Brooksville 2: Arikareean, Florida

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia
Herpetotherium sp. Cope 1873
said to be new
Mammalia - Chiroptera - Emballonuridae
Emballonuridae indet. Dobson 1875
a bat
Mammalia - Chiroptera - Phyllostomidae
Phyllostomatidae indet. Savage 1951
Czaplewski and Morgan 2002
synonym of Phyllostomidae
a bat; large sized
Mammalia - Chiroptera - Mormoopidae
Mormoopidae indet. Saussure 1860
a bat
Koopmanycteris palaeomormoops n. gen., n. sp. Morgan et al. 2019
Morgan et al. 2019 42 specimens
UF 157765 (holotype), right dentary with p4-m3 and alveoli for i1-p3
Mammalia - Chiroptera - Vespertilionidae
Vespertilionidae indet. Gray 1821
a bat
Mammalia - Perissodactyla - Equidae
Miohippus sp. Marsh 1874
Mammalia - Hypertragulidae
Nanotragulus loomisi Lull 1922
Mammalia - Camelidae
Nothokemas waldropi Frailey 1978
Mammalia - Merycoidodontidae
Phenacocoelinae indet. Schultz and Falkenbach 1950
invalid subgroup of Merycoidodontidae
Mammalia - Carnivora - Palaeogalidae
Palaeogale minuta (Gervais 1848)
(1 measurement)
Mammalia - Carnivora - Mustelidae
Arikarictis chapini n. sp. Hayes 2000
(3 measurements)
Acheronictis webbi n. sp. Hayes 2000
(3 measurements)
Mammalia - Carnivora - Canidae
Osbornodon wangi n. sp. Hayes 2000
(3 measurements)
Enhydrocyon cf. pahinsintewakpa (Macdonald 1963)
Phlaocyon taylori n. sp. Hayes 2000
(7 measurements)
Mammalia - Geolabididae
Centetodon magnus (Clark 1936)
Mammalia - Erinaceidae
Parvericius montanus Koerner 1940
(1 measurement)
Mammalia - Lagomorpha - Leporidae
Megalagus abaconis n. sp. Hayes 2000
(6 measurements)
Mammalia - Rodentia - Sciuridae
Sciuridae indet. Gray 1821
Mammalia - Rodentia - Florentiamyidae
Florentiamyidae indet. Wood 1936
Mammalia - Rodentia - Eomyidae
Eomyidae indet. Winge 1887
Mammalia - Rodentia - Heliscomyidae
Heliscomyidae indet. Korth et al. 1991
Mammalia - Rodentia - Castoridae
Agnotocastor sp. Stirton 1935
Mammalia - Rodentia - Entoptychidae
Entoptychinae indet. Miller and Gidley 1918
spelled with current rank as Entoptychidae
said to be new
Reptilia - Scincidae
Scincidae indet. Gray 1825
Chovanec 2014 5 specimens
"UF 274043 (partial right dentary with splenial), UF 274063 (partial right dentary), UF 274239 (partial right dentary), UF 274240 (partial right dentary), UF 274252 (right premaxilla)"
Reptilia
Scolecophidia indet. (Duméril and Bibron 1844)
Mead 2013 3 specimens
UF 268995, -96, 269073, three trunk vertebrae
Reptilia - Polychrotidae
Anolis informal morphotype A
Chovanec 2014 36 specimens
"UF 274039 (partial dentary), 274041 (partial dentary), 274061 (frontal), 274073 (partial dentary), 274074 (partial dentary), 275606 (partial maxilla), 275608 (partial dentary), 275611 (partial dentary), 275620 (partial dentary), 275627 (left postorbital), 275629 (partial dentary), 275645-275647 (partial dentaries), 275649-650 (partial dentaries), 275653 (partial left maxilla), 275667 (braincase), 275677-275681 (frontals), 275682-275684 (left quadrates), 275685-275686 (frontals), 275693-275694 (partial maxillae), 275697 (partial dentary), 275699-275701 (partial dentaries), 278727 (left prefrontal), 278745 (left prefrontal)"
cf. Anolis informal morphotype B
Chovanec 2014 5 specimens
"UF 275662 (partial right dentary), UF 275695 (partial right dentary), UF 275696 (partial right dentary), UF 275619 (partial left dentary), UF 275665 (partial left dentary)"
Reptilia - Iguanidae
Iguanidae indet. Oppel 1811
Chovanec 2014 3 specimens
"UF 275612 (partial left dentary), UF 274077 (partial right dentary), UF 275644 (partial right dentary)"
Iguaninae indet. Cope 1886
Chovanec 2014 2 specimens
"UF 274049 (partial right dentary), UF 275635 (partial left dentary)"
Reptilia - Eublepharidae
Eublepharidae indet. Boulenger 1883
Chovanec 2014 3 specimens
"UF 275603 (partial left maxilla), UF 274247 (partial right dentary), UF 278770 (partial left dentary)"
Reptilia - Rhineuridae
Rhineuridae indet. Vanzolini 1951
Chovanec 2014 3 specimens
"UF 268989-94 (vertebrae), UF 274047 (vertebra), UF 275673 (partial right dentary)"
Reptilia - Helodermatidae
Helodermatidae indet. Gray 1837
Bourque 2013
Reptilia - Testudines - Testudinidae
Testudinidae indet. Batsch 1788
Bourque 2013
"small tortoise (from sub-localities 1A and 1B)
Reptilia - Testudines - Kinosternoidea
Xenochelys floridensis n. sp. Bourque 2013
Bourque 2013
UF 150006-150007, holotype
Reptilia
Alligatoridae indet. Gray 1844
Bourque 2013
"dwarf alligatorid"
    = Alligator sp. Daudin 1809
Whiting et al. 2016
UF 422816 and additional remains
Amphibia - Temnospondyli - Rhinophrynidae
Rhinophrynus sp. Dumeril and Bibron 1841
Blackburn et al. 2019 12 specimens
from Quarry 1A, localities HE003 and HE008
Echinoidea - Spatangoida - Schizasteridae
Agassizia (Aniaster) mossomi Cooke 1942
Cooke 1942
see common names

Geography
Country:United States State/province:Florida County:Hernando
Coordinates: 28.5° North, 82.2° West (view map)
Paleocoordinates:28.7° North, 77.0° West
Basis of coordinate:based on political unit
Geographic resolution:outcrop
Time
10 m.y. bin:Cenozoic 4-5
*Period:Tertiary *Epoch:Oligocene
*Local age/stage:Middle Arikareean
Key time interval:Arikareean
Age range of interval:29.50000 - 18.50000 m.y. ago
Age estimate:27 ± 1 Ma (other)
* legacy (obsolete) database fields
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: Middle Arikareean
The age of the deposit was considered to be 26–25 Ma by Tedford et al. (2004) based on mammalian biochronology. Later analysis by Czaplewski and Morgan (2012) and Morgan et al. (2019) suggested a slightly older age range of 28–26 Ma
Lithology and environment
Primary lithology:planar lamination claystone
Secondary lithology: sandstone
Includes fossils?Y
Includes fossils?Y
Lithology description: laminated clay and sand
Environment:fissure fill
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:general faunal/floral analysis
Metadata
Database number:17476
Authorizer:J. Alroy, M. Uhen, P. Mannion, J. Bloch, P. Holroyd, P. Barrett, B. Allen Enterer:J. Alroy, R. Hulbert, M. Uhen, T. Cleary, P. Mannion, P. Holroyd, B. Allen
Modifier:B. Allen Research group:vertebrate
Created:2002-05-15 00:00:00 Last modified:2023-07-25 09:38:26
Access level:the public Released:2002-05-15 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:
4229. F. G. Hayes. 2000. The Brooksville 2 local fauna (Arikareean, latest Oligocene) Hernando County, Florida. Bulletin of the Florida Museum of Natural History 43(1):1-47 [J. Alroy/J. Alroy/J. Alroy]

Secondary references:

71280 D.C. Blackburn, L. Roberts, M. C. Vallejo-Pareja and E. L. Stanley. 2019. First Record of the Anuran Family Rhinophrynidae from the Oligocene of Eastern North America . Journal of Herpetology 53(4):316-323 [P. Holroyd/P. Holroyd]
58392 J. R. Bourque. 2013. Fossil Kinosternidae from the Oligocene and Miocene of Florida, USA. In D. B. Brinkman, P. A. Holroyd, and J. Gardner (eds.), Morphology and Evolution of Turtles, Vertebrate Paleobiology and Paleoanthropology 459-475 [P. Holroyd/P. Holroyd]
59473 K. Chovanec. 2014. East Tennessee State University. [P. Barrett/T. Cleary]
32917 C. W. Cooke. 1942. Cenozoic irregular echinoids of the Eastern United States. Journal of Paleontology 16:1-62 [L. Villier/L. Villier]
7290 N. J. Czaplewski and G. Morgan. 2002. Phyllostomatid bats from the Oligocene and early Miocene of Florida. Journal of Vertebrate Paleontology 22(3):48A [J. Alroy/J. Alroy/J. Alroy]
84078 A. K. Hastings, B. W. Schubert, J. R. Bourque and R. C. Hulbert, Jr. 2023. Oldest record of Alligator in southeastern North America. Palaeontologia Electronica 26(1):a6:1-19 [P. Mannion/P. Mannion]
50261 J. I. Mead. 2013. Scolecophidia (Serpentes) of the late Oligocene and early Miocene, North America, and a fossil history overview. Geobios 46:225-231 [J. Bloch/R. Hulbert]
85393 G. S. Morgan, N. J. Czaplewski, and N. B. Simmons. 2019. A New Mormoopid Bat from the Oligocene (Whitneyan and Early Arikareean) of Florida, and Phylogenetic Relationships of the Major Clades of Mormoopidae (Mammalia: Chiroptera). Bulletin of the American Museum of Natural History 434:1-146 [B. Allen/B. Allen]
59566 E. T. Whiting, D. W. Steadman, and K. A. Vliet. 2016. Cranial Polymorphism and Systematics of Miocene and Living Alligator in North America. Journal of Herpetology 50(2):306-315 [P. Mannion/P. Mannion]