PPP# 170 - Escudo de Veraguas [Escudo de Veraguas Fm]: Piacenzian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Turbonillidae indet.
3 specimens
? Turbonillidae indet.
2 specimens
originally entered as "? Turbonillidae sp."
Anthozoa - Scleractinia - Caryophylliidae
Asterosmilia profunda (Duncan 1864)
Scaphopoda - Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758
9 specimens
sensu-lato
Scaphopoda - Gadilida - Gadilidae
Gadilopsis sp. (Woodring 1925)
9 specimens
Scaphopoda - Gadilida - Entalinidae
Bathoxiphus sp. Pilsbry 1897
2 specimens
Bivalvia - Arcida - Arcidae
Anadara (Rasia) sp. (Gray 1857)
4 specimens
Bivalvia - Nuculanida - Nuculanidae
Saccella sp. Woodring 1925
4 specimens
Bivalvia - Nuculida - Nuculidae
Nucula (Lamellinucula) sp. (Schenck 1944)
1 specimen
Gastropoda - Trochidae
Microgaza sp. Dall 1881
5 specimens
Gastropoda - Opisthobranchia - Cylichnidae
Acteocina sp. Gray 1847
1 specimen
Gastropoda - Opisthobranchia - Retusidae
Volvulella sp. Newton 1891
6 specimens
Gastropoda - Opisthobranchia - Cavoliniidae
Styliola sp. Gray 1850
3 specimens
Cavolinia sp. Avilgaard 1791
1 specimen
Gastropoda - Heterostropha - Architectonicidae
Architectonica sp. Röding 1798
9 specimens
Pseudotorinia sp. Sacco 1892
1 specimen
Gastropoda - Heterostropha - Pyramidellidae
Pyramidellidae indet. Gray 1840
2 specimens
Eulimella (Ebalina) sp. (Thiele 1929)
11 specimens
Gastropoda - Heterostropha - Acteonidae
Acteon sp. Montfort 1810
10 specimens
Gastropoda - Cocculinidae
? Cocculina sp. Dall 1882
3 specimens
Gastropoda - Eulimidae
Eulimidae indet. Philippi 1853
1 specimen
Gastropoda - Naticidae
Natica (Naticarius) sp. (Dumeril 1806)
5 specimens
original and current combination Naticarius
Gastropoda - Rissoidae
Rissoidae indet. Gray 1847
7 specimens
Gastropoda - Neogastropoda - Columbellidae
Cotonopsis sp. Olsson 1942
2 specimens
Gastropoda - Neogastropoda - Buccinidae
Metula sp. Adams and Adams 1853
3 specimens
Antillophos (Antillophos) sp. Woodring 1928
6 specimens
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
1 specimen
sensu-lato
Gastropoda - Neogastropoda - Olividae
Olivella (Minioliva) sp. Olsson 1956
3 specimens
Olivella (Macgintiella) sp. Olsson 1956
10 specimens
Oliva (Oliva) sp. Bruguière 1789
2 specimens
Gastropoda - Neogastropoda - Turridae
Polystira sp. Woodring 1928
8 specimens
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
16 specimens
Gastropoda - Neogastropoda - Drilliidae
Agladrillia (Agladrillia) sp. Woodring 1928
4 specimens
Gastropoda - Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891
3 specimens
Gastropoda - Neogastropoda - Mangeliidae
Ithycythara sp. Woodring 1928
1 specimen
Gastropoda - Neogastropoda - Marginellidae
Marginellidae indet. Fleming 1828
6 specimens
Volvarina sp. (Hinds 1844)
20 specimens
Gastropoda - Neogastropoda - Muricidae
Haustellum sp. Schumacher 1817
6 specimens
Gastropoda - Personidae
Distorsio clathrata (Lamarck 1816)
Beu 2010 1 specimen
Distorsio decussata (Valenciennes 1832)
Beu 2010 1 specimen
Gastropoda - Cassidae
Sconsia grayi Adams 1855
4 specimens
Gastropoda - Bursidae
Marsupina bufo (Bruguiére 1792)
Beu 2010 1 specimen
Gastropoda - Tonnidae
Malea ringens (Swainson 1822)
Beu 2010 1 specimen
Gastropoda - Pseudomelaniidae
Miraclathurella sp. Woodring 1928
1 specimen
? Compsodrillia sp. Woodring 1928
2 specimens
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.5° West (view map)
Paleocoordinates:9.8° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian Nannofossil zone: NN15
Age range of interval:3.60000 - 2.58000 m.y. ago
Age estimate:3.6 Ma (age-depth)3.6 to 3.5 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), planktic foraminifera suggest 1.8 Ma near the top and 3.5 Ma at the base. PPP dataset indicates an age range between 3.6 and 3.5 Ma, nannofossil zone NN15.
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, prevasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:BMNH,NMB,USNM
Collection method comments: COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 8/7/87. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),
Taxonomic list comments:NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 17622; 87-09-03
Database number:61337
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2006-06-05 20:34:43 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2006-12-05 20:34:43
Creative Commons license:CC BY
Reference information

Primary reference:
17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

30037 A. G. Beu. 2010. Neogene Tonnoidean Gastropods of Tropical and South America; contributions to the Dominican Republic and Panama Paleontology Projects and Uplift of the Central American Isthmus. Bulletins of American Paleontology (377-378)1-550 [A. Hendy/A. Hendy/A. Hendy]