PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Protictis

Mammalia - Viverravidae

Taxonomy

Didymictis (Protictis) was named by Matthew (1937).

It was reranked as Protictis by MacIntyre (1966), Flynn and Galiano (1982), Flynn (1998), Meehan and Wilson (2002), Wesley-Hunt and Werdelin (2005) and Solé et al. (2014).

It was assigned to Didymictis by Matthew (1937); to Didymictidae by Flynn and Galiano (1982) and Flynn (1998); and to Viverravidae by MacIntyre (1966), Meehan and Wilson (2002), Wesley-Hunt and Werdelin (2005) and Solé et al. (2014).

Subtaxa

P. (Bryanictis), P. (Protictis), P. (Protictoides), P. agastor, P. haydenianus (syn. Didymictis primus, Prolimnocyon macfaddeni), P. minor, P. paralus, P. simpsoni

Sister taxa

P. (Bryanictis), P. (Protictis), P. (Protictoides)

Synonymy list

Year	Name and author
1937	Didymictis (Protictis) Matthew
1966	Protictis MacIntyre
1982	Protictis Flynn and Galiano p. 24
1998	Protictis Flynn
2002	Protictis Meehan and Wilson p. 1092
2005	Protictis Wesley-Hunt and Werdelin p. 842
2014	Protictis Solé et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()

Rank	Name	Author
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Theriamorpha	(Rowe 1993)
—	Theriiformes	()
—	Trechnotheria
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Ferae	()
—	Carnivoramorpha	Wyss and Flynn 1993
family	Viverravidae
genus	Protictis	(Matthew)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Protictis Matthew

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Subg. †Protictis (Bryanictis) MacIntyre 1966

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†Bryanictis microlestes Simpson 1935

†Bryanictis paulus Meehan and Wilson 2002

Subg. †Protictis (Protictis) Matthew 1937

Subg. †Protictis (Protictoides) Flynn and Galiano 1982

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†Protictoides aprophatos Flynn and Galiano 1982

†Protictis agastor Gingerich and Winkler 1985

†Protictis haydenianus Cope 1882

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Invalid names: Didymictis primus Cope 1884 [synonym], Prolimnocyon macfaddeni Rigby, Jr. 1980 [synonym]

†Protictis minor Meehan and Wilson 2002

†Protictis paralus Holtzman 1978

†Protictis simpsoni Meehan and Wilson 2002

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Life habit:

ground dwellingf

Diet:

carnivoref

Reproduction:

viviparoussubc

Created:

2005-08-30 14:47:14

Modified:

2005-08-30 16:47:14

Source: f = family, subc = subclass, c = class, subp = subphylum

References: Flynn 1998, Lillegraven 1979, Hendy et al. 2009, Ji et al. 2002, Carroll 1988

Age range: base of the Ti3 to the top of the Uintan or 61.70000 to 39.70000 Ma

Collections (54 total)

Time interval	Ma	Country or state	Original ID and collection number
Danian	66.0 - 61.6	USA (North Dakota)	P. paralus (14962)
To2	63.8 - 60.9	Canada (Alberta)	P. sp. (14894)
Torrejonian	63.8 - 60.9	Canada (Alberta)	Bryanictis microlestes (143200) P. sp. (14822)
Torrejonian	63.8 - 60.9	USA (Wyoming)	Bryanictis microlestes, P. haydenianus (14912) Prolimnocyon macfaddeni, P. haydenianus (14886) P. haydenianus, P. microlestes (14896) P. microlestes (14899 14900 14901 14904 14905) P. sp., P. microlestes (14897 14898)
Torrejonian	63.8 - 60.9	USA (Utah)	P. haydenianus (14830)
Torrejonian	63.8 - 60.9	USA (Texas)	Bryanictis sp. (14829)
Torrejonian	63.8 - 60.9	USA (New Mexico)	Bryanictis paulus, Didymictis sp. b, Didymictis sp. a (14816) Didymictis haydenianus (14819 14873) P. haydenianus (14812 14815 14824 14832 14833 14835 14862 14865 14893) P. sp. (14868)
Torrejonian	63.8 - 60.9	USA (Montana)	Didymictis haydenianus (14919 14920 14923) Didymictis microlestes (14917) Didymictis microlestes, Didymictis haydenianus (14913) P. haydenianus, P. microlestes (14906)
Ti3	61.7 - 56.8	Canada (Alberta)	P. paralus (14928 14992)
Ti4	61.7 - 56.8	Canada (Saskatchewan)	P. paralus (14978)
Tiffanian	61.7 - 56.8	USA (North Dakota)	P. microlestes (15000) P. paralus (14931)
Tiffanian	61.7 - 56.8	USA (Colorado)	P. proteus (174227)
Tiffanian	61.7 - 56.8	USA (Wyoming)	P. agastor, P. paralus (14936) P. haydenianus (14974) P. microlestes (15014 15067) P. paralus (15027) P. paralus, P. haydenianus (15017)
Tiffanian	61.7 - 56.8	Canada (Alberta)	P. sp. (15005)
Tiffanian	61.7 - 56.8	USA (California)	P. agastor, P. paralus (154724)
Tiffanian	61.7 - 56.8	USA (Montana)	P. haydenianus, P. microlestes (15048) P. sp. (15025 15064)
Wasatchian	55.8 - 50.3	USA (Wyoming)	P. sp. (46782)
Uintan	46.2 - 39.7	USA (Wyoming)	P. aprophatos (16630)