PBDB Collection

Sand Quarry: Wasatchian, Colorado

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Reptilia - Anguidae

	Anguidae indet. Gray 1825	ElShafie 2024
Mammalia - Multituberculata - Eucosmodontidae

	Neoliotomus ultimus (Granger and Simpson 1928)
	original and current combination Eucosmodon ultimus
	includes "cf. Prochetodon sp.": Krause 1982
Mammalia - Multituberculata - Neoplagiaulacidae

	Ectypodus cf. hazeni Jepsen 1940
	= ? Neoplagiaulacidae indet. Ameghino 1890	Krause 1982

	Parectypodus childei (Kühne 1969)
	= Parectypodus lunatus Krause 1982	Krause 1982
	replaces ID of Sloan 1981

	Ectypodus tardus (Jepsen 1940)	Krause 1982
Mammalia - Leptictidae

	Palaeictops tauricinerei (Jepsen 1930)
	= cf. Prodiacodon tauricinerei Jepsen 1930	Alroy 2002

	Didelphodus absarokae (Cope 1881)
Mammalia - Apatemyidae

	Apatemys cf. kayi (Simpson 1929)
	= Labidolemur kayi Simpson 1929	Gunnell 1989
Mammalia - Esthonychidae

	Esthonyx spatularius Cope 1880	Gunnell 1989
Mammalia - Macroscelidea - Amphilemuridae

	Entomolestes nitens Matthew 1918
	= cf. Macrocranion nitens Matthew 1918	Alroy 2002
Mammalia - Macroscelidea - Apheliscidae

	Apheliscus sp. Cope 1875	Rose 1981
	: "A. nitidus"

	Haplomylus speirianus (Cope 1880)
Mammalia - Condylarthra - Hyopsodontidae

	Hyopsodus miticulus (Cope 1874)
	= Hyopsodus simplex Loomis 1905	Redline 1997
	replaces ID of Gunnell 1989

	Hyopsodus loomisi McKenna 1960
Mammalia - Creodonta

	Prototomus viverrinus Cope 1874
	= Creodonta indet. Cope 1875	Gingerich and Deutsch 1989
	could be Prolimnocyon atavus; not discussed by Polly 1993
Mammalia - Creodonta - Hyaenodontidae

	Prototomus secundarius Cope 1875	Polly 1993
	robustus [Polly 1993: "Sinopa sp."
Mammalia

	Miacis exiguus Matthew and Granger 1915
	probably OK, see Gingerich 1983a
Mammalia - Viverravidae

	Viverravus acutus Matthew and Granger 1915

	Didymictis protenus (Cope 1874)
	= Didymictis leptomylus Cope 1880	Polly 1997
Mammalia - Perissodactyla - Equidae

	Hyracotherium angustidens (Cope 1875)
	= Hyracotherium sp. Owen 1841	Alroy 2002
	= Arenahippus grangeri Gingerich 1989	Froehlich 2002
	recombined as Sifrhippus grangeri
Mammalia - Perissodactyla

	Homogalax primaevus (Wortman 1896)
	= Cardiolophus radinskyi Gingerich 1991	Gingerich 1991
Mammalia - Diacodexeidae

	Bunophorus robustus (Sinclair 1914)	Stucky and Krishtalka 1990
	apparently described by McKenna as "Wasatchia sp.," but not on his list

	Diacodexis metsiacus (Cope 1882)
	= Diacodexis primus Krishtalka and Stucky 1985	Krishtalka and Stucky 1985
Mammalia - Procreodi - Arctocyonidae

	Thryptacodon antiquus Matthew and Granger 1915

	Chriacus gallinae Matthew and Granger 1915
Mammalia - Rodentia - Ischyromyidae

	Paramys taurus Korth 1984	Korth 1984
	species "A" of McKenna
Mammalia - Primates - Notharctidae

	Pelycodus mckennai Gingerich and Simons 1977
	= Cantius mckennai Gingerich and Simons 1977	Gingerich and Simons 1977
	"P. ralstoni"
Mammalia - Primates - Omomyidae

	Tetonius homunculus (Cope 1882)
	= Tetonius matthewi Bown and Rose 1987	Bown and Rose 1987
Mammalia - Primates - Microsyopidae

	Cynodontomys alfi McKenna 1960
	= Microsyops angustidens Matthew 1915	Gunnell 1985
Mammalia - Primates - Paromomyidae

	Phenacolemur jepseni Simpson 1955
	= cf. Simpsonlemur jepseni Simpson 1955	Alroy 2002
Mammalia

	Peradectes protinnominatus McKenna 1960

	Peradectes elegans Matthew and Granger 1921	Krishtalka and Stucky 1983

	Herpetotherium innominatum (Simpson 1928)
	= Copedelphys innominata Simpson 1928	Krishtalka and Stucky 1983
	recombined as Herpetotherium innominatum
	includes Peratherium macgrewi as a subspecies
see common names

Geography

Country:	United States	State/province:	Colorado	County:	Moffat
Coordinates:	40.6° North, 108.1° West (view map)
Paleocoordinates:	45.4° North, 91.9° West
Basis of coordinate:	based on political unit
Geographic resolution:	small collection

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Ypresian	10 m.y. bin:	Cenozoic 2
*Period:	Tertiary
Key time interval:	Wasatchian
Age range of interval:	54.90000 - 50.50000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:

Wasatch

Lithology and environment

Environment:

terrestrial indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils

Collection methods and comments

Collection size:	594 specimens
Museum repositories:	UCMP

Metadata

Also known as:	UCMP V-5421
Database number:	15725
Authorizer:	J. Alroy, J. Marcot, P. Holroyd	Enterer:	J. Alroy, P. Holroyd, J. Marcot
Modifier:	J. Alroy	Research group:	vertebrate
Created:	1993-09-27 00:00:00	Last modified:	2009-07-15 19:53:42
Access level:	the public	Released:	1993-09-27 00:00:00
Creative Commons license:	CC BY

Reference information

Primary reference:

2528.

M. C. McKenna. 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, Eocene of northwest Colorado. University of California Publications in Geological Sciences 37(1):1-130 [J. Alroy/J. Alroy/M. Uhen]

Secondary references:

6294		J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
1123		T. M. Bown and K. D. Rose. 1987. Patterns of Dental Evolution in Early Eocene Anaptomorphine Primates (Omomyidae) from the Bighorn Basin, Wyoming. Paleontological Society Memoir 23:1-162 [J. Alroy/J. Alroy/J. Ju]
87093		S. J. ElShafie. 2024. Body size estimation from isolated fossil bones reveals deep time evolutionary trends in North American lizards. PLoS ONE 19(1:e0296318):1-25 [P. Holroyd/P. Holroyd/P. Holroyd]
53105		D. J. Froehlich. 2002. Quo vadis Eohippus? The systematics and taxonomy of the early Eocene equids (Perissodactyla). Zoological Journal of the Linnean Society 134:141-256 [M. Carrano/M. Carrano/M. Carrano]
1660		P. D. Gingerich. 1991. Systematics and Evolution of Early Eocene Perissodactyla (Mammalia) in the Clarks Fork Basin, Wyoming. Contributions from the Museum of Paleontology, University of Michigan 28(8):181-213 [J. Alroy/J. Alroy/P. Mannion]
1666		P. D. Gingerich and H. A. Deutsch. 1989. Systematics and Evolution of Early Eocene Hyaenodontidae (Mammalia, Creodonta) in the Clarks Fork Basin, Wyoming. Contributions from the Museum of Paleontology, University of Michigan 27(13):327-391 [J. Alroy/J. Alroy/J. Marcot]
1675		P. D. Gingerich and E. L. Simons. 1977. Systematics, phylogeny, and evolution of early Eocene Adapidae (Mammalia, Primates) in North America. Contributions from the Museum of Paleontology, University of Michigan 24(22):245-279 [J. Alroy/J. Alroy/M. Uhen]
1728		G. F. Gunnell. 1985. Systematics of Early Eocene Microsyopinae (Mammalia, Primates) in the Clark's Fork Basin, Wyoming. Contributions from the Museum of Paleontology, University of Michigan 27(2):51-71 [J. Alroy/J. Alroy/M. Uhen]
1730		G. F. Gunnell. 1989. Evolutionary History of Microsyopoidea (Mammalia, ?Primates) and the Relationship Between Plesiadapiformes and Primates. University of Michigan Papers on Paleontology 27:1-157 [J. Alroy/J. Alroy/P. Mannion]
2105		W. W. Korth. 1984. Earliest Tertiary evolution and radiation of rodents in North America. Bulletin of Carnegie Museum of Natural History 24:1-71 [J. Alroy/J. Alroy/J. Marcot]
2158		D. W. Krause. 1982. Multituberculates from the Wasatchian Land-Mammal Age, Early Eocene, of Western North America. Journal of Paleontology 56(2):271-294 [J. Alroy/J. Alroy/J. Ju]
2179		L. Krishtalka and R. K. Stucky. 1983. Paleocene and Eocene marsupials of North America. Annals of Carnegie Museum 52(10):229-264 [J. Alroy/J. Alroy/M. Uhen]
2180		L. Krishtalka and R. K. Stucky. 1983. Revision of the Wind River faunas, early Eocene of central Wyoming. Part 3. Marsupialia. Annals of Carnegie Museum 52(9):205-227 [J. Alroy/J. Alroy/J. Alroy]
2183		L. Krishtalka and R. K. Stucky. 1985. Revision of the Eocene Wind River Faunas, Early Eocene of Central Wyoming. Part 7. Revision of Diacodexis (Mammalian, Artiodactyla). Annals of Carnegie Museum 54(14):413-486 [J. Alroy/J. Alroy/J. Marcot]
2785		P. D. Polly. 1993. Hyaenodontidae (Creodonta, Mammalia) from the Early Eocene Four Mile Fauna and their biostratigraphic implications. PaleoBios 14(4):1-10 [J. Alroy/J. Alroy/J. Ju]
2787		P. D. Polly. 1997. Ancestry and Species Definition in Paleontology: A Stratocladistic Analysis of Paleocene-Eocene Viverravidae (Mammalia, Carnivora) from Wyoming. Contributions from the Museum of Paleontology, University of Michigan 30(1):1-53 [J. Alroy/J. Alroy/M. Uhen]
6223		A. D. Redline. 1997. Revision of the Wind River faunas, early Eocene of central Wyoming. Part 13. Systematics and phylogenetic pattern of early Eocene Hyopsodus (Mammalia: Condylarthra). Annals of Carnegie Museum 66(1):1-81 [J. Alroy/J. Alroy/J. Alroy]
2953		K. D. Rose. 1981. The Clarkforkian Land-Mammal Age and Mammalian Faunal Composition Across the Paleocene-Eocene Boundary. University of Michigan Papers on Paleontology 26:1-197 [J. Alroy/J. Alroy/P. Mannion]
3335		R. K. Stucky and L. Krishtalka. 1990. Revision of the Wind River Faunas, Early Eocene of Central Wyoming. Part 10. Bunophorus (Mammalia, Artiodactyla). Annals of Carnegie Museum 59(2):149-171 [J. Alroy/J. Alroy/J. Marcot]