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Tupelocetus

Mammalia - Cetacea - Protocetidae

Taxonomy
Tupelocetus was named by Gibson et al. (2019). Its type is Tupelocetus palmeri.

It was assigned to Protocetidae by Gibson et al. (2019).

Species
T. palmeri (type species)

Synonymy list
YearName and author
2019Tupelocetus Gibson et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
RankNameAuthor
Mammaliaformes
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Artiodactylamorpha
Artiodactyla()
Whippomorpha
orderCetacea
familyProtocetidae
genusTupelocetus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Tupelocetus Gibson et al. 2019
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Tupelocetus palmeri Gibson et al. 2019
Diagnosis
ReferenceDiagnosis
M. L. Gibson et al. 2019Tupelocetus palmeri is unique among protocetids in having the following autapomorphy: a deeply excavated central fossa of the occiput. Differs from Protocetus atavus, Geor- giacetus vogtlensis, Kharodacetus sahnii, and Dhedacetus hyaeni in lacking a posteriorly positioned, accessory cusp on P2 (although heavily worn in Protocetus). The enamel of the P2 in Tupelocetus is more coarsely ornamented than those, or adjacent, teeth of Artiocetus clavis, Rodhocetus, Babiacetus indicus, Maiace- tus inuus, Aegyptocetus tarfa, and Dhedacetus hyaeni, and teeth referred to Togocetus traversei. Differs from Qaisracetus arifi, Rodhocetus, Maiacetus inuus, Artiocetus clavis, Gaviacetus razai, and Georgiacetus vogtlensis in having large nasal processes of the frontals. Unlike Protocetus atavus, Carolinacetus gingerichi, Rodhocetus, and Qaisracetus arifi, Tupelocetus has a parietal ridge. Further differs from Carolinacetus gingerichi and Maiacetus inuus in having the ascending process of the premaxilla terminate at the level of P2 or P3, instead of more anteriorly. The basicra- nium is only slightly narrower at the external auditory meatus, as compared with the abrupt narrowing seen in Georgiacetus vogtlensis, Protocetus atavus, Artiocetus clavis, Gaviacetus razai, and Maiacetus inuus. Differs from Artiocetus clavis, Qaisracetus arifi, and Gaviacetus razai in a foramen ‘pseudo-ovale’ that is open ventrally. Further differs from Carolinacetus gingerichi, Georgiacetus vogtlensis, Artiocetus clavis, and Gaviacetus razai in having a lateral lip of the entoglenoid process that slightly over- hangs adjacent parts of the squamosal, with the skull in ventral view. Tupelocetus can be differentiated from the fragmentary holotype of Indocetus ramani in having a much longer external auditory meatus and posterior process of the tympanic, relative to the distances from these structures to the entoglenoid process. Likely differs from Crenatocetus rayi, Takracetus simus, and Togocetus traversei in being significantly larger, although this cannot be confirmed because comparable elements are lacking. Preserved material does not allow Tupelocetus palmeri to be distinguished from Natchitochia jonesi and Pappocetus lugardi (see Discussion).