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Scrotifera

Mammalia

Taxonomy
Scrotifera was named by Waddell et al. (1999). It is extant.

It was assigned to Laurasiatheria by Waddell et al. (1999) and Asher and Helgen (2010).

Sister taxa
Bulbulodentata, Euungulata (syn. Ungulata, Eparctocyona), Ferae (syn. Ostentoria), Fereuungulata

Synonyms
Synonymy list
YearName and author
1999Scrotifera Waddell et al. p. cover
2010Scrotifera Asher and Helgen p. 3 figs. Figure 1

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
RankNameAuthor
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Scrotifera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Scrotifera Waddell et al. 1999
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Or. †Acreodi Matthew 1909
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G. †Andrewsarchus Osborn 1924
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Andrewsarchus crassum Ding et al. 1977
Unr. †Mesonychia [mesonychian]
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Fm. †Hapalodectidae Szalay and Gould 1966 [condylarth]
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G. †Hapalodectes Matthew 1909
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Hapalodectes anthracinus Zhou and Gingerich 1991
Hapalodectes auctus Matthew and Granger 1925
Hapalodectes compressus Matthew 1909
Hapalodectes dux Lopatin 2001
Hapalodectes hetangensis Ting and Li 1987
Hapalodectes huanghaiensis Tong and Wang 2006
Hapalodectes leptognathus Osborn and Wortman 1892
Hapalodectes lopatini Solé et al. 2017
Hapalodectes paleocenus Beard et al. 2010
Hapalodectes paradux Lopatin 2023
Hapalodectes serus Matthew and Granger 1925
G. †Hapalorestes Gunnell and Gingerich 1996
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Hapalorestes lovei Gunnell and Gingerich 1996
Fm. †Mesonychidae Cope 1875 [condylarth]
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G. †Dissacus Cope 1881
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Dissacus argenteus O'Leary and Rose 1995
Dissacus blayaci Stehlin 1926
Dissacus bohaiensis Tong and Wang 2006
Dissacus europaeus Lemoine 1891
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Invalid names: Hyaenodictis gaudryi Lemione 1891 [synonym]
Dissacus filholi Lemoine 1891
Dissacus indigenus Dashzeveg 1976
Dissacus magushanensis Yan and Tang 1976
Dissacus navajovius Cope 1881
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Invalid names: Dissacus carnifex Cope 1882 [synonym]
Dissacus praenuntius Matthew and Granger 1915
Dissacus progressus Crusafont-Pairo and Golpe-Posse 1968
Dissacus raslanloubatieri Solé et al. 2018
Dissacus rotundus Wang 1975
Dissacus rougierae Solé et al. 2018
Dissacus saurognathus Wortman 1897
Dissacus serior O'Leary and Rose 1995
Dissacus serratus Chow and Qi 1978
Dissacus shanghoensis Chow et al. 1973
Dissacus willwoodensis O'Leary and Rose 1995
Dissacus zanabazari Geisler and McKenna 2007
Dissacus zengi Ting 2004
Invalid names: Hyaenodictis Lemoine 1891 [synonym], Hyoenodictis Lemoine 1880 [synonym], Plesidissacus Lemoine 1894 [synonym]
G. †Dissacusium Chow et al. 1977
G. †Harpagolestes Wortman 1901
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Harpagolestes brevipes Thorpe 1923
Harpagolestes immanis Matthew 1909
Harpagolestes koreanicus Shikama 1943
Harpagolestes leei Jin 2005
Harpagolestes leotensis Peterson 1931
Harpagolestes orientalis Szalay and Gould 1966
G. †Hessolestes Peterson 1931
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Hessolestes ultimus Peterson 1931
G. †Honanodon Chow 1965
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Honanodon hebetis Chow 1965
Honanodon macrodontus Chow 1965
G. †Jiangxia Zhang et al. 1979
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Jiangxia chaotoensis Zhang et al. 1979
G. †Lohoodon Chow et al. 1973
Subfm. †Mesonychinae Cope 1880
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G. †Ankalagon Van Valen 1980
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Invalid names: Ancalagon Van Valen 1978 [synonym]
G. †Guilestes Zheng and Chi 1978
G. †Hukoutherium Chow et al. 1973
G. †Mesonyx Cope 1872
G. †Mongolestes Szalay and Gould 1966
G. †Mongolonyx Szalay and Gould 1966
G. †Yantanglestes Ideker and Yan 1980
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Invalid names: Lestes Yan and Tang 1976 [replaced]
G. †Metahapalodectes Dashzeveg 1976
G. †Olsenia Matthew and Granger 1925
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Olsenia mira Matthew and Granger 1925
G. †Pachyaena Cope 1874
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Pachyaena gigantea Osborn and Wortman 1892
Pachyaena gracilis Matthew and Granger 1915
Pachyaena nemegetica Dashzeveg 1976
Pachyaena ossifraga Cope 1874
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Invalid names: Pachyaena intermedia Wortman 1899 [synonym]
G. †Plagiocristodon Chow and Qi 1978
G. †Sinonyx Zhou et al. 1995
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Sinonyx jiashanensis Zhou et al. 1995
G. †Synoplotherium Cope 1872
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Synoplotherium canius Cope 1872
Synoplotherium lanius Cope 1872
Synoplotherium vorax Marsh 1876
Invalid names: Dromocyon Marsh 1876 [synonym]
Invalid names: Hapalodectinae Szalay and Gould 1966 [empty]
G. †Simidectes Stock 1933
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Simidectes magnus Peterson 1919
Simidectes medius Peterson 1919
Simidectes merriami Stock 1933
Invalid names: Pleurocyon Peterson 1919 [synonym]
G. †Wyolestes Gingerich 1981
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Wyolestes apheles Gingerich 1981
Wyolestes dioctes Gingerich 1982
Wyolestes iglesius Novacek et al. 1991
Invalid names: Andrewsarchidae Szalay and Gould 1966 [empty], Wyolestidae [empty]
G. †Mondegodon Tabuce et al. 2011
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Mondegodon eutrigonus Tabuce et al. 2011
Fm. †Triisodontidae Scott 1892
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G. †Eoconodon Matthew and Granger 1921
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Eoconodon copanus Van Valen 1978
Eoconodon coryphaeus Cope 1885
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Invalid names: Mioclaenus bathygnathus Cope 1888 [synonym], Triisodon biculminatus Cope 1888 [synonym]
Eoconodon gaudrianus Cope 1888
Eoconodon ginibitohia Clemens and Williamson 2005
Eoconodon hutchisoni Clemens 2011
Eoconodon nidhoggi Van Valen 1978
G. †Goniacodon Scott 1892
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Goniacodon hiawathae Van Valen 1978
Goniacodon levisanus Cope 1883
G. †Stelocyon Gingerich 1978
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Stelocyon arctylos Gingerich 1978
G. †Triisodon Cope 1881
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Invalid names: Triisodon rusticus Cope 1884 [synonym]
Triisodon quivirensis Cope 1881
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Invalid names: Sarcothraustes antiquus Cope 1882 [synonym], Triisodon conidens Cope 1882 [synonym]
Invalid names: Sarcothraustes Cope 1882 [synonym]
Invalid names: Oxyclaenidae Scott 1892 [invalid subgroup]
Unr. †Bulbulodentata Archibald 1998
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Fm. †Periptychidae Cope 1882 [condylarth]
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G. †Alticonus Eberle and Lillegraven 1998
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Alticonus gazini Eberle and Lillegraven 1998
G. †Ampliconus Eberle and Lillegraven 1998
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Ampliconus antoni Eberle and Lillegraven 1998
Ampliconus browni Eberle and Lillegraven 1998
Subfm. †Anisonchinae Osborne and Earle 1895
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G. †Haploconus Cope 1882
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Haploconus angustus Cope 1881
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Invalid names: Haploconus corniculatus Cope 1888 [synonym], Haploconus inopinatus Gazin 1939 [synonym], Haploconus xiphodon Cope 1882 [synonym]
Haploconus elachistus Gazin 1941
Haploconus entoconus Cope 1882
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Invalid names: Anisonchus coniferus Cope 1882 [synonym]
Haploconus lineatus Cope 1882
G. †Pseudanisonchus Zhang et al. 1979
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Pseudanisonchus antelios Zhang et al. 1979
G. †Anisonchus Cope 1881
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Anisonchus athelas Van Valen 1978
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Invalid names: Anisonchus eowynae Van Valen 1978 [synonym]
Anisonchus fortunatus Simpson 1932
Anisonchus oligistus Gazin 1941
Anisonchus onostus Gazin 1939
Anisonchus sectorius Cope 1881
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Invalid names: Anisonchus dracus Gazin 1939 [synonym], Mioclaenus mandibularis Cope 1881 [synonym], Zetodon gracilis Cope 1883 [synonym]
Anisonchus willeyi Rigby, Jr. 1980
Invalid names: Zetodon Cope 1883 [synonym]
G. †Auraria Middleton and Dewar 2004
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Auraria urbana Middleton and Dewar 2004
G. †Beornus Atteberry and Eberle 2021
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Beornus honeyi Atteberry and Eberle 2021
G. †Conacodon Matthew 1897
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Conacodon cophater Cope 1884
Conacodon delphae Eberle and Lillegraven 1998
Conacodon harbourae Eberle and Lillegraven 1998
Conacodon hettingeri Atteberry and Eberle 2021
Conacodon kohlbergeri Archibald et al. 1983
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Invalid names: Conacodon utahensis Robison 1986 [synonym]
Conacodon matthewi Middleton and Dewar 2004
G. †Escatepos Reynolds 1936
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Escatepos campi Reynolds 1936
G. †Goleroconus McKenna and Honey 2008
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Goleroconus alfi McKenna and Honey 2008
G. †Hemithlaeus Cope 1882
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Hemithlaeus josephi Van Valen 1978
G. †Miniconus Atteberry and Eberle 2021
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Miniconus archibaldi Middleton and Dewar 2004
Miniconus jeanninae Atteberry and Eberle 2021
G. †Mithrandir Cifelli 1983
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Mithrandir gillianus Cope 1882
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Invalid names: Anisonchus apiculatus Cope 1884 [synonym]
Invalid names: Gillisonchus Rigby 1981 [synonym]
G. †Oxyacodon Osborn and Earle 1895
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Oxyacodon agapetillus Cope 1884
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Invalid names: Fimbrethil ambaronae Van Valen 1978 [synonym]
Oxyacodon apiculatus Osborn and Earle 1895
Oxyacodon ferronensis Archibald et al. 1983
Oxyacodon marshater Van Valen 1978
Oxyacodon priscilla Matthew 1937
Invalid names: Fimbrethil Van Valen 1978 [synonym]
G. †Paleoungulatum Kelly 2014
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Paleoungulatum hooleyi Kelly 2014
Subfm. †Periptychinae Osborn and Earle 1895
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G. †Ectoconus Cope 1884
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Ectoconus cavigellii Eberle and Lillegraven 1998
Ectoconus ditrigonus Cope 1882
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Invalid names: Ectoconus majusculus Matthew 1937 [synonym]
Ectoconus symbolus Gazin 1941
G. †Periptychus Cope 1881
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+
Invalid names: Catathlaeus rhabdodon Cope 1881 [synonym], Eohyus robustus Marsh 1894 [synonym], Periptychus gilmorei Gazin 1938 [synonym], Periptychus superstes Matthew 1935 [synonym]
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Invalid names: Periptychus brabensis Cope 1888 [synonym], Periptychus hamaxitus Gazin 1941 [synonym], Periptychus matthewi Simpson 1936 [synonym], Plagioptychus matthewi Simpson 1936 [synonym]
Invalid names: Carsioptychus Simpson 1936 [synonym], Catathlaeus Cope 1881 [synonym], Plagioptychus Matthew 1936 [synonym]
G. †Tinuviel Van Valen 1978
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Tinuviel eurydice Van Valen 1978
Invalid names: Conacodontinae Archibald et al. 1983 [empty]
Unr. Euungulata Waddell et al. 2001
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Unr. Artiodactylamorpha Spaulding et al. 2009
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Unr. Artiodactyla Owen 1848 [even-toed ungulate]
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Subfm. †Antiacodontinae
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G. †Sarcolemur Cope 1875
Superfm. †Cainotherioidea Camp and VanDerHoof 1940
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Fm. †Cainotheriidae Cope 1881
Fm. †Robiacinidae Weppe et al. 2019
Fm. Camelidae Gray 1821 [camel]
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G. Auchenia Illiger 1811
G. †Australocamelus Patton 1969
Subfm. Camelinae Gray 1821
Subfm. †Floridatragulinae Maglio 1966
G. †Matthewlabis Prothero 2011
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Invalid names: Paralabis Lull 1921 [replaced]
Subfm. †Miolabinae Hay 1902
Subfm. †Nothokemadinae White 1947
G. †Oxydactylus Peterson 1904
G. †Palauchenia del Castillo 1869
G. †Paratylopus Matthew 1904
G. †Poebrodon Gazin 1955
G. †Poebrotherium Leidy 1847
G. †Priscocamelus Stevens et al. 1969
Subfm. †Protolabinae Zittel 1893
Subfm. †Stenomylinae Matthew 1910
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Invalid names: Pseudolabidinae Simpson 1945 [synonym]
G. †Stilauchenia Ameghino 1889
G. †Teleopternus Cope 1899
Invalid names: Aepycamelinae Webb 1965 [invalid subgroup], Altomeryx Frick 1929 [nomen nudum], Aucheniina Bonaparte 1850 [empty], Camelina Gray 1825 [empty], Eschatiidae Cope 1887 [synonym], Gomphotherium Cope 1886 [invalid subgroup], Gomphotherium serus Douglass 1899 [nomen dubium], Merycotheriina Bonaparte 1850 [empty], Nothokematinae Prothero 1996 [empty], Poebrotheriinae Zittel 1893 [invalid subgroup], Protolabididae Cope 1887 [synonym], Protomeryx Leidy 1856 [nomen dubium]
Superfm. †Choeropotamoidea
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Fm. †Choeropotamidae Owen 1845
Fm. †Climacoceratidae Hamilton 1978
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G. †Climacoceras MacInnes 1936
G. †Prolibytherium Arambourg 1961
Invalid names: Climacoceratinae Hamilton 1978 [empty], Sperrgebietomerycinae Morales et al. 1999 [empty]
Fm. †Diacodexeidae Krishtalka and Stucky 1985
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G. †Bunophorus Sinclair 1914
G. †Diacodexis Cope 1882
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Invalid names: Trigonolestes Cope 1894 [synonym], Wasatchia Sinclair 1914 [synonym]
G. †Gujaratia Bajpai et al. 2005
G. †Magwetherium Ducrocq et al. 2016
G. †Neodiacodexis West and Atkins 1970
G. †Simpsonodus Krishtalka and Stucky 1986
G. †Tapochoerus McKenna 1959
Invalid names: Bunophorinae Stucky and Krishtalka 1990 [empty]
Superfm. †Dichobunoidea Gill 1872
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G. †Acoessus Cope 1881
Fm. †Cebochoeridae Lydekker
Fm. †Dichobunidae Gill 1872
Fm. †Helohyidae Marsh 1877
Fm. †Homacodontidae Marsh 1849
Fm. †Leptochoeridae Marsh 1894
Fm. †Mixtotheriidae Pearson 1927
Fm. †Raoellidae Sahni et al. 1981
G. †Eolantianus Averianov 1996
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Eolantianus russelli Averianov 1996
G. †Erlianhyus Li and Li 2022
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Erlianhyus primitivus Li and Li 2022
G. †Hsanotherium Ducrocq et al. 2000
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Hsanotherium parvum Ducrocq et al. 2000
Fm. †Merycoidodontidae Cook 1912 [oreodont]
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Subfm. †Aclistomycterinae Lander 1998
Subfm. †Brachycrurinae Stevens and Stevens 1996
Subfm. †Eporeodontinae Schultz and Falkenbach 1968
Subfm. †Leptaucheniinae Schultz and Falkenbach 1940
Subfm. †Merychyinae Simpson 1945
Subfm. †Merycochoerinae Schultz and Falkenbach 1940
Subfm. †Merycoidodontinae Hay 1902
Subfm. †Miniochoerinae Schultz and Falkenbach 1956
Subfm. †Oreonetinae Schultz and Falkenbach 1956
Subfm. †Promerycochoerinae Schultz and Falkenbach 1949
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Invalid names: Desmatochoerinae Schultz and Falkenbach 1954 [synonym]
Subfm. †Ticholeptinae Schultz and Falkenbach 1941
Subfm. †Ustatochoerinae Stevens and Stevens 1996
Invalid names: Merychyus pariogonus Cope 1884 [nomen dubium], Oreodontidae Leidy 1869 [synonym], Phenacocoelinae Schultz and Falkenbach 1950 [invalid subgroup]
G. †Myanmarius Tsubamoto et al. 2013
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Myanmarius chitseini Tsubamoto et al. 2013
Subor. Omnivora
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G. †Hadrohyus Leidy 1872
G. †Raphenacodus Gabunia 1971
Unr. Ruminantiamorpha Spaulding et al. 2009
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Fm. †Protoceratidae Marsh 1891 [protoceratid]
Unr. Ruminantia Scopoli 1777 [ruminant]
Subor. Suiformes Jaeckel 1911
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Invalid names: Anthracotherioidea Gill 1872 [empty], Palaeodonta Matthew [synonym]
Superfm. Suoidea Gray 1821
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G. †Doliochoerus Filhol 1882
G. †Egatochoerus Ducrocq 1994
G. †Palaeochoerus Pomel 1847
G. †Schizochoerus Crusafont-Pairo and Lavocat 1954
Fm. Suidae Gray 1821 [pig]
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Invalid names: Phacochoeridae Gray 1868 [synonym]
G. †Taucanamo Simpson 1945
Invalid names: Palaeochoeridae Matthew 1924 [invalid subgroup], Sanitheriidae Simpson 1945 [empty], Suina Gray 1868 [invalid subgroup], Suinamorpha Spaulding et al. 2009 [synonym]
Fm. †Tapirulidae Cope 1879
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G. †Obotherium Bai et al. 2023
G. †Tapiruloides Bai et al. 2023
G. †Tapirulus Gervais 1850
G. †Tragulohyus Gervais 1874
Subor. Tylopoda Illiger 1811
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Fm. †Amphimerycidae Stehlin 1910
Superfm. †Anoplotherioidea Bonaparte 1850
Fm. †Oromerycidae Gazin 1955
Invalid names: Camelidamorpha Spaulding et al. 2009 [synonym], Cameloidea [empty], Oreodontoidea Gill 1872 [empty], Xiphodontoidea Flower 1883 [empty]
Invalid names: Achaenodontidae [empty], Ancodonta Matthew [empty], Antiacodontidae Krishtalka and Stucky 1985 [empty], Booidea [empty], Bunomerycidae Gentry and Hooker 1988 [empty], Caenotheriidae [empty], Cetartiodactyla Montgelard et al. 1997 [synonym], Eurytheriidae Cope 1889 [empty], Haplobunodontidae Sudre 1978 [empty], Hyodonta Matthew 1929 [empty], Leptochoerinae [empty], Merycoidodontoidea [empty], Neoselenodontia Webb and Taylor 1980 [empty], Nonruminantia Gregory 1910 [empty], Trigonolestidae [empty]
Unr. †Panperissodactyla Welker et al. 2015
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Or. †Panameriungulata de Muizon and Cifelli 2000
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Fm. †Didolodontidae Simpson 1934
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G. †Archaeohyracotherium Ameghino 1906
G. †Argyrolambda Ameghino 1904
G. †Depaulacoutoia Cifelli and Ortiz-Jaureguizar 2014
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Invalid names: Paulacoutoia Cifelli 1983 [replaced]
G. †Didolodus Ameghino 1897
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Invalid names: Cephanodus Ameghino 1902 [synonym], Lonchoconus Ameghino 1901 [synonym], Nephacodus Ameghino 1902 [synonym]
G. †Enneoconus Ameghino 1901
G. †Ernestokokenia Ameghino 1901
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Invalid names: Notoprotogonia Ameghino 1904 [synonym]
G. †Escribania Bonaparte et al. 1993
G. †Lamegoia Paula Couto 1952
G. †Paulogervaisia Ameghino 1901
G. †Proectocion Ameghino 1904
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Invalid names: Oxybunotherium Pascual 1965 [synonym]
G. †Raulvaccia Bonaparte et al. 1993
G. †Ricardocifellia Mones 2015
G. †Salladolodus Soria and Hoffstetter 1983
G. †Saltaodus Gelfo et al. 2020
G. †Umayodus Gelfo and Sigé 2011
Invalid names: Bunolitopternidae Schlosser 1823 [synonym], Didolodidae Scott 1913 [replaced]
Subor. †Litopterna Ameghino 1889
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Fm. †Adianthidae Ameghino 1891
G. †Eolicaphrium Ameghino 1902
Subor. †Lopholipterna Cifelli 1983
Fm. †Macraucheniidae Ameghino 1889
Fm. †Mesorhinidae Ameghino 1891
Fm. †Notonychopidae Soria 1989
Fm. †Proterotheriidae Ameghino 1887
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Invalid names: Bunodontheridae Mercerat 1891 [synonym]
Fm. †Protolipternidae Cifelli 1983
Fm. †Sparnotheriodontidae Soria 1980
G. †Thoatheriopsis Soria 2001
Invalid names: Eolitopterna Soria 2001 [empty], Lambdaconus elegans Roth 1904 [nomen dubium], Paulogervaisia celata Ameghino 1901 [nomen dubium], Proacrodon Roth 1899 [nomen vanum], Proacrodon transformatus Roth 1899 [nomen vanum], Proadiantus gibbus Ameghino 1901 [nomen vanum]
Superor. Perissodactylamorpha Rose et al. 2020
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Or. Perissodactyla Owen 1848 [odd-toed ungulate]
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Fm. †Anthracobunidae Wells and Gingerich 1983
Fm. †Brontotheriidae Marsh 1873 [brontothere]
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Invalid names: Titanotheriidae [synonym]
Fm. †Cambaytheriidae Bajpai et al. 2005
G. †Cambaytherium Bajpai et al. 2005
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Invalid names: Indobune Rose et al. 2006 [synonym]
Or. †Desmostylia Reinhart 1953 [desmostylian]
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Invalid names: Desmostyliformes Hay 1923 [synonym]
Superfm. Equoidea Hay 1902
G. †Ghazijhippus Missiaen and Gingerich 2014
G. †Hallensia Franzen and Haubold 1986
Subor. †Lophodontomorpha Hooker and Dashzeveg 2004
G. †Mesolambdolophus Holbrook and Lapergola 2011
G. †Nakusia Ginsburg et al. 1999
G. †Paschatherium Russell 1963
G. †Perissobune Missiaen and Gingerich 2014
G. †Skopaiolophus Métais et al. 2006
Subor. †Solidungula Cuvier
Unr. Tapiromorpha Haeckel 1873
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Invalid names: Moropomorpha [synonym]
G. †Teilhardimys Kretzoi and Kretzoi 2000
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Invalid names: Microhyus Teilhard de Chardin 1927 [replaced]
Invalid names: Chelodactyla Scott 1913 [empty], Equina Bonaparte 1850 [empty], Hippopotamus iravaticus Noetling 1897 [nomen dubium], Hyracotheriidae Cope 1881 [empty], Mesaxonia Marsh 1884 [empty], Rhinocerotida Owen 1845 [empty], Selenida McKenna and Bell 1997 [invalid subgroup], Stereopterna Ameghino 1889 [empty], Triplopodidae Cope 1881 [empty]
G. †Radinskya McKenna et al. 1989
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Radinskya yupingae McKenna et al. 1989
Invalid names: Anthracobunia Ginsburg et al. 1999 [empty]
Unr. Phenacodonta McKenna 1975
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G. †Olbitherium Tong et al. 2004
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Olbitherium millenariusum Tong et al. 2004
Fm. †Phenacodontidae Cope 1881 [condylarth]
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G. †Almogaver Crusafont-Pairo and Villalta 1954
G. †Copecion Gingerich 1989
G. †Eodesmatodon Zheng and Chi 1978
G. †Lophocion Wang and Tong 1997
Subfm. †Meniscotheriinae Cope 1882
G. †Phenacodus Cope 1873
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Invalid names: Eohyus Marsh 1894 [synonym], Opisthotomus Cope 1875 [synonym], Trispondylus Cope 1884 [replaced]
G. †Tetraclaenodon Scott 1892
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Invalid names: Euprotogonia Cope 1893 [synonym], Protogonia Cope 1881 [synonym]
Invalid names: Eparctocyona McKenna 1975 [synonym], Ungulata Linnaeus 1766 [synonym]
Unr. Ferae Linnaeus 1758
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Unr. Carnivoramorpha Wyss and Flynn 1993
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Unr. †Carnivoraformes Flynn et al.
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G. †Africtis Mattingly et al. 2020
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Africtis sirtensis Mattingly et al. 2020
Or. Carnivora Bowdich 1821 [carnivoran]
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G. †Adracon Filhol 1884
Fm. Canidae Fischer de Waldheim 1817 [canid]
Subor. Caniformia Kretzoi 1943
G. †Cryptailurus Martin and Lim 2001
Subor. Feliformia Kretzoi 1945
Subor. Fissipedia Blumenbach 1791
Superfm. †Miacoidea Cope
G. †Monachiopsis Kretzoi 1941
Fm. †Percrocutidae Werdelin and Solounias 1991
G. †Pristinictis Fox and Youzwyshyn 1994
G. †Ravenictis Fox and Youzwyshyn 1994
G. †Thos Bogachev
Invalid names: Eosictis Scott 1945 [nomen dubium], Felidi Jaekel 1911 [empty], Phocidi Jaekel 1911 [empty], Protelidi Jaekel 1911 [empty], Protoferi Jaekel 1911 [empty], Ursina Gray 1825 [empty], Ursocanidi Jaekel 1911 [empty], Viverra memphitis Linnaeus 1758 [nomen vanum]
G. †Dormaalocyon Solé et al. 2014
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Dormaalocyon latouri Quinet 1966
G. †Gracilocyon Smith and Smith 2010
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Gracilocyon igniculus Beard and Dawson 2009
Gracilocyon rosei Heinrich et al. 2008
Gracilocyon rundlei Hooker 2010
Gracilocyon solei Smith and Smith 2010
Gracilocyon winkleri Gingerich 1983
G. Lycarion Matthew 1909
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Lycarion medius Matthew 1909
G. †Miacis Cope 1872
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Miacis bathygnathus Scott 1888
Miacis boqinghensis Huang et al. 1999
Miacis deutschi Gingerich 1983
Miacis edax Leidy 1872
Miacis exiguus Matthew and Granger 1915
Miacis gracilis Clark 1939
Miacis hookwayi Stock 1934
Miacis invictus Matthew and Granger 1925
Miacis latidens Matthew and Granger 1915
Miacis lushiensis Chow 1975
Miacis parvivorus Cope 1872
Miacis petilus Gingerich 1983
Miacis rundlei Hooker 2010
Miacis vorax Leidy 1872
Miacis vulpinus Scott and Osborn 1887
Invalid names: Harpalodon Marsh 1872 [synonym], Prodaphaenus Wortman and Matthew 1899 [synonym]
G. †Neovulpavus Wortman 1901
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Neovulpavus mccarrolli Tomiya et al. 2021
Neovulpavus washakius Wortman 1901
G. †Palaearctonyx Matthew 1909
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Palaearctonyx meadi Matthew 1909
G. †Paramiacis Filhol 1876
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Paramiacis exilis Filhol 1876
Paramiacis teilhardi Mathis 1987
G. †Procynodictis Wortman and Matthew 1899
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Procynodictis vulpiceps Wortman and Matthew 1899
Invalid names: Plesiomiacis Simpson 1945 [synonym]
G. †Quercygale Kretzoi 1945
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Quercygale angustidens Filhol 1872
Quercygale hastingsae Davies 1884
Quercygale helvetica Rütimeyer 1862
Quercygale smithi Solé 2014
Invalid names: Humbertia De Beaumont 1965 [objective synonym]
G. †Uintacyon Leidy 1873
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Uintacyon acutus Thorpe 1923
Uintacyon asodes Gazin 1952
Uintacyon gingerichi Heinrich et al. 2008
Uintacyon hookeri Solé 2014
Uintacyon jugulans Matthew 1909
Uintacyon rudis Rose 1981
G. †Vassacyon Matthew 1909
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Vassacyon bowni Heinrich et al. 2008
Vassacyon prieuri Solé et al. 2016
Vassacyon promicrodon Wortman and Matthew 1899
Vassacyon taxidiotis Solé et al. 2013
G. †Vulpavus Marsh 1871
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Subg. †Vulpavus (Phlaodectes) Matthew 1909
Vulpavus australis Matthew and Granger 1915
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Invalid names: Vulpavus asius Gazin 1952 [synonym]
Vulpavus canavus Cope 1881
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Invalid names: Miacis brevirostris Cope 1881 [synonym]
Vulpavus completus Matthew 1909
Vulpavus farsonensis Gunnell 1998
Vulpavus hargeri Wortman 1901
Vulpavus ovatus Matthew 1909
Vulpavus palustris Marsh 1871
Vulpavus profectus Matthew 1909
G. †Xinyuictis Zheng et al. 1975
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Xinyuictis tenuis Zheng et al. 1975
G. †Zodiocyon Tong and Wang 2006
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Zodiocyon zetesios Tong and Wang 2006
G. †Dawsonicyon Spaulding et al. 2010
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Dawsonicyon isami Spaulding et al. 2010
G. †Oodectes Wortman 1901
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Oodectes herpestoides Wortman 1901
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Invalid names: Oodectes proximus Matthew 1909 [synonym], Paeneprolimnocyon amissadomus Guthrie 1967 [synonym], Triacodon nanus Marsh 1872 [synonym]
Oodectes jepseni Guthrie 1967
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Invalid names: Paeneprolimnocyon iudei Guthrie 1967 [synonym]
Oodectes pugnax Wortman and Matthew 1899
Invalid names: Paeneprolimnocyon Guthrie 1967 [synonym]
G. †Tapocyon Stock 1934
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Tapocyon dawsonae Wesley and Flynn 2003
Tapocyon robustus Peterson 1919
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Invalid names: Tapocyon occidentalis Stock 1934 [synonym]
Fm. †Viverravidae Wortman and Matthew 1899
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G. †Didymictis Cope 1875
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Didymictis altidens Cope 1880
Didymictis dellensis Dorr 1952
Didymictis leptomylus Cope 1880
Didymictis protenus Cope 1874
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Invalid names: Didymictis curtidens Cope 1882 [synonym]
Didymictis proteus Polly 1997
Didymictis vancleveae Robinson 1966
G. †Orientictis Huang and Zheng 2005
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Orientictis spanios Huang and Zheng 2005
G. †Preonictis Tong and Wang 2006
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Preonictis youngi Tong and Wang 2006
G. †Protictis Matthew
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Subg. †Protictis (Bryanictis) MacIntyre 1966
Subg. †Protictis (Protictis) Matthew 1937
Subg. †Protictis (Protictoides) Flynn and Galiano 1982
Protictis agastor Gingerich and Winkler 1985
Protictis haydenianus Cope 1882
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Invalid names: Didymictis primus Cope 1884 [synonym], Prolimnocyon macfaddeni Rigby, Jr. 1980 [synonym]
Protictis minor Meehan and Wilson 2002
Protictis paralus Holtzman 1978
Protictis simpsoni Meehan and Wilson 2002
G. †Simpsonictis MacIntyre 1962
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Simpsonictis jaynanneae Rigby, Jr. 1980
Simpsonictis pegus Gingerich and Winkler 1985
Simpsonictis tenuis Simpson 1935
G. †Variviverra Tong and Wang 2006
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Variviverra vegetatus Tong and Wang 2006
G. †Viverravus Marsh 1872
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Viverravus acutus Matthew and Granger 1915
Viverravus gracilis Marsh 1872
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Invalid names: Didymictis dawkinsianus Cope 1881 [synonym], Harpalodon vulpinus Marsh 1872 [synonym], Triacodon fallax Marsh 1871 [synonym], Ziphacodon rugatus Marsh 1872 [synonym]
Viverravus lawsoni Hooker 2010
Viverravus laytoni Gingerich and Winkler 1985
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Invalid names: Viverravus bowni Gingerich 1987 [synonym]
Viverravus lutosus Gazin 1952
Viverravus minutus Wortman 1901
Viverravus nitidus Marsh 1872
Viverravus politus Matthew and Granger 1915
Viverravus rosei Polly 1997
Viverravus schaffi Gingerich and Winkler 1985
Viverravus sicarius Matthew 1909
Invalid names: Ziphacodon Marsh 1872 [synonym]
G. †Viverriscus Beard and Dawson 2009
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Viverriscus omnivorus Beard and Dawson 2009
Invalid names: Deltatherium durini Van Valen 1978 [nomen dubium], Didymictidae Flynn and Galiano 1982 [synonym]
Or. †Creodonta Cope 1875 [creodont]
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G. †Altacreodus Fox 2015
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Altacreodus magnus Clemens and Russell 1965
Or. †Hyaenodonta Van Valen 1967
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G. †Eoproviverra Solé et al. 2014
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Eoproviverra eisenmanni Godinot 1981
Superfm. †Hyaenodontoidea Leidy 1869
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Fm. †Hyaenodontidae Leidy 1869
Subfm. †Proviverrinae Schlosser 1886
Superfm. †Hyainailouroidea Borths et al. 2016
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Unr. †Hyainailouridae Solé et al. 2015
G. †Kyawdawia Egi et al. 2005
G. †Metasinopa Osborn 1909
G. †Paratritemnodon Ranga Rao 1973
Unr. †Teratodontinae Savage 1965
Subfm. †Limnocyoninae Wortman 1902
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G. †Iridodon Morlo and Gunnell 2003
G. †Limnocyon Marsh 1872
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Invalid names: Telmatocyon Marsh 1899 [synonym]
G. †Oxyaenodon Matthew 1899
G. †Prolaena Xu et al. 1979
G. †Prolimnocyon Matthew and Granger 1915
G. †Thinocyon Marsh 1872
Invalid names: Limnocyonini Wortman 1902 [empty], Machaeroidini Matthew 1909 [empty]
Unr. †Oxyaenodonta Van Valen 1971
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Fm. †Oxyaenidae Cope 1877
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G. †Dipsalidictis Matthew and Granger 1915
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Invalid names: Dipsalidictides Denison 1938 [synonym]
G. †Dipsalodon Jepsen 1930
G. †Dormaalodon Lange-Badré 1987
G. †Galethylax Gervais 1850
G. †Machaeroides Matthew 1909
Subfm. †Machaeroidinae
Subfm. †Oxyaeninae Trouessart 1885
Subfm. †Palaeonictinae Denison 1938
G. †Sarkastodon Granger 1938
G. †Tytthaena Gingerich 1980
Invalid names: Ambloctonidae Cope 1877 [empty], Amblyctonidae Cope 1877 [empty], Arctocyonoidae Trouessart 1885 [empty], Hyaenodontida Leidy 1869 [empty], Limnocyonidae [empty], Oxyaenoidae Cope 1877 [empty], Uintacyonidae Hay 1902 [empty]
Unr. †Pholidotamorpha Gaudin et al. 2009
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Subor. †Palaeanodonta Matthew 1918
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G. †Amelotabes Rose 1978
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Amelotabes simpsoni Rose 1978
G. †Arcticanodon Rose et al. 2004
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Arcticanodon dawsonae Rose et al. 2004
Fm. †Epoicotheriidae Simpson 1927
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G. †Alocodontulum Rose et al. 1978
G. †Auroratherium Tong and Wang 1997
G. †Dipassalus Rose et al. 1991
G. †Epoicotherium Simpson 1927
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Invalid names: Pseudochrysochloris Turnbull and Reed 1967 [synonym], Xenotherium Douglass 1905 [replaced]
G. †Tetrapassalus Simpson 1959
G. †Tubulodon Jepsen 1932
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Invalid names: Alocodon Rose et al. 1977 [synonym], Pentapassalus Gazin 1952 [synonym]
G. †Xenocranium Colbert 1942
G. †Ernanodon Ding 1979
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Ernanodon antelios Ding 1979
Fm. †Escavadodontidae Rose and Lucas 2000
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G. †Escavadodon Rose and Lucas 2000
G. †Melaniella Fox 1984
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Melaniella timosa Fox 1984
Fm. †Metacheiromyidae Wortman 1903
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G. †Brachianodon Gunnell and Gingerich 1993
G. †Metacheiromys Wortman 1903
G. †Mylanodon Secord et al. 2002
G. †Palaeanodon Matthew 1918
G. †Propalaeanodon Rose 1979
Or. Pholidota Weber 1904 [pangolin]
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Unr. †Eupholidota Gaudin et al. 2009
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G. †Eomanis Storch 1978
Unr. †Manoidea Gaudin et al.
G. †Eurotamandua Storch 1981
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Eurotamandua joresi Storch 1981
G. †Leptomanis Filhol 1894
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Leptomanis edwardsi Filhol 1894
Invalid names: Eomanidae Storch 2003 [empty], Squamata Huxley 1872 [synonym]
Invalid names: Ostentoria [synonym]
Unr. Fereuungulata Waddell et al. 1999
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Invalid names: Zooamata Waddell et al. 1999 [empty]
Invalid names: Variamana [synonym]
Diagnosis
No diagnoses are available