PBDB Collection

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Miomegaderma gaillardi	Ginsburg 1964

	Hemicyonidae indet.	Ginsburg 1964

	Schlossericyon viverroides	Ginsburg 1964
Angiospermae - Rosales - Ulmaceae

	Celtis sp. seed/fruit Linnaeus 1753	Ginsburg 1964
Amphibia - Temnospondyli - Albanerpetontidae

	Albanerpeton inexpectatum Estes and Hoffstetter 1976	Rage and Hossini 2000	33 specimens
	Sa 23401 - Sa 23403, Sa 23404a, Sa 23405, Sa 23407 - Sa 23409
Reptilia

	Anguimorpha indet. (Fürbringer 1900)	Augé and Rage 2000
	; published as Anguimorpha incertae sedis; 1 anterior left dentary fragment
Reptilia - Anguidae

	Anguinae indet. Gray 1825	Augé and Rage 2000

	Pseudopus laurillardi (Lartet 1851)	Augé and Rage 2000

	Ophisaurus sp. Daudin 1803	Augé and Rage 2000
	24 jaw elements (maxillae or dentary fragments) entered as Ophisaurus sp. 1; 5 jas elements (maxillae or dentary fragments) entered as Ophisaurus sp. 2
Reptilia - Amphisbaenidae

	Amphisbaenidae indet. Gray 1865	Augé and Rage 2000
	3 dentaries; 1 maxilla; approximately 800 vertebrae
Reptilia

	Blanus sp. Wagler 1830	Augé and Rage 2000
	23 dentaries

	Blanus antiquus	Böhme 1999
Reptilia - Boidae

	Albaneryx depereti Hoffstetter and Rage 1972	Augé and Rage 2000
Reptilia - Crotalidae

	Vipera sp. Laurenti 1768	Augé and Rage 2000
	= Viperinae informal subtaxon (Vipera sp. "oriental vipers" or Daboia sp.)	Rage and Bailon 2005
	informal subtaxon of Vipera sp. in Augé & Rage (2000) is "oriental viper"

	Vipera aegertica n. sp.	Augé and Rage 2000
	Holotype (Sa 23714) one trunk vertebra; also 20 referred vertebrae
Reptilia - Elapidae

	Micrurus gallicus Rage and Holman 1984	Augé and Rage 2000
Reptilia - Colubridae

	Paleoheterodon arcuatus Rage and Holman 1984	Augé and Rage 2000

	Colubrinae indet. Oppel 1811	Augé and Rage 2000
	includes "Colubrinae incertae sedis nomen dubium" applied to Scaptophis Rochebrune, 1880; also includes 103 indeterminate vertebrae

	Texasophis meini Rage and Holman 1984	Augé and Rage 2000

	Coluber pouchetii	Augé and Rage 2000

	Coluber (aff. Coluber) pouchetii	Augé and Rage 2000

	Elaphe sp. Wagler 1833	Augé and Rage 2000

	Palaeonatrix lehmani (Rage and Roček 1983)	Augé and Rage 2000

	Palaeonatrix (aff. Palaeonatrix) lehmani (Rage and Roček 1983)	Augé and Rage 2000

	Natricinae indet. Bonaparte 1838	Augé and Rage 2000

	Neonatrix natricoides n. sp.	Augé and Rage 2000
	Holotype MNHN Sa 23683, one mid-trunk vertebra; 79 referred vertebrae

	Neonatrix europaea Rage and Holman 1984	Augé and Rage 2000

	Natrix sansaniensis	Augé and Rage 2000

	Natrix longivertebrata Szyndlar 1984	Augé and Rage 2000

	Natrix (aff. Natrix) longivertebrata Szyndlar 1984	Augé and Rage 2000
Reptilia - Gekkonidae

	Gekkonidae indet. Gray 1825	Augé and Rage 2000
	17 maxilla or dentary fragments; 34 trunk vertebrae; 4 caudal vertebrae
Reptilia

	Lacertilia indet. (Owen 1842)	Augé and Rage 2000
	fragment of humerus
Reptilia - Lacertidae

	Amblyolacerta sp. Roček 1984	Augé and Rage 2000
	1 left dentary

	Edlartetia n. gen. sansaniensis Augé and Rage 2000	Augé and Rage 2000
	neotype designated as MNHN Sa 23595, an incomplete right dentary

	Lacerta sp. Linnaeus 1758	Augé and Rage 2000
	265 dentaries entered as Lacerta sp. 1; 54 dentaries entered as Lacerta sp. 2
Reptilia - Scincidae

	? Scincidae sp. Gray 1825	Augé and Rage 2000
	18 dentaries entered as ? Scincidae sp. 1; one dentary entered as ? Scincidae sp. 2
Reptilia - Anseriformes - Anatidae

	Anas sansaniensis Milne-Edwards 1867	Zelenkov et al. 2018
	recombined as Chenoanas sansaniensis
Reptilia - Accipitriformes - Accipitridae

	Haliaeetus piscator Milne-Edwards 1869	Mlíkovsky 1997
Reptilia - Testudines - Geoemydidae

	Clemmydopsis turnauensis (Meyer 1847)	Broin 1977

	Ptychogaster sansaniensis (Lartet 1851)	Broin 1977
	nomen dubium belonging to Chelydridae

	Ocadia sansaniensis n. sp. (Lartet 1851)	Lartet 1851
	nomen dubium belonging to Chelydridae
	MNHN SA 15648

	Ptychogaster cf. batalleri Bergounioux 1931	Broin 1977
	"Ptychogaster (Temnoclemys) cf. batalleri"
Reptilia - Testudines - Testudinidae

	Testudo canetotiana Lartet 1851	Broin 1977
	recombined as Paleotestudo canetotiana

	Testudo cf. larteti Pictet 1853	Lydekker 1889
	BMNH 40772, left precoracoid
Reptilia - Testudines - Chelydridae

	Leptochelys braneti n. gen., n. sp. Bergounioux and Crouzel 1965	Bergounioux and Crouzel 1965
	nomen dubium belonging to Chelydridae
	MNHN SA2023 (holotype), a partial hyo/hypoplastron

	Trionyx sansaniensis n. sp. Bergounioux 1935	Bergounioux 1935
	synonym of Chelydropsis murchisoni

	Broilia robusta n. gen., n. sp. Bergounioux and Crouzel 1965	Bergounioux and Crouzel 1965
	nomen dubium belonging to Chelydridae
	MNHN SA uncat. (lectotype), a string of “caudal” vertebrae
Mammalia - Proboscidea - Mammutidae

	Turicius turicensis	Ginsburg 1964
	= Zygolophodon turicensis Schintz 1824	Tassy 1977
Mammalia - Proboscidea - Deinotheriidae

	Deinotherium levius Jourdan 1868	Ginsburg 1964
	synonym of Deinotherium giganteum
	entered as "Dinotherium levius"
Mammalia - Proboscidea - Gomphotheriidae

	Trilophodon angustidens (Cuvier 1806)	Ginsburg 1964
	recombined as Tetrabelodon angustidens
Mammalia - Chiroptera

	informal Chiroptera informal I nov. gen. nov. sp. (Ginsburg 1964)	Ginsburg 1964

	informal Chiroptera informal II ("Vespertilio" noctuloides Lartet)(Ginsburg 1964)(Ginsburg 1964)	Ginsburg 1964

	informal Chiroptera informal III ("Vespertilio" murinoides Lartet) (Ginsburg 1964)	Ginsburg 1964
Mammalia - Primates - Pliopithecidae

	Pliopithecus antiquus (Blainville 1839)	Ginsburg 1964
Mammalia - Rodentia - Sciuridae

	Sciuropterus sansaniensis	Ginsburg 1964
Mammalia - Rodentia - Cricetidae

	Eumyarion medius Lartet 1851	Heißig 1989

	Cricetodon minus Lartet 1851	Ginsburg 1964
	recombined as Democricetodon minor

	Cricetodon cf. breve Schaub 1925	Ginsburg 1964
	recombined as Democricetodon brevis

	Cricetodon helveticus	Ginsburg 1964

	Cricetodon gaillardi	Ginsburg 1964

	Cricetodon cf. affine	Ginsburg 1964

	Cricetodon sansaniensis Lartet 1851	Ginsburg 1964

	Megacricetodon minor Lartet 1851	Bollinger 1994
Mammalia - Rodentia - Gliridae

	Muscardinus sansaniensis Lartet 1856	Ginsburg 1964
Mammalia - Rodentia - Castoridae

	Steneofiber sansaniensis	Ginsburg 1964
Mammalia - Lagomorpha - Ochotonidae

	Prolagus oeningensis (König 1825)	Ginsburg 1964
Mammalia - Dimylidae

	Plesiodimylus chantrei Gaillard 1899	Ginsburg 1964
Mammalia - Erinaceidae

	Lantanotherium sansaniensis (Lartet 1851)	Ginsburg 1964

	Galerix exilis de Blainville 1839	Ginsburg 1964
Mammalia - Talpidae

	Talpa minuta Blainville 1840	Ginsburg 1964

	Proscapanus sansaniensis Lartet 1851	Ginsburg 1964

	Mygalea antiqua Pomel 1848	Ginsburg 1964
Mammalia - Soricidae

	Heterosorex sansaniensis	Ginsburg 1964

	Sorex grivensis	Ginsburg 1964
Mammalia - Carnivora - Amphicyonidae

	Amphicyon major De Blainville 1841	Ginsburg 1964

	Pseudarctos bavaricus Schlosser 1899	Ginsburg 1964

	Pseudocyon sansaniensis Lartet 1851	Ginsburg 1964
Mammalia - Carnivora

	Proputorius sansaniensis Filhol 1891	Ginsburg 1964
Mammalia - Carnivora - Ursidae

	Hemicyon sansaniensis Lartet 1851	Ginsburg 1964

	Plithocyon armagnacensis Ginsburg 1955	Ginsburg 1964
Mammalia - Carnivora - Ailuridae

	Alopecocyon goeriachensis (Toula 1884)	Ginsburg 1964
	recombined as Alopecodon goeriachensis
Mammalia - Carnivora - Mustelidae

	Melidellavus leptorhynchus	Ginsburg 1964

	Mionictis dubia de Blainville 1841	Ginsburg 1964

	Mustelinae indet. Fischer 1817	Ginsburg 1964

	Lutra dubia Blainville 1841	Heizmann and Morlo 1998
	recombined as Lartetictis dubia

	Ischyrictis zibethoides Blainville 1842	Ginsburg 1964

	Martes muncki	Ginsburg 1964

	Martes sansaniensis Lartet 1851	Ginsburg 1964

	Taxodon sansaniensis Lartet 1851	Ginsburg 1964
Mammalia - Carnivora - Viverridae

	Semigenetta mutata	Ginsburg 1964
Mammalia - Carnivora - Nimravidae

	Sansanosmilus palmidens (Blainville 1841)	Ginsburg 1964
Mammalia - Carnivora - Percrocutidae

	Pseudailurus quadridentatus	Ginsburg 1964

	Pseudailurus lorteti	Ginsburg 1964

	Pseudailurus turnauensis	Ginsburg 1964
Mammalia - Bovidae

	Strogulognathus sansaniensis	Ginsburg 1964

	Eotragus sansaniensis Lartet 1851	Ginsburg 1964
Mammalia - Moschidae

	Micromeryx flourensianus Lartet 1851	Ginsburg 1964
Mammalia - Palaeomerycidae

	Palaeomeryx bojani	Ginsburg 1964
Mammalia - Lagomerycidae

	Heteroprox larteti Filhol 1890	Ginsburg 1964

	Dicrocerus elegans Lartet 1837	Ginsburg 1964
Mammalia - Tragulidae

	Dorcatherium crassum Lartet 1851	Ginsburg 1964
Mammalia

	Taucanamo sansaniense Lartet 1851	Ginsburg 1964
Mammalia - Suidae

	Listriodon splendens	Ginsburg 1964
Mammalia - Perissodactyla - Equidae

	Anchitherium aurelianense Cuvier 1825	Ginsburg 1964
Mammalia - Perissodactyla - Rhinocerotidae

	Dicerorhinus sansaniensis	Ginsburg 1964

	Brachypotherium brachypus Lartet 1848	Ginsburg 1964

	Aceratherium tetradactylum (Lartet 1837)	Ginsburg 1964
	recombined as Hoploaceratherium tetradactylum
Mammalia - Perissodactyla - Chalicotheriidae

	Chalicotherium grande (de Blainville 1849)	Ginsburg 1964
Gastropoda - Neogastropoda - Olividae

	Ancilla sp. Lamarck 1799
Gastropoda - Heterostropha - Lymnaeidae

	Lymnaea aff. stagnalis
Gastropoda - Heterostropha - Planorbidae

	Planorbis sp. Muller 1774
Gastropoda

	Bulimus lubricus (Müller 1774)
unclassified

	Cyclostoma elegans (Muller 1774)
see common names

Geography

Country:	France	State/province:	Midi-Pyrenees	County:	Gers
Coordinates:	43.5° North, 0.6° East (view map)
Paleocoordinates:	43.7° North, 0.6° West
Basis of coordinate:	based on nearby landmark
Geographic resolution:	outcrop

Time

Period:	Neogene	Epoch:	Miocene
10 m.y. bin:	Cenozoic 5
*Period:	Late/Upper Tertiary - Late/Upper Tertiary	*Epoch:	Middle Miocene - Middle Miocene
*International age/stage:	Langhian - Serravallian	*Local age/stage:	Middle Helvetian - Middle Helvetian
Key time interval:	Langhian - Serravallian
Age range of interval:	15.98000 - 11.63000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Stratigraphic resolution:

member

Lithology and environment

Primary lithology:	gray,blue unlithified claystone
Secondary lithology:	marl
Includes fossils?	Y
Includes fossils?	Y
Lithology description: The fossils come from different, superposed lithologies. Those include: purple clays, gray clays, marls, but also limestones and molasse. No distinction is made about the exact provenance of the fossils from these various levels.
Environment:	lacustrine - small

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils
Disassociated major elements:	many
Disassociated minor elements:	many

Collection methods and comments

Collection methods:	field collection
Reason for describing collection:	taxonomic analysis
Taxonomic list comments:The ref. Ginsburg (1964)is an update of previous works (Lartet 1836-1858) and Filhol (1891), but it was published as new excavations were taking place.

Metadata

Database number:	32191
Authorizer:	E. Fara, M. Uhen, C. Bell, P. Holroyd, M. Carrano, J. Alroy, J. Mueller, A. Dunhill, T. Cleary, E. Vlachos	Enterer:	E. Fara, T. Liebrecht, B. Allen, T. Cleary, E. Vlachos, S. Kuemmell, P. Holroyd, M. Carrano, M. Uhen, C. Bell
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2003-06-09 23:23:54	Last modified:	2021-07-14 11:09:55
Access level:	the public	Released:	2003-06-09 23:23:54
Creative Commons license:	CC BY

Reference information

Primary reference:

11567.

E. Lartet. 1836. [Lettre de E. Lartet a M. Michelin au sujet de la localité de Sansan (Gers), en lecture a la seance du 16 Mai 1836]. Bulletin de la Societe geologique de France 1(7):217-220 [E. Fara/E. Fara/E. Fara]

Secondary references:

43532		M. Augé and J.-C. Rage. 2000. Les squamates (Reptilia) du Miocène moyen de Sansan. Mémoires du Muséum National d'Histoire Naturelle 183:263-313 [C. Bell/C. Bell]
60872		F. M. Bergounioux. 1935. Contribution a l'étude paléontologique des chéloniens: chéloniens fossiles du bassin d'Aquitaine. Mémoires de la Société Géologique de France 25:1-214 [E. Vlachos/E. Vlachos]
64326		F. M. Bergounioux and F. Crouzel. 1965. Chéloniens de Sansan. Annales de Paléontologie 51:153-187 [E. Vlachos/E. Vlachos]
67837		M. Böhme. 1999. Doppelschleichen (Sauria, Amphisbaenia) aus dem Untermiozän von Stubersheim 3 (Süddeutschland). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historischen Geologie 39:85-90 [T. Cleary/T. Cleary]
11712	5% 6560	T.h. Bollinger. 1994. Die Obere Süßwassermolasse in Bayern und der Ostschweiz: bio- und lithostratigraphische Korrelationen (Upper Freshwater molasse in Bavaria and Eastern Switzerland: bio- and lithostratigraphic correlations). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 34:109-144 [J. Alroy/S. Kuemmell/S. Kuemmell]
60047		F. Broin. 1977. Contribution a l'etude des Cheloniens. Cheloniens continentaux due Cretace et du Tertiare de France. Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie, Serie C, Sciences de la terre 38:1-366 [P. Holroyd/P. Holroyd/P. Holroyd]
52039		J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano]
77386		J. D. Gardner, A. Villa, S. Colombero, M. Venczel, and M. Delfino. 2021. A Messinian (latest Miocene) occurrence for Albanerpeton Estes & Hoffstetter, 1976 (Lissamphibia: Albanerpetontidae) at Moncucco Torinese, Piedmont Basin, northwestern Italy, and a review of the European Cenozoic record for albanerpetontids. Geodiversitas 43(14):391-404 [M. Carrano/M. Carrano]
28567		P. Gervais. 1852. Zoologie et Paléontologie Françaises (Animaux Vertébrés) ou Nouvelles Recherches sur les Animaux Vivants et Fossiles de la France [French Zoology and Paleontology (Vertebrate Animals) or New Research on the Living and Fossil Animals of France] 1–3:1-274+ [M. Uhen/M. Uhen/M. Carrano]
8672	5% 41560	L. Ginsburg. 1964. Les mammiferes fossiles recoltes a Sansan au cours du XIXe siecle. Bulletin de la Societe geologique de France 5(1):3-15 [E. Fara/E. Fara/J. Alroy]
13262		K. Heißig. 1989. The faunal succession of the Bavarian Molasse reconsidered - correlation of the MN5 and MN6 faunas. Lindsay, E. H., Fahlbusch, V. & Mein, P.: European Neogene Mammal Chronology. Nato ASI-series, Plenum Press; New York 181-192 [J. Alroy/S. Kuemmell/S. Kuemmell]
24579		E. J. Heizmann and M. Morlo. 1998. Die semisquatische Lartetictis dubia (Mustelinae, Carnivora, Mammala) vom Goldberg/Ries (Baden-Württemberg). Mainzer Naturwissenschaftliches Archiv Beiheft 21:141-153 [M. Uhen/M. Uhen/M. Uhen]
64331		E. Lartet. 1851. Notice sur la colline de Sansan, suivie d’une récapitulation des diverses espèces d’animaux vertébrés fossiles, trouvés soit a Sansan, soit dans d’autres gisements du terrain Tertiaire Miocène dans le Bassin sous-Pyrénéen. 1-45 [E. Vlachos/E. Vlachos]
41531		R. Lydekker. 1889. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History) Part III. The Order Chelonia 1-239 [R. Benson/R. Benson/M. Uhen]
65507		J. Mlíkovsky. 1997. Taxonomic identity of Haliaeetus angustipes Jánossy, 1983 (Aves: Accipitridae) from the early Pleistocene of the Czech Republic. Buteo 9:51-56 [M. Uhen/M. Uhen]
47304		J. C. Rage and S. Hossini. 2000. Les Amphibians du Miocene moyen de Sansan. Memoires du Museum national d'Histoire naturelle 183:177-217 [M. Carrano/M. Carrano]
43777		J.-C. Rage and S. Bailon. 2005. Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France). Geodiversitas 27(3):413-441 [J. Mueller/T. Liebrecht]
73265		N. V. Zelenkov, T. A. Stidham, N. Martynovich, N. V. Volkova, Q. Li and Z. Qui. 2018. The middle Miocene duck, Chenoanas (Aves, Anatidae): New species, phylogeny and geographical range. Papers in Paleontology 4(3):309-326 [M. Uhen/M. Uhen]