Bergeroniaspis sp., Bergeroniellus sp. (90438) Triangulaspis sp. (174246 232979)
Botomian	516.0 - 512.9	Russian Federation	Bergeroniaspis dualis (233054) Bergeroniellus expansus (225138) Lermontovia dzevanovskii (225139) Triangulaspis lermontovae, Triangulaspis annio (233047)
Botomian	516.0 - 512.9	Poland	Ellipsocephalus sanctacrucensis, Strenuaeva orlowinensis (153007)
Botomian	516.0 - 512.9	Morocco	Berabichia vertumnia (80850 80851)
Lenian	516.0 - 510.2	Morocco	Hamatolenus sp. (161461) Kingaspidoides neglectus, Protolenus termierelloides, Protolenus dimarginatus, Protolenus hupei, Latoucheia epichara, Latoucheia ourikaensis, Ourikaia calva (9801) Kingaspis sp., Hamatolenus vincenti, Protolenus densigranulatus (161460) Latikingaspis sulcatus (9793)
Lenian	516.0 - 510.2	Sweden	Comluella scanica, Ellipsocephalus lunatus (87214) Comluella scanica, Ellipsocephalus lunatus, Strenuaeva sp. (87123) Comluella scanica, Strenuaeva sp. (87187)
Toyonian	512.9 - 510.2	Norway (Mjøsa)	Ellipsocephalus linnarssoni, Ellipsocephalus gripi (174259)
Toyonian	512.9 - 510.2	Spain (Zaragoza)	Protolenus jilocanus, Protolenus dimarginatus, Protolenus termierelloides, Kingaspis campbelli (233072)
Toyonian	512.9 - 510.2	Russian Federation (Khakasiya)	Glabrella mrassina (81331)
Toyonian	512.9 - 510.2	Antarctica	Kingaspis convexus (56127)
Toyonian	512.9 - 510.2	Morocco	Protolenus densigranulatus (9123)
Delamaran	511.2 - 506.5	Morocco	Hamatolenus maroccanus, Hamatolenus sp., Kingaspidoides laetus, Kingaspidoides neglectus, Hamatolenus vincenti (174203) Hamatolenus maroccanus, Hamatolenus sp., Kingaspidoides neglectus, Kingaspidoides laetus, Latoucheia epichara, Protolenus densigranulatus (174223) Hamatolenus maroccanus, Kingaspidoides laetus, Hamatolenus meridionalis, Hamatolenus vincenti, Protolenus densigranulatus, Ornamentaspis angustigena (174208) Hamatolenus maroccanus, Kingaspidoides laetus, Latoucheia epichara, Hamatolenus meridionalis, Hamatolenus vincenti, Protolenus densigranulatus (174210) Hamatolenus maroccanus, Latoucheia epichara, Hamatolenus lotzei, Ornamentaspis frankenwaldensis (174294) Hamatolenus sp., Latoucheia epichara (174204) Kingaspidoides laetus (174222) Kingaspidoides laetus, Hamatolenus meridionalis, Hamatolenus lotzei, Hamatolenus vincenti, Ornamentaspis frankenwaldensis (174293) Kingaspidoides laetus, Hamatolenus vincenti, Ornamentaspis angustigena (174220) Kingaspidoides laetus, Hamatolenus vincenti, Ornamentaspis frankenwaldensis (174290) Kingaspidoides laetus, Latoucheia epichara, Hamatolenus meridionalis, Hamatolenus vincenti (174292) Kingaspidoides laetus, Latoucheia epichara, Protolenus densigranulatus (174217) Kingaspidoides laetus, Protolenus densigranulatus (174291) Kingaspidoides neglectus (174205 174206 174207) Kingaspidoides neglectus, Kingaspidoides laetus, Latoucheia epichara, Hamatolenus draensis, Protolenus densigranulatus, Ornamentaspis angustigena (174219)
Amgan	510.2 - 504.5	Germany	Cambrunicornia saxonica (162995) Protolenus bergstroemi (162997) Protolenus bergstroemi, Cambrunicornia saxonica (162994 162996) Protolenus bergstroemi, Cambrunicornia saxonica, Ornamentaspis todraensis (162993)
Amgan	510.2 - 504.5	Kyrgyzstan	Glabrella ventrosa (156378)
Amgan	510.2 - 504.5	Sweden (Oland)	Ellipsocephalus polytomus (225039 225040)
Amgan	510.2 - 504.5	Czech Republic (Bohemia)	Ellipsocephalus vetustus, Acadolenus snajdri (67709) Ellipsocephalus vetustus, Ellipsocephalus hoffi, Acadolenus snajdri (67708) Ellipsocephalus vetustus, Germaropyge sp. nov. (67738)
Amgan	510.2 - 504.5	Poland	Ellipsocephalus polytomus, Ellipsocephalus sandomiri, Ellipsocephalus guerichi, Ellipsocephalus jugoszovi, Strenuella samsonowiczi, Protolenus polonicus, Kingaspis henningsmoeni (72270)
Solvan	510.2 - 504.5	Sweden (Norrbotten)	Ellipsocephalus polytomus (9782)
Solvan - Menevian	510.2 - 499.5	Czech Republic	Ellipsocephalus hoffi (73537 73539) Ellipsocephalus vetustus (73545 73561) Ellipsocephalus vetustus, Germaropyge germari (73529 73557) Germaropyge germari (203343)
Middle Cambrian	509.0 - 497.0	Czech Republic (Litavka)	Ellipsocephalus vetustus (76453) Ellipsocephalus vetustus, Acadolenus snajdri (76454)
Middle Cambrian	509.0 - 497.0	Czech Republic (Central Boemian)	Ellipsocephalus vetustus (76456)
Middle Cambrian	509.0 - 497.0	Poland	Ellipsocephalus polytomus, Ellipsocephalus guerichi, Ellipsocephalus puschi, Ellipsocephalus jugoszovi, Strenuella samsonowiczi, Strenuella hupei, Protolenus polonicus, Kingaspis henningsmoeni (72269) Ellipsocephalus polytomus, Ellipsocephalus sandomiri, Strenuella samsonowiczi (72271)
Middle Cambrian	509.0 - 497.0	India	Bhargavia prakritika (116546 116547 116548 116549)
Middle Cambrian	509.0 - 497.0	Spain	Kymataspis bernardi (55265)
Middle Cambrian	509.0 - 497.0	Italy (Iglesiente)	Protolenus pisidianus (139878)
Middle Cambrian	509.0 - 497.0	Czech Republic (Jince)	Ellipsocephalus vetustus (76483)
Middle Cambrian	509.0 - 497.0	Czech Republic (Bohemia)	Germaropyge germari (67775)
Middle Cambrian	509.0 - 497.0	Germany (Bayern)	Latikingaspis sp., Ornamentaspis frankenwaldensis (97598)
Middle Cambrian	509.0 - 497.0	Spain (Aragon)	Hamatolenus lotzei, Alueva hastata, Alueva moratrix (120553) Hamatolenus sp., Alueva hastata, Alueva moratrix (120551)
St Davids	509.0 - 497.0	USA (Massachusetts)	Kingaspis avalonensis (49340)
St Davids	509.0 - 497.0	Spain (Leon)	Acadolenus decorus (55503)
Undillan	503.0 - 500.5	Germany	Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186152) Kingaspidoides frankenwaldensis (186149 186150 186153 186154 186159 186165 186166 186170) Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186167) Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186151 186157 186158) Kingaspidoides sp. (186162 197474) Latikingaspis alatus, Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186169) Ornamentaspis crassilimbata, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186155 186156 186163) Ornamentaspis crassilimbata, Kingaspidoides frankenwaldensis (186148 186160) Ornamentaspis crassilimbata, Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186168) Ornamentaspis sp., Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186164)