Akespe: Late/Upper Oligocene, Kazakhstan
collected by N.S. Shevyreva 1968

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Urodela
Mioproteus sp. Estes and Darevsky 1978
Vasilyan et al. 2017
GNM, GIN
Mammalia - Perissodactyla - Eggysodontidae
Eggysodon sp. Roman 1910
Lucas et al. 1998
Mammalia - Perissodactyla - Paraceratheriidae
Paraceratherium sp. Forster Cooper 1911
Lucas et al. 1998
Paraceratherium prohorovi
Lucas et al. 1998
Mammalia - Perissodactyla - Rhinocerotidae
Protaceratherium sp. (Abel 1910)
Lucas et al. 1998
Aprotodon borissiaki
Lucas et al. 1998
Gigantamynodon akespensis n. sp. Bayshashov 1993
Lucas and Emry 1996
nomen dubium belonging to Rhinocerotidae
KAN PK 11/6 - type
Aceratherium aralense
Lucas et al. 1998
Mammalia - Archaeomerycidae
Lucas et al. 1998
Mammalia
Prodremotherium sp. Filhol 1877
Lucas et al. 1998
Mammalia - Lophiomerycidae
Lophiomeryx sp. Pomel 1853
Lucas et al. 1998
Mammalia - Moschidae
Micromeryx sp. Lartet 1851
Lucas et al. 1998
Mammalia - Palaeomerycidae
Amphitragulus sp. Pomel 1847
Lucas et al. 1998
Mammalia - Erinaceidae
Amphechinus sp. Aymard 1850
Lopatin 2004
PIN, no. 210/265 and 737
Amphechinus akespensis n. sp. Lopatin 1999
Lopatin 1999
PIN, no. 210/735 - type; referred material: PIN, no. 210/736 and 210/774
Amphechinus minimus Bohlin 1942
Lucas et al. 1998
Palaeoscaptor acridens Matthew and Granger 1924
Lucas et al. 1998
Mammalia - Lagomorpha - Ochotonidae
Sinolagomys pachygnathus Li and Qiu 1980
Lopatin 1998
PIN, no. 210/773
Mammalia - Lagomorpha - Leporidae
Desmatolagus aff. schargaltensis
Lucas et al. 1998
Agispelagus simplex n. gen., n. sp. Argyropulo 1940
Lopatin 1998
recombined as Desmatolagus simplex
PIN, no. 210/609 - type
Mammalia - Rodentia - Cricetidae
Eucricetodon aff. caducus (Shevyreva 1967)
Lucas et al. 1998
original and current combination Cricetops caducus
Eucricetodon aff. asiaticus (Matthew and Granger 1923)
Lucas et al. 1998
Argyromys woodi n. sp.
Lucas et al. 1998
PIN no. 210/262 (type), 765 and 766
Argyromys aralensis n. sp. Argyropulo 1939
Lucas et al. 1998
PIN, no. 210/261 - type
Protalactaga borissiaki n. sp. Argyropulo 1939
Lucas et al. 1998
synonym of Argyromys aralensis
PIN no. 210/260 - type
Eumyarion tremulus Lopatin 1996
Lopatin 2004
PIN, no. 210/764
Mammalia - Rodentia - Muridae
Eumysodon spurius n. gen., n. sp. Argyropulo 1939
Lucas et al. 1998
PIN, no. 210/264 - type
Eumysodon orlovi n. sp. Argyropulio 1939
Lucas et al. 1998
synonym of Eumysodon spurius
PIN no. 210/264a - type
Aralomys gigas n. sp.
Lucas 1996
synonym of Tachyoryctoides
PIN, no. 210/263 - type; referred specimen: PIN, no. 210/772
Tachyoryctoides obrutschewi Bohlin 1937
Lucas et al. 1998
Mammalia - Rodentia - Castoridae
Capatanka schokensis Bendukidze 1993
Lucas et al. 1998
recombined as Propalaeocastor schokensis
Steneofiber kumbulakensis (Lytschev 1970)
Lucas et al. 1998
original and current combination Propalaeocastor kumbulakensis
PIN no. 210/767-769
Steneofiber sp. Geoffroy 1833
Lucas et al. 1998
Agnotocastor sp. Stirton 1935
Lucas et al. 1998
see common names

Geography
Country:Kazakhstan
Coordinates: 46.8° North, 60.6° East (view map)
Paleocoordinates:47.6° North, 58.1° East
Basis of coordinate:estimated from map
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Oligocene
Stage:Chattian 10 m.y. bin:Cenozoic 4
Key time interval:Late/Upper Oligocene
Age range of interval:27.82000 - 23.03000 m.y. ago
Stratigraphy
Formation:Aral
Stratigraphic resolution:bed
Stratigraphy comments: late Oligocene
Lithology and environment
Primary lithology:gray,green,yellow calcareous "shale"
Secondary lithology: claystone
Includes fossils?Y
Lithology description: grayish yellow-green calcareous shale and claystone
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Reason for describing collection:taxonomic analysis
Collectors:N.S. Shevyreva Collection dates:1968
Metadata
Also known as:Agyspe; Kumbulak Cliffs
Database number:64413
Authorizer:M. Uhen, E. Vlachos, P. Mannion Enterer:M. Uhen, E. Vlachos, P. Mannion
Modifier:P. Mannion Research group:vertebrate
Created:2006-09-08 07:15:12 Last modified:2016-01-20 10:37:04
Access level:the public Released:2006-09-08 07:15:12
Creative Commons license:CC BY
Reference information

Primary reference:
57669. A. V. Lopatin. 1996. The Stratigraphy and Small Mammals from the Aral Formation, the Altynshokysu Locality, Northern Aral Area. Stratigraphy and Geological Correlation 4(2):65-79 [P. Mannion/P. Mannion]

Secondary references:

57695 A. V. Lopatin. 1998. A Revision of the Early Miocene Lagomorpha (Mammalia) from the North Aral Region. Paleontological Journal 32(3):291-304 [P. Mannion/P. Mannion]
24211 A. V. Lopatin. 1999. Oligocene and Early Miocene Insectivores (Mammalia) from Western Kazakhstan. Paleontological Journal 33(2):182-191 [L. van den Hoek Ostende/L. van den Hoek Ostende/P. Mannion]
57712 A. V. Lopatin. 2004. Early Miocene small mammals from the North Aral region (Kazakhstan) with special reference to their biostratigraphic significance. Paleontological Journal 38(S3):S217-S323 [P. Mannion/P. Mannion]
57112 S. G. Lucas. 1996. Fossil mammals and the age of the Changxindian Formation, Northeastern China. Palaeo Vertebrata 25(2-4):133-140 [P. Mannion/P. Mannion/P. Mannion]
58493 S. G. Lucas and R. J. Emry. 1996. Biochronological significance of Amynodontidae (Mammalia, Perissodactyla) from the Paleogene of Kazakhstan. Journal of Paleontology 70(4):691-696 [P. Mannion/P. Mannion]
18566 S. G. Lucas, E. G. Kordikova, and R. J. Emry. 1998. Oligocene stratigraphy, sequence stratigraphy, and mammalian biochronology north of the Aral Sea, Western Kazakstan. Bulletin of the Carnegie Museum of Natural History 34:313-348 [M. Uhen/M. Uhen]
62113 D. Vasilyan, V. S. Zazhigin, and M. Bohme. 2017. Neogene amphibians and reptiles (Caudata, Anura, Gekkota, Lacertilia, and Testudines) from the south of Western Siberia, Russia, and Northeastern Kazakhstan. PeerJ [E. Vlachos/E. Vlachos]