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Carettochelyidae
Taxonomy
Carettochelyidae was named by Gill (1889). It is extant. Its type is Carettochelys. It was considered monophyletic by Karl et al. (2012).
It was corrected as Carettochelydidae by Perrier (1928); it was reranked as the unranked clade Carettochelyidae by Joyce (2014).
It was assigned to Pleurodira by Gadow (1898); to Pleurodiri by Perrier (1928); to Trionychia by Kuhn (1946); to Carettochelyoidea by Kuhn (1966); to Testudines by Frank and Ramus (1996); to Chelonii by Alonso Santiago and Alonso Andrés (2005); to Trionychia by Joyce et al. (2004), Pérez-García et al. (2013) and Joyce (2014); and to Trionychoidea by Gaffney (1975), Carroll (1988), Karl et al. (2006), Tong et al. (2010), Karl et al. (2012), Vremir (2013) and van Dijk et al. (2014).
It was corrected as Carettochelydidae by Perrier (1928); it was reranked as the unranked clade Carettochelyidae by Joyce (2014).
It was assigned to Pleurodira by Gadow (1898); to Pleurodiri by Perrier (1928); to Trionychia by Kuhn (1946); to Carettochelyoidea by Kuhn (1966); to Testudines by Frank and Ramus (1996); to Chelonii by Alonso Santiago and Alonso Andrés (2005); to Trionychia by Joyce et al. (2004), Pérez-García et al. (2013) and Joyce (2014); and to Trionychoidea by Gaffney (1975), Carroll (1988), Karl et al. (2006), Tong et al. (2010), Karl et al. (2012), Vremir (2013) and van Dijk et al. (2014).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1847 | Apholidemys granosa Pomel |
| 1847 | Apholidemys sublaevis Pomel |
| 1872 | Anosteira radulina Cope p. 555 |
| 1873 | Anostira radulina Cope |
| 1884 | Anostira radulina Cope |
| 1889 | Carettochelyidae Gill |
| 1898 | Carettochelydidae Gadow p. 21 |
| 1902 | Anosteira radulina Hay p. 447 |
| 1908 | Anosteira radulina Hay p. 279 figs. Plate 43, figs 1,2; text-figs. 352-354. |
| 1928 | Carettochelydidae Perrier p. 3115 |
| 1930 | Anosteira radulina Hay p. 93 |
| 1946 | Carettochelyidae Kuhn p. 59 |
| 1966 | Carettochelyidae Kuhn p. 29 |
| 1975 | Carettochelyidae Gaffney p. 428 |
| 1987 | Chorlakkichelys de Broin pp. 174-177 figs. Pl 1, figs.1-16 |
| 1987 | Chorlakkichelys shahi de Broin pp. 174-177 figs. Pl 1, figs.1-16 |
| 1988 | Carettochelyidae Carroll |
| 1996 | Carettochelyidae Frank and Ramus |
| 2004 | Carettochelyidae Joyce et al. |
| 2005 | Carettochelyidae Alonso Santiago and Alonso Andrés |
| 2006 | Carettochelyidae Karl et al. |
| 2010 | Carettochelyidae Tong et al. p. 280 |
| 2012 | Carettochelyidae Karl et al. |
| 2013 | Carettochelyidae Pérez-García et al. |
| 2013 | Carettochelyidae Vremir |
| 2014 | Carettochelyidae Joyce |
| 2014 | Carettochelyidae van Dijk et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Carettochelyidae Gill 1889
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G. †Anosteira Leidy 1871
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†Anosteira manchuriana Zangerl 1947
†Anosteira maomingensis Chow and Liu 1955
†Anosteira ornata Leidy 1871
†Anosteira pulchra Clark 1932
†Anosteira shuvalovi Chkhikvadze 1979
Subfm. †Anosteirinae Lydekker 1889
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G. †Kyzilkumemys Nesov 1977
G. †Pseudanosteira Clark 1932
G. †Pseudotrionyx Dollo 1886
Unr. †Carettochelyinae Boulenger 1887
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Tr. †Allaeochelyini Godinot et al. 2018
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G. †Allaeochelys Noulet 1867
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†Allaeochelys crassesculpta Harrassowitz 1922
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Invalid names: Anosteira gracilis Harrassowitz 1922 [synonym]
†Allaeochelys delheidi Dollo 1886
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Invalid names: Allaeochelys casasecai Jimenez Fuentes 1971 [synonym], Allaeochelys jimenezi Alonso Santiago and Alonso Andrés 2005 [synonym], Anosteira anglica Lydekker 1889 [synonym]
Allaeochelys libyca Havlik et al. 2014
†Allaeochelys lingnanica Young and Chow 1962
†Allaeochelys magnifica Hutchison et al. 2004
†Allaeochelys parayrei Noulet 1867
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Invalid names: Allaeochelys nouleti Bergounioux 1931 [synonym], Castresia munieri De Stefano 1902 [synonym]
†Allaeochelys rouzilhacensis Godinot et al. 2018
G. †Burmemys Hutchison et al. 2004
G. †Hemichelys Lydekker 1887
G. Carettochelys Ramsay 1886 [pig-nose turtle]
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Carettochelys insculpta Ramsay 1886 [Pignose turtle]
†Carettochelys niahensis White et al. 2023
G. †Kizylkumemys Nessov 1977
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†Kizylkumemys khoratensis Tong et al. 2005
†Kizylkumemys schultzi Nessov 1977
Invalid names: Kizylkumemys schultzi mirabilis Nessov 1981 [nomen dubium]
Invalid names: Allaeochelyinae Bergounioux 1952 [empty], Anosteira radulina Cope 1872 [nomen dubium], Apholidemys granosa Pomel 1847 [nomen nudum], Apholidemys sublaevis Pomel 1847 [nomen nudum], Chorlakkichelys de Broin 1987 [nomen dubium], Chorlakkichelys shahi de Broin 1987 [nomen dubium]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| E. Perrier 1928 | Carapace recouverte par un tégument ayant la consistance du cuir, sans boucliers cornés, et contituée par des séries régulières de pièces osseuses; les plaques osseuses vertébrales au nombre de 6 seulement, toutes séparées les unes des autres par les plaques costales, qui se joignent sur la ligne médiane. Plastron formé des 9 os habituels. Membres en forme de palettes nageuses; les doigts très allongés, les deux internes seuls armés d'ongles. Cou non rétractile; alliés sous tous les autres rapports aux PLEURODIRA. |
Measurements
No measurements are available
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| Source: f = family, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Uetz 2005, Carroll 1988 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Barremian to the top of the Holocene or 130.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma
Collections (137 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Barremian | Thailand (Kalasin) | Kizylkumemys sp. (84443) | |
| Aptian | China (Guangxi) | Kizylkumemys sp. (24901 228852) | |
| Aptian | Thailand (Nakhon Ratchasima) | Kizylkumemys khoratensis (99337) | |
| Aptian - Albian | Laos (Savannakhet) | Carettochelyidae indet. (45092 118323) | |
| Aptian - Albian | Thailand (Nakhon Ratchasima) | Kizylkumemys khoratensis (48875) | |
| Aptian - Albian | Thailand (Ubon Ratchathani) | Kizylkumemys khoratensis (52346) | |
| Late/Upper Albian - Cenomanian | Uzbekistan (Navoi) | Kizylkumemys schultzi (58471) | |
| Cenomanian - Santonian | Mongolia (Dornogov) | Kizylkumemys schultzi (118167) | |
| Cenomanian - Santonian | Mongolia (Omnogov) | Anosteira shuvalovi (67974) | |
| Early/Lower Cenomanian | Uzbekistan (Navoi) | Kizylkumemys schultzi (58208 64685) | |
| Early/Lower Cenomanian | Uzbekistan (Karakalpakstan) | Kizylkumemys schultzi (58490 64689) | |
| Turonian | Japan (Iwate) | Carettochelyidae indet. (165694) | |
| Early/Lower Turonian | Uzbekistan | Carettochelyidae indet. (58596) | |
| Early/Lower Turonian | Uzbekistan (Karakalpakstan) | Carettochelyidae indet. (59698) | |
| Middle Paleocene | China (Guangdong) | Anosteira lingnanica (71846) | |
| Ypresian | France (Île-de-France) | Carettochelyidae indet. (68437) | |
| Ypresian | India (Gujarat) | Carettochelyidae indet. (180565) | |
| Ypresian | France (Alsace-Champagne-Ardenne-Lorraine) | Allaeochelys sp. (178052) | |
| Ypresian | France (Aude) | Allaeochelys sp. (78236) | |
| Ypresian | France (Marne) | Allaeochelys sp. (35533 35535 35539 35540 181108 181112 181653) | |
| Ypresian | France (Hérault) | Eulalichelys labarrerei (181563) | |
| Ypresian - Lutetian | France (Occitanie) | Allaeochelys rouzilhacensis (195457) | |
| Ypresian - Lutetian | Romania | Allaeochelys sp. (178481) | |
| Early/Lower Eocene | China (Jiangxi) | Anosteira sp. (182565) | |
| Early/Lower Eocene | France (Oise) | Apholidemys granosa, Apholidemys sublaevis (181647) | |
| Early/Lower Eocene | China (Shandong) | Anosteira shantungensis (154958) | |
| Early/Lower Eocene | France (Marne) | Allaeochelys sp. (181633) | |
| MP 10 | France (Picardie) | Allaeochelys sp. (68440) | |
| MP 8 + 9 | France (Picardie) | Allaeochelys sp. (84997) | |
| Wasatchian - Bridgerian | Canada (Northwest Territories) | Anosteirinae indet. (15584) | |
| Bridgerian | USA (Wyoming) | Anosteira ornata (15837 16201 16228 16235 16272 16273 27269 70266) Anosteira radulina (131861) Anostira ornata (130809) | |
| MP 11 | Germany (Hessen) | Anosteira crassesculpta (120036) Anosteira gracilis (22701) | |
| Lutetian | Pakistan (North-West Frontier) | Chorlakkichelys shahi (28978) | |
| Lutetian | Spain (Huesca) | Allaeochelys sp. (179886 179887) | |
| Lutetian | Spain (Zamora) | Allaeochelys casasecai (41308) Allaeochelys casasecai, Allaeochelys jimenezi (179692) Allaeochelys indet. (45232) Allaeochelys sp. (41073 41078 45233 45237 79704 224597) Carettochelyidae indet. (45097) | |
| Lutetian | France (Gironde) | Akrochelys neuvillei (181638) | |
| Lutetian | China (Liaoning) | Anosteira manchuriana (162504) | |
| Lutetian | Belgium (Brabant) | Pseudotrionyx delheidi (124459) | |
| Lutetian | Germany (Bavaria) | Allaeochelys sp. (187573) | |
| Lutetian - Bartonian | USA (Virginia) | Allaeochelys sp. (167197) | |
| Lutetian - Bartonian | China (Guangdong) | Anosteira maomingensis (151737 167230) | |
| Middle Eocene | China (Inner Mongolia) | Anosteira mongoliensis (178831) | |
| Middle Eocene | Spain (Zamora) | Allaeochelys casasecai, Allaeochelys jimenezi (207764) | |
| Middle Eocene | Mongolia | Anosteira mongoliensis (127656) | |
| Early/Lower Uintan | USA (Utah) | Anosteira pulchra (16572) | |
| Late/Upper Uintan | Canada (Saskatchewan) | Pseudanosteira sp. (16626) | |
| Late/Upper Uintan | USA (Texas) | Allaeochelys sp. (16542) Carettochelyidae indet. (16614) | |
| Uintan | USA (Wyoming) | Anosteira pulchra (198273) | |
| Uintan | USA (Utah) | Anosteira ornata (185691) Anosteira pulchra (186893 186895 186896 213058 213059 213060 213061 213062 213063 213064 213065 213066 213067 213068 213069 213070 213071 213112 213113 213134 213135 222799 222802) Pseudanosteira pulchra (16594) | |
| Bartonian | France (Midi-Pyrenees Region) | Allaeochelys parayrei, Castresia munieri (31634) | |
| Bartonian | Myanmar | Anosteira sp., Burmemys magnifica (41320 41456 41460 41461 41462 41463) Burmemys magnifica (41307 41310 41331 41455 41457 41459 41466) | |
| Bartonian | France (Occitanie) | Allaeochelys nouleti (184187) | |
| Bartonian | China (Henan) | Anosteira sp. (177036) | |
| Duchesnean | USA (Utah) | Carettochelyidae indet. (16877) | |
| Priabonian | United Kingdom (England) | Anostira anglica (39643) | |
| Chadronian | USA (South Dakota) | Dermatemydidae indet. (16885 16938) | |
| Late/Upper Eocene | China (Guangdong) | Anosteira maomingensis (144065) | |
| Early/Lower Miocene | Libya | Testudines indet. (13065) | |
| Burdigalian | Saudi Arabia | Carettochelyidae indet. (47021 47023 47074 47091 47092) | |
| Burdigalian | Germany | Allaeochelys parayrei (47425 49101) | |
| Burdigalian | Germany (Baden-Württemberg) | Allaeochelys parayrei (198614) | |
| MN 5 | Germany | Trionychidae indet. (46955) | |
| Late/Upper Pleistocene - Holocene | Malaysia | Carettochelys niahensis (192522) |