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Scaldiphora

Taxonomy
Scaldiphora is an unranked clade. It is extant.

It was assigned to Cycloneuralia by Nielsen (2012).

Subtaxa
Priapula, Priapulida
Sister taxa
Priapula

Synonymy list
YearName and author
2012Scaldiphora Nielsen p. 290 figs. Fig. 47.3

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
RankNameAuthor
—Ecdysozoa
—CycloneuraliaAhlrichs 1995
—Scaldiphora
—Scaldiphora

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Ph. Priapulida Théel 1906 [priapulid worm]
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G. †Acosmia Chen and Zhou 1997
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†Acosmia maotiania Chen and Zhou 1997
Unr. †Archaeopriapulida Conway Morris 1977
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Fm. †Fieldiidae Conway Morris 1977
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G. †Fieldia Walcott 1912
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†Fieldia lanceolata Walcott 1912
G. †Lecythioscopa Conway Morris 1977
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†Lecythioscopa simplex Conway Morris 1977
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Invalid names: Canadia simplex Walcott 1931 [replaced]
Fm. †Miskoidae Walcott 1911
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G. †Louisella
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†Louisella pedunculata Conway Morris 1977
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Invalid names: Miskoia pedunculata Walcott 1911 [replaced]
G. †Miskoia Walcott 1911
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†Miskoia preciosa Walcott 1911
Fm. †Ottoidae Walcott 1911
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G. †Banffia Walcott 1911
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†Banffia constricta Walcott 1911
†Banffia episoma Conway Morris et al. 2015
G. †Ottoia Walcott 1911
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†Ottoia minor Walcott 1911
†Ottoia prolifica Walcott 1911
†Ottoia tenuis Walcott 1911
G. †Scolecofurca Conway Morris 1977
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†Scolecofurca rara Conway Morris 1977
Fm. †Selkirkiidae Conway Morris 1977
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G. †Selkirkia Walcott 1911
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†Selkirkia columbia Conway Morris 1977
†Selkirkia fragilis Walcott 1911
†Selkirkia major Walcott 1908
G. †Singuuriqia Peel 2017
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†Singuuriqia simoni Peel 2017
Invalid names: Ancalagonidae Conway Morris 1977 [empty]
G. †Archotuba Hou et al. 1999
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†Archotuba conoidalis Hou et al. 1999
G. †Corynetis Luo and Hu 1999
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†Corynetis brevis Luo and Hu 1999
†Corynetis fortis Hu et al. 2012
Invalid names: Anningvermis Huang et al. 2004 [synonym]
G. †Eximipriapulus Ma et al. 2014
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†Eximipriapulus globocaudatus Ma et al. 2014
G. †Omnidens Hou and Others 2006
Unr. †Palaeoscolecidomorpha Shi et al. 2022
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†Cricocosmia jinningensis Hou and Sun 1988
G. †Houscolex
G. †Tabelliscolex Han et al. 2003
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†Tabelliscolex chengjiangensis Han et al. 2007
†Tabelliscolex hexagonus Han et al. 2003
G. †Tylotites Luo and Hu 1999
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†Tylotites petiolaris Luo and Hu 1999
G. †Maotianshania Sun and Huo 1987
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†Maotianshania cylindrica Sun and Huo 1987
G. †Markuelia Valkov 1983
Cl. †Palaeoscolecida Conway-Morris and Robison 1986
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G. †Corallioscolex Müller and Hintz-Schallreuter 1993
G. †Cowiella Hinz 1987
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†Cowiella granulata Hinz 1987
†Cowiella reticulata Hinz 1987
†Cowiella undulata Hinz 1987
†Cowiella verrucata Hinz 1987
Or. †Cricocosmida Han et al. 2007
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Fm. †Palaeoscolecidae Whittard 1953
Invalid names: Tylotitidae Han and Others 2003 [empty]
G. †Milaculum
Invalid names: Palaeoscolecida [invalid subgroup]
G. †Paraselkirkia Luo and Hu 1999
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†Paraselkirkia jinningensis Hou et al. 1999
Fm. Priapulidae Gosse 1855
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G. Acanthopriapulus van der Land 1970
G. †Priapulites Schram 1973
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†Priapulites konecniorum Schram 1973
G. Priapulopsis Koren and Danielssen 1875
G. Priapulus Lamarck 1816
G. †Xiaoheiqingella Hu 2002
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Invalid names: Yunnanpriapulus halteroformis Huang et al. 2004 [synonym]
Invalid names: Yunnanpriapulus Huang et al. 2004 [synonym]
Or. †Priapulomorpha Slavini-Plawen 1974
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G. †Ancalagon Conway Morris 1977
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†Ancalagon minor Walcott 1911
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†Sicyophorus rara Luo and Hu 1999
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Invalid names: Protopriapulites haikouensis Hou et al. 1999 [synonym]
Invalid names: Palaeopriapulites Hou et al. 1999 [synonym], Protopriapulites Hou et al. 1999 [synonym]
Fm. Tubiluchidae Van der Land 1970
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G. †Paratubiluchus Han et al. 2004
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†Paratubiluchus bicaudatus Han et al. 2004
Invalid names: Halicryptidae Salvini-Plawen 1974 [empty], Maccabeidae Por and Bromley 1974 [empty], Xiaoheiqingidae Chen 2002 [empty]
Invalid names: Priapula [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Meishucunian to the top of the Westphalian D or 530.70000 to 307.50000 Ma
Minimum age of oldest fossil (stem group age): 525.5 Ma

Collections (167 total)


Time interval Ma Country or state Original ID and collection number
Nemakit-Daldynian538.8 - 525.5Russian Federation (Siberia) Markuelia sp. (50340)
Early/Lower Cambrian538.8 - 509.0China (Yunnan) Paraselkirkia sp., Sicyophorus sp. (185938 185939) Xiaoheiqingella sp. (228573)
Waucoban538.8 - 509.0Canada (Northwest Territories) Palaeoscolecida indet. (227949)
Caerfai - St Davids538.8 - 497.0China (Yunnan) Maotianshania cylindrica (9940 9941) Maotianshania cylindrica, Palaeoscolex sinensis (9937) Palaeoscolex sinensis, Cricocosmia jinningensis (9939)
Meishucunian530.7 - 521.0China (Shaanxi) Houscolex lepidotus, Houscolex sp. (6851)
Tommotian525.5 - 521.0Russian Federation (Khabarovsk Krai) Markuelia secunda (50314)
Tommotian525.5 - 521.0China (Yunnan) Maotianshania cylindrica (6817 6818 6819) Palaeoscolex sinensis (9784 9785 9786)
Tommotian - Caerfai525.5 - 509.0China (Yunnan) Maotianshania cylindrica (9783)
Atdabanian521.0 - 516.0China (Yunnan) Cricocosmia jiningensis, Banffia confusa, Maotianshania cylindrica, Paraselkirkia sinica, Palaeoscolex sinensis, Tylotites petiolaris, Tabelliscolex hexagonus, Sicyophorus rara, Acosmia maotiania, Corynetis brevis (68083) Cricocosmia jiningensis, Maotianshania cylindrica, Palaeoscolex sinensis (86851) Cricocosmia jiningensis, Palaeoscolex sinensis, Tabelliscolex hexagonus (86845) Cricocosmia jinningensis, Sicyophorus rara (90136) Mafangscolex sinensis (90138) Mafangscolex sinensis, Cricocosmia jinningensis (90149 90150 90164 90165) Maotianshania cylindrica (90494) Maotianshania cylindrica, Mafangscolex sinensis (90161 90163 90166) Maotianshania cylindrica, Palaeoscolex sinensis (9938) Maotianshania cylindrica, Palaeoscolex sinensis, Sicyophorus rara, Palaeopriapulites parvus (86846) Maotianshania sp., Acosmia sp. (86933) Paraselkirkia sinica, Mafangscolex sinensis (90152) Paraselkirkia sinica, Mafangscolex sinensis, Cricocosmia jinningensis, Sicyophorus rara, Corynetis brevis (90133 90135 90137) Paraselkirkia sinica, Mafangscolex sinensis, Cricocosmia jinningensis, Sicyophorus rara, Corynetis brevis, Xiaoheiqingella peculiaris (90151) Priapulida indet., Cricocosmia sp., Palaeoscolex sp., Banffia sp., Maotianshania sp., Acosmia sp., Corynetis sp., Paraselkirkia sp., Sicyophorus sp., Anningvermis sp. (86929) Priapulida indet., Palaeoscolex sp., Maotianshania sp., Corynetis sp., Paraselkirkia sp., Protopriapulites sp. (86930)
Atdabanian521.0 - 516.0Mongolia Markuelia sp. (219046)
Atdabanian521.0 - 516.0United Kingdom (England) Cowiella granulata (79940) Cowiella reticulata, Cowiella verrucata, Cowiella undulata (79933)
Atdabanian521.0 - 516.0China (Hubei) Maotianshania cylindrica (68086)
Nangaoian521.0 - 514.0China (Hubai) Selkirkia sp., Banffia sp., Archotuba sp., Corynetis sp. (219687)
Nangaoian521.0 - 514.0China (Yunnan) Mafangscolex sinensi, Cricocosmia jinningensis, Corynetis brevis, Archotuba conoidalis, Paraselkirkia jinningensis (194160) Mafangscolex sinensis (132328) Mafangscolex sinensis, Cricocosmia jinningensis (194140) Mafangscolex sinensis, Cricocosmia jinningensis, Archotuba conoidalis (132329 194162) Mafangscolex sinensis, Cricocosmia jinningensis, Archotuba conoidalis, Paraselkirkia jinningensis (194150 194155) Mafangscolex sinensis, Cricocosmia jinningensis, Corynetis brevis, Palaeopriapulites parvus, Xiaoheiqingella peculiaris, Archotuba conoidalis, Paraselkirkia jinningensis (132330) Mafangscolex sinensis, Cricocosmia jinningensis, Palaeopriapulites parvus, Archotuba conoidalis, Paraselkirkia jinningensis (194143 194161 194163) Mafangscolex sinensis, Cricocosmia jinningensis, Xiaoheiqingella peculiaris, Archotuba conoidalis, Paraselkirkia jinningensis (194141) Mafangscolex sinensis, Cricocosmia jinningensis, Yunnanpriapulus halteroformis, Paraselkirkia jinningensis (194153) Mafangscolex sinensis, Tabelliscolex hexagonus, Cricocosmia jinningensis, Archotuba conoidalis (132327)
Atdabanian - Caerfai521.0 - 509.0Greenland Singuuriqia simoni (9780)
Branchian517.1 - 509.0United Kingdom (England) Cowiella reticulata (79935 79937) Hadimopanella apicata (79931) Milaculum sp. indet., Cowiella reticulata, Cowiella verrucata (79928)
Botomian516.0 - 512.9Russian Federation (Yakutia) Palaeoscolecida indet., Palaeoscolex sp., Corallioscolex sp., Palaeoscolex lubovae, Palaeoscolex spinosus, Sahascolex labyrinthus (90438)
Botomian516.0 - 512.9China (Yunnan) Guanduscolex minor (90357) Palaeoscolex sp. (90363 90367 90368 90369 90370 90371) Paramaotianshania zijunia, Corynetis fortis (139521)
Botomian516.0 - 512.9Russian Federation (Siberia) Hadimopanella knappologica (174262)
Botomian516.0 - 512.9Australia (South Australia) Palaeoscolex antiquus, Wronascolex iacoborum (187498)
Lenian516.0 - 510.2Antarctica (South Shetland Islands) Hadimopanella antarctica (49692)
Dyeran515.3 - 511.2USA (Nevada) Ottoia sp. (10560) Selkirkia sp. (9820)
Dyeran515.3 - 511.2Greenland Hadimopanella apicata (49466 49467 49469 49470 49473 67782 67783 67787 67795 67797 67798 67799 67800)
Dyeran515.3 - 511.2USA (Pennsylvania) Ottoia sp., Selkirkia pennsylvanica (192 226194)
Duyunian514.0 - 509.0China (Shaanxi) Palaeoscolecida indet. (90680)
Toyonian512.9 - 510.2Turkey (Konya) Hadimopanella oezgueli (93621)
Toyonian512.9 - 510.2Australia (South Australia) Kaimenella reticulata (64472 97986 97987 118171 118173)
Delamaran511.2 - 506.5USA (Idaho) Selkirkia sp. (207) Selkirkia spencei (76096)
Delamaran511.2 - 506.5Canada (British Columbia) Ottoia prolifica, Selkirkia columbia (84414 84416 84417 84434 84437) Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84423 84436) Priapulida indet. A, Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84425) Priapulida indet. B, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84418) Selkirkia columbia (84413 84415 84420) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia (84421) Selkirkia sp. A, Ancalagon minor, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84424) Selkirkia sp. A, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia (84427) Selkirkia sp. A, Fieldia lanceolata, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84422) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia (84412 84419 84426 84430 84431 84433) Selkirkia sp. A, Ottoia prolifica, Selkirkia columbia, Louisella pedunculata (84428 84429 84432 84435)
Delamaran511.2 - 506.5USA (Utah) Selkirkia spencei, Selkirkia columbia, Palaeoscolex ratcliffei (77218) Selkirkia willoughby, Selkirkia sp. (77219)
Amgan510.2 - 504.5Kyrgyzstan Hadimopanella oezgueli (156378)
Amgan510.2 - 504.5Russian Federation (Siberia) Hadimopanella knappologica (110619)
Solvan510.2 - 504.5Russian Federation (Siberia) Hadimopanella knappologica (49429)
Solvan510.2 - 504.5USA (Nevada) Ottoia sp. (9215)
Taijangian509.0 - 504.5China (Guizhou) Ottoia sp. (79770)
Middle Cambrian509.0 - 497.0Canada (Brithish Columbia) Selkirkia sp. (188664)
Middle Cambrian509.0 - 497.0Canada (British Columbia) Priapulida indet., Selkirkia sp., Banffia sp. (171308)
Middle Cambrian509.0 - 497.0USA (Utah) Banffia episoma (80122)
St Davids509.0 - 497.0Canada (British Columbia) Ottoia sp. (122017 122072) Ottoia sp., Ottoia prolifica (122094)
Menevian504.5 - 499.5USA (Utah) Ottoia sp., Selkirkia sp., Palaeoscolecida gen. et sp. nov. A (230681)
Marjumian504.5 - 497.0USA (Utah) Ottoia prolifica (77206) Selkirkia willoughbyi, Scathascolex minor, Ottoia prolifica (230679)
Guzhangian500.5 - 497.0China (Yunnan) Palaeoscolecida indet. (219647)
Merioneth497.0 - 485.4Russian Federation (Siberia) Protoscolex tchopkoensis (10009)
Floian477.7 - 470.0China (Hunan) Ottoia sp., Palaeoscolecida indet. (226639)
Darriwilian467.3 - 458.4United Kingdom (Wales) Priapulida NMW.2021.3 G.97, Palaeoscolecida NMW.2021.3 G.89, Archaeopriapulida NMW.2021.3 G.19 (230215)
Llandeilo461.1 - 458.4United Kingdom (Wales) Palaeoscolecidae gen. et sp. nov. Botting et al. in press (216511)
Caradoc458.4 - 449.5USA (New York) Protoscolex sp. (340)
Turinian457.3 - 453.8Canada (British Columbia) Milaculum sp. (192195)
Rocklandian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90362)
Rocklandian - Shermanian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90360)
Shermanian457.3 - 449.6USA (Wisconsin) Milaculum sp. (90366)
Edenian451.0 - 450.2USA (Ohio) Protoscolex sp. (383 384)
Maysvillian450.2 - 449.6Canada (Ontario) Protoscolex sp. (26302)
Richmondian449.6 - 445.5Canada (Nunavut) Milaculum ethinclarki (360)
Rhuddanian - Aeronian443.4 - 438.5USA (Iowa) Protoscolex sp. (26860)
Telychian438.5 - 433.4USA (Wisconsin) Palaeoscolecida indet. (91415)
Ludlow427.4 - 423.0USA (Illinois) Protoscolex ruedemanni (76450) Protoscolex sp. (26861)
Westphalian D309.8 - 307.5USA (Illinois) Priapulites konecniorum (122493 122538 151852 168004)