Masia de l'Hereuet: Ypresian, Spain

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Squamata
Squamata indet. Oppel 1811
Bolet 2017 9 specimens
IPS 49742, fragment of left dentary; IPS 49746, anterior portion of a right dentary with two teeth; IPS 59530, vertebra; IPS 59532, tooth; IPS 59539, tooth; IPS 83542, seven fragments of toothbearing bones; IPS 83548, four fragments of dentary; IPS 83550, two fragments of tooth-bearing bone; IPS 83541, roughly rounded osteoderm
Reptilia - Anguidae
Anguinae indet. Gray 1825
Bolet 2017 3 specimens
IPS 83557, partial parietal; IPS 59538, two partial vertebrae; IPS 83540, two body osteoderms
Glyptosaurinae indet. Marsh 1872
Bolet 2017 9 specimens
IPS 35741, three fragments of body osteoderm; IPS 49738 fragment of body osteoderm; IPS 59536, four fragments of body osteoderms; IPS 83547, seven body osteoderms or fragments of body osteoderms, one partial skull osteoderm; IPS 83549, 10 body osteoderms and fragments of osteoderm; IPS 49744, tooth-bearing bone (?dentary) fragment with one tooth; IPS 83539, several dozen osteoderms; IPS 83551, four fragments of osteoderms; IPS 83554, body osteoderms
Glyptosaurini indet. Sullivan 1979
Bolet 2017 1 specimen
IPS 3742, partial skull osteoderm
Reptilia - Necrosauridae
"Necrosauridae" indet. Hoffstetter 1943
Bolet 2017 2 specimens
IPS 83545, fragment of dentary; IPS 49741, partial osteoderm
Reptilia
"Scincomorpha" indet. (Camp 1923)
Bolet 2017 2 specimens
IPS 49743, fragment of ?maxilla bearing two teeth; IPS 83544, fragment of left dentary with four teeth (three of them are broken)
Amphisbaenia indet. (Gray 1844)
Bolet 2017 1 specimen
IPS 59529, dorsal vertebra
Reptilia - Iguanidae
Geiseltaliellus sp. Kuhn 1944
Bolet 2017 6 specimens
IPS 49740, partial left dentary with 10 tooth; IPS 59535, fragment of right dentary; IPS 83552, partial left maxilla with six teeth; IPS 83558, two partial right and two partial left dentaries, one partial maxilla; less certainly IPS 49745, fragment of ?maxilla with two teeth; IPS 83553, partial right dentary
Reptilia - Agamidae
Agamidae indet. Gray 1827
Bolet 2017 2 specimens
IPS 83546, fragment of tooth-bearing bone with one tooth; IPS 83543, fragment of left dentary with two teeth
Reptilia
Scincoidea indet. Oppel 1811
Bolet 2017 1 specimen
IPS 49761, fragment of tooth-bearing bone with one tooth
Mammalia - Primates - Paromomyidae
Arcius sp. Godinot 1984
Mammalia - Primates - Notharctidae
Agerinia marandati n. sp. Femenias-Gual et al. 2017
Femenias-Gual et al. 2017
ICP IPS-82801 - type
Mammalia - Rodentia - Ischyromyidae
? Ischyromyidae sp. Alston 1876
Mammalia - Lipotyphla
? Lipotyphla sp. Haeckel 1866
synonym of Eulipotyphla
Mammalia - Diacodexeidae
Diacodexis sp. Cope 1882
Mammalia - Perissodactyla - Palaeotheriidae
Propachynolophus sp. Lemoine 1891
Mammalia - Perissodactyla - Lophiodontidae
Lophiodon sp. Cuvier 1822
Mammalia - Apatemyidae
Heterohyus ilerdensis n. sp. (López-Torres and Silcox 2018)
Beard and Métais 2024
Mammalia
? Marsupialia sp. (Illiger 1811)
see common names

Geography
Country:Spain State/province:Lleida County:Pallars Jossà
Coordinates: 42.0° North, 0.7° East (view map)
Paleocoordinates:37.3° North, 0.9° West
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
Key time interval:Ypresian Mammal zone: MP 8 + 9
Age range of interval:56.00000 - 47.80000 m.y. ago
Age estimate:51 ± 56 Ma (unknown)
Stratigraphy
Formation:Corca
Stratigraphic resolution:bed
Stratigraphy comments: "The site was assigned to the Grauvian (MP10) by Antunes et al. (1997) on the basis of its mammalian fauna. However, after revision of the artiodactyl, perissodactyl, and rodent assemblage, Badiola et al. (2009) concluded that this level is older than previously considered, corresponding to the Neustrian (MP8+9, 51-56 Ma), and represents the oldest continental locality of the Àger sub-basin" (p.430)
Lithology and environment
Primary lithology: claystone
Secondary lithology: sandstone
Includes fossils?Y
Includes fossils?Y
Environment:fluvial indet.
Taphonomy
Modes of preservation:body
Size of fossils:mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Metadata
Database number:142031
Authorizer:J. Alroy, P. Mannion, P. Barrett, K. Beard Enterer:A. Garcia Selles, P. Mannion, K. Beard, T. Cleary
Modifier:P. Mannion Research group:vertebrate
Created:2013-04-04 11:29:45 Last modified:2020-06-17 06:53:58
Access level:the public Released:2013-04-04 11:29:45
Creative Commons license:CC BY
Reference information

Primary reference:
46138. J. Marigó, R. Minwer-Barakat, S. Moyà-Solà and S. López-Torres. 2012. First record of Plesiadapiformes (Primates, Mammalia) from Spain. Journal of Human Evolution 62(3):429-433 [J. Alroy/A. Garcia Selles]

Secondary references:

87057 K. C. Beard and G. Métais. 2024. Oldest record of Apatemyidae (Mammalia, Apatotheria) from Spain and the taxonomic status of Spanish paromomyids (Mammalia, Primatomorpha). Journal of Vertebrate Paleontology [K. Beard/K. Beard]
62826 A. Bolet. 2017. First early Eocene lizards from Spain and a study of the compositional changes between late Mesozoic and early Cenozoic Iberian lizard assemblages . Palaeontologia Electronica 20(2):1-22 [P. Barrett/T. Cleary]
62202 J. Femenias-Gual, R. Minwer-Barakat, J. Marigó, M. Poyatos-Moré, and S. Moyà-Solà. 2017. Agerinia marandati sp. nov., a new early Eocene primate from the Iberian Peninsula, sheds new light on the evolution of the genus Agerinia. PeerJ 5:e3239:1-35 [P. Mannion/P. Mannion]
66265 S. López-Torres and M. T. Silcox. 2018. The European Paromomyidae (Primates, Mammalia): taxonomy, phylogeny, and biogeographic implications. Journal of Paleontology [P. Mannion/P. Mannion]