Gatun [middle Gatun Fm]: Tortonian, Panama
collected by Olsson

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bivalvia - Arcida - Glycymerididae
Glycymeris canalis Brown and Pilsbry 1911
recombined as Tucetona pectinata canalis
Bivalvia - Arcida - Arcidae
Arca (Scapharca) dariensis (Brown and Pilsbry 1911)
recombined as Anadara (Rasia) dariensis
Bivalvia - Pectinida - Pectinidae
Pecten gatunensis (Toula 1909)
recombined as Leopecten gatunensis
Pecten (Chlamys) levicostatus (Toula 1909)
recombined as Argopecten levicostatus
Amusium luna (Brown and Pilsbry 1913)
synonym of Amusium toulae
Bivalvia - Pectinida - Propeamussiidae
Pecten oligolepis (Brown and Pilsbry 1913)
recombined as Cyclopecten oligolepis
Bivalvia - Pholadida - Corbulidae
Corbula hexacyma Brown and Pilsbry 1913
recombined as Hexacorbula hexacyma
Bivalvia - Cardiida - Cardiidae
Protocardia (Lophocardium) gurabica (Maury 1917)
recombined as Nemocardium (Lophocardium) gurabicum
Cardium (Trachycardium) dominicense Gabb 1873
recombined as Trachycardium (Dallocardia) dominicense
Cardium (Trachycardium) stiriatum Brown and Pilsbry 1911
recombined as Trachycardium (Phlogocardia) stiriatum
Bivalvia - Cardiida - Tellinidae
Tellina (Eurytellina) dariena Conrad 1855
synonym of Eurytellina punicea
Bivalvia - Cardiida - Veneridae
Chione mactropsis (Conrad 1855)
recombined as Lirophora (Panchione) mactropsis
Dosinia acetabulum (Conrad 1832)
Chione rowleei Olsson 1922
Callocardia gatunensis (Dall 1903)
recombined as Callocardia (Agriopoma) gatunensis
Pitaria circinata (Born 1778)
recombined as Lamelliconcha circinata
Clementia dariena (Conrad 1855)
recombined as Clementia (Clementia) dariena
Bivalvia - Cardiida - Chamidae
Echinochama yaquensis (Maury 1917)
recombined as Arcinella yaquensis
Bivalvia - Nuculanida - Nuculanidae
Leda balboae Brown and Pilsbry 1911
recombined as Saccella balboae
Gastropoda - Heterostropha - Architectonicidae
Architectonica granulata (Lamarck 1822)
synonym of Architectonica nobilis
Gastropoda - Turritellidae
Turritella mimetes Brown and Pilsbry 1911
recombined as Turritella (Broderiptella) mimetes
Turritella gatunensis Conrad 1857
Turritella altilira Conrad 1857
recombined as Turritella (Bactrospira) altilira
Gastropoda - Naticidae
Natica guppyana (Toula 1909)
recombined as Stigmaulax guppiana
Natica canrena (Linnaeus 1758)
recombined as Naticarius canrena
Polinices subclausa (Sowerby 1850)
synonym of Polinices hepaticus
Gastropoda - Vermetidae
Petaloconchus sculpturatus Lea 1846
Gastropoda - Neogastropoda - Mitridae
Mitra longa Gabb 1873
recombined as Subcancilla longa
Mitra dariensis Brown and Pilsbry 1911
recombined as Subcancilla dariensis
Gastropoda - Neogastropoda - Columbellidae
Strombina lessepsiana Brown and Pilsbry 1911
recombined as Strombina (Strombina) lessepsiana
Gastropoda - Neogastropoda - Buccinidae
Metula gabbi Brown and Pilsbry 1911
Phos subsemicostatus Brown and Pilsbry 1911
recombined as Cymatophos subsemicostatus
Solenosteira dalli Brown and Pilsbry 1911
recombined as Hanetia dalli
Phos gatunensis Toula 1909
recombined as Antillophos (Antillophos) candei gatunensis
Gastropoda - Neogastropoda - Fasciolariidae
Fasciolaria gorgasiana Brown and Pilsbry 1913
recombined as Pleuroploca gorgasiana
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria dariena Toula 1908
recombined as Bivetiella dariena
Cancellaria epistomifera Guppy 1876
    = Bivetiella dariena Toula 1908
Landau et al. 2012
Gastropoda - Neogastropoda - Turridae
Turris albida (Perry 1811)
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Oldest record of Polystira albida is Cayo Agua Fm (c. 4.25 Ma) per Todd & Johnson, 2013
Gastropoda - Neogastropoda - Conidae
Conus imitator Brown and Pilsbry 1911
Conus concavitectum Brown and Pilsbry 1911
synonym of Conus molis
Conus veatchi Olsson 1922
Gastropoda - Neogastropoda - Drilliidae
Drillia venusta (Sowerby 1850)
recombined as Clathrodrillia venusta
Gastropoda - Neogastropoda - Terebridae
Terebra gatunensis (Toula 1908)
recombined as Strioterebrum gatunensis
Gastropoda - Neogastropoda - Marginellidae
Marginella leander Brown and Pilsbry 1911
recombined as Volvarina leander
Marginella mindiensis Cossmann 1913
Marginella latissima Dall 1896
recombined as Prunum latissimum
SUBSPECIES: Marginella latissima pilsbryi
Gastropoda - Neogastropoda - Muricidae
Murex messorius Sowerby 1841
recombined as Vokesimurex messorius
Typhis alatus Sowerby 1850
recombined as Typhina alata
Gastropoda - Strombidae
Strombus gatunensis Toula 1908
Gastropoda - Personidae
Distortrix simillima (Sowerby 1850)
recombined as Distorsio simillima
Gastropoda - Cassidae
Sconsia laevigata (Guppy 1850)
SUBSPECIES: Sconsia laevigata gabbi
    = Sconsia grayi Adams 1855
Beu 2010
Gastropoda - Neotaenioglossa - Calyptraeidae
Crepidula plana Say 1822
see common names

Geography
Country:Panama
Coordinates: 9.3° North, 79.9° West (view map)
Paleocoordinates:9.0° North, 78.5° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.63000 - 7.24600 m.y. ago
Age estimate:9.9 Ma (other)
Stratigraphy
Formation:Gatun Member:middle Gatun
Stratigraphic resolution:formation
Stratigraphy comments: LITHOSTRATIGRAPHY: From the Gatun Formation, which unconformably overlies either volcanic basement of Caimito Formation (late Oligocene), and is unconformably overlain by the Chagres Formation. AGE: Late Miocene; constrained by biostratigraphy of multiple groups (foraminifera, nannoplankton); refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 9.9 Ma (Hendy, unpublished data). STRATIGRAPHIC POSITION: Probably from top of middle member (enterer's opinion).
Lithology and environment
Primary lithology: "siliciclastic"
Includes fossils?Y
Lithology description: LITHIFICATION: Unknown. LITHOLOGY: No lithologic data provided, but assumed to be siliciclastic.
Environment:coastal indet.
Geology comments: No paleoenvironmental data provided, but assumed to be from shallow marine facies (enterer's opinion).
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Museum repositories:PRI
Collectors:Olsson
Collection method comments: COLLECTOR: Olsson, A. REPOSITORY: PRI.
Taxonomic list comments:NOMENCLATURE: Authoritative publication, with species-resolution identifications, and antiquated nomenclature. COVERAGE: Exhaustive for mollusk.
Metadata
Database number:134661
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2012-10-12 06:14:30 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2014-10-12 06:14:30
Creative Commons license:CC BY
Reference information

Primary reference:
43229. A. A. Olsson. 1922. The Miocene of Northern Costa Rica. Bulletins of American Paleontology 9(39):1-309 [A. Hendy/A. Hendy]

Secondary references:

30037 A. G. Beu. 2010. Neogene Tonnoidean Gastropods of Tropical and South America; contributions to the Dominican Republic and Panama Paleontology Projects and Uplift of the Central American Isthmus. Bulletins of American Paleontology (377-378)1-550 [A. Hendy/A. Hendy/A. Hendy]
43491 B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy]
48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]