Le Quesnoy (QNY2): Ypresian, France
collected 1997 1998

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Osteichthyes - Lepisosteiformes - Lepisosteidae
    = Lepisosteidae indet. Cuvier 1825
Dutheil et al. 2006
Osteichthyes - Amiiformes - Amiidae
Osteichthyes - Palaeolabridae
Palaeolabrus sp. Casier 1967
Dutheil et al. 2006
Osteichthyes - Perciformes
Percoidei indet. (Bleeker 1859)
    = Diaphyodus sp. Schafhäutl 1863
Dutheil et al. 2006
Osteichthyes - Scombriformes - Trichiuridae
Eutrichiurides sp. Casier 1944
Dutheil et al. 2006
Osteichthyes - Albuliformes - Albulidae
Albulidae indet. Bleeker 1859
Dutheil et al. 2006
Osteichthyes - Elopiformes - Phyllodontidae
Egertonia sp. Cocchi 1866
Dutheil et al. 2006
unclassified
Osteichthyes - Testudines
'chelydroide' gen, sp. indet.
Osteichthyes - Testudines - Geoemydidae
Palaeochelys sp. sensu lato Meyer 1847
    = Merovemys ploegi n. gen., n. sp. Hervet 2006
Hervet 2006
Osteichthyes - Testudines - Trionychidae
Trionyx s.l. sp. 1 and sp. 2
    = Trionychidae indet. Gray 1825
Alroy 2009
erroneously spelled Palaeotrionyx
Osteichthyes - Eolacertidae
Eolacerta sp. Nöth 1940
Augé 2005 3 specimens
incomplete right maxilla, two dorsal vertebrae
Osteichthyes - Lacertidae
Lacertidae indet. Bonaparte 1831
Plesiolacerta sp. Hoffstetter 1942
Osteichthyes - Tropidophiidae
Osteichthyes - Boidae
Boidae indet. Gray 1825
Groupe Paleryx-Palaeopythori; gen., sp. indet
Osteichthyes - Cordylidae
Osteichthyes - Necrosauridae
Osteichthyes
Osteichthyes - Anguidae
Osteichthyes - Gekkonidae
Osteichthyes - Iguanidae
Osteichthyes
Oiseaux Fam. indet.: gen., sp. indet.
MNHN QNY 1C7
Diplocynodon sp. Pomel 1847
MNHN QNY 1C4
MNHN QNY 2C1
MNHN QNY 1C5 & QNY 2C4
Osteichthyes - Multituberculata
Osteichthyes
Peratherium sp. Aymard 1850
or Amphiperatherium sp.
Osteichthyes - Macroscelidea - Amphilemuridae
cf. Macrocranion sp. Weitzel 1949
Osteichthyes - Primates - Plesiadapidae
Platychoerops sp. Charlesworth 1855
new species
Osteichthyes - Primates - Notharctidae
Cantius sp. Simons 1962
Osteichthyes - Primates - Omomyidae
Teilhardina aff. belgica (Teilhard de Chardin 1927)
Osteichthyes - Rodentia - Ischyromyidae
gen., sp. indet. 1 and 2
Osteichthyes - Pantolestidae
Palaeosinopa sp. Matthew 1901
Osteichthyes - Nyctitheriidae
Osteichthyes - Perissodactyla - Palaeotheriidae
Osteichthyes - Perissodactyla - Equidae
Bronnert et al. 2017
MNHN QNY2-2696 - type
Osteichthyes - Diacodexeidae
Osteichthyes - Procreodi - Arctocyonidae
cf. Landenodon sp. Quinet 1968
Osteichthyes
Solé et al. 2013
MNHN QNY2-2548 - type; referred specimens: MNHN QNY2-2547, 2505, 2532, 2535, 2502 and 2507
    = Miacis latouri Quinet 1966
Solé et al. 2013
MNHN QNY2- 2619, 2534, 2553, 2554, 2523, 2624 and 2511
Solé et al. 2013
MNHN QNY2- 2520, 2517, 2546, 2510, 2618, 2522, 2528 and 2533
Osteichthyes - Hyaenodonta - Hyaenodontidae
Solé et al. 2013
MNHN QNY2- 2504, 2508, 2514, 2570, 2617, 2623 and 2625
Solé et al. 2013
MNHN QNY2-2501, 2512, 2516 and 2519
gen., sp. indet. 1 and 2
Solé et al. 2013
MNHN QNY2-2509, 2529-2531 and 2542
Osteichthyes - Oxyaenidae
Oxyaena woutersi (Lange-Badré and Godinot 1982)
Solé et al. 2011
MNHN QNY2-2568, 2527, 2569, 2561, 2603, 2584, 2586, 2598, 2551, 2571, 2574, 2590, 2572, 2573, 2600, 2503, 2549, 2622, 2565, 2595, 2579, 2524, 2526, 2636, 2633, 2635 and 2638
Palaeonictis cf. gigantea de Blainville 1842
    = Palaeonictis gigantea de Blainville 1842
Solé et al. 2011
MNHN QNY2-2562 (and many more specimens)
Osteichthyes - Metacheiromyidae
Gheerbrant et al. 2005
Osteichthyes - Condylarthra - Hyopsodontidae
Hyopsodus sp. Leidy 1870
Osteichthyes - Apatotheria
cf. Apatotheria indet. Scott and Jepsen 1936
Osteichthyes - Esthonychidae
Esthonyx sp. Cope 1874
Osteichthyes - Pantodonta
Osteichthyes - Temnospondyli - Pelobatidae
Pelobatidae indet. Lataste 1879
Osteichthyes - Temnospondyli
Caudata indet. Scopoli 1777
fam., gen., sp. indet.
Osteichthyes - Temnospondyli - Urodela
gen., sp. indet.
unclassified
"dicotyledon" wood
Angiospermae - Fagales - Juglandaceae
pollen
Angiospermae - Fagales - Myricaceae
Myricaceae indet. Martinov 1817
pollen: Myricaceae and/or Symplocaceae
Angiospermae - Celastrales - Celastraceae
pollen
Angiospermae - Rosales
fruit
Angiospermae - Gentianales - Apocynaceae
Apocynaceae indet. de Jussieu 1789
pollen; Apocynaceae and/or Tiliaceae
Angiospermae - Arecaceae
Arecaceae indet. von Berchtold and Presl 1820
wood
Angiospermae - Laurales - Lauraceae
Lauraceae indet. Jussieu 1789
leaf
unclassified
wood
Charales - Feistiellaceae
Nitellopsis (Tectochara)
see common names

Geography
Country:France State/province:Oise
Coordinates: 49.3° North, 2.7° East (view map)
Paleocoordinates:40.4° North, 9.6° East (Wright 2013)
Basis of coordinate:based on nearby landmark
Time
Period: Paleogene Epoch: Eocene
Stage: Ypresian 10 m.y. bin: Cenozoic 2
Key time interval: Ypresian
Age range of interval: 56 - 48.07 m.y. ago
Stratigraphy
Formation:Argiles d'lignite du Soissonnais
Stratigraphic resolution:group of beds
Stratigraphy comments: MP7
Lithology and environment
Primary lithology:lenticular,gray sandstone
Includes fossils?Y
Lithology description: a succession of lenticular bodies showing two main facies: 1) clayey sands rich in frequently pyritised lignite, together with amber; 2) grey clayey sands with less lignite (1 to 12 % of the sediment), with continental vertebrate fauna. These facies, the rarity of mollusc shells and charophytes, probably due to decarbonatation and the presence of pyrrhotite (FeS4), reflect a hypoxic environment.
Environment:fine channel fill
Geology comments: Strata deposited at bottom of two channels cutting into underlying Thanetian marine green sands. The channels prograde toward the northeast and were discovered under Oise River Quaternary deposits.
Taphonomy
Modes of preservation:body,soft parts
Size of fossils:macrofossils,microfossils
Collection methods and comments
Collection excludes:some genera,some macrofossils,some microfossils
Collection methods:bulk,field collection
Reason for describing collection:general faunal/floral analysis
Museum repositories:MNHN
Collection dates:1997 and 1998
Collection method comments: Screen-washing noted by Gheerbrant et al. 2005
Taxonomic list comments:Original faunal list only notes vertebrates; other organisms mentioned in text. Amber specimens in a different collection.
Metadata
Database number:68174
Authorizer:P. Holroyd, P. Mannion, M. Clapham, P. Barrett, J. Alroy Enterer:P. Holroyd, T. Cleary, J. Alroy, M. Clapham, P. Mannion
Modifier:M. Clapham Research group:paleobotany,paleoentomology,vertebrate
Created:2007-01-06 17:04:04 Last modified:2026-02-03 17:33:58
Access level:the public Released:2007-01-06 17:04:04
Creative Commons license:CC0
Reference information

Primary reference:
19606. A. Nel, G. de Plöeg, J. Dejax, D. Duthiel, D. Franceschi, E. Gheerbrant, M. Godinot, S. Hervet, J.-J. Menier, M. Augé, G. Bignot, C. Cavagnetto, S. Duffaud, J. Gaudant, S. Hua, A. Jossang, F. Lapparent Broin, J.-P. Pozzi, J.-C. Paicheler, F. Beuchet, and J.-C. Rage. 1999. Un gisement sparnacien exceptionnel à plantes, arthropodes et vertébrés (Éocène basal, MP7): Le Quesnoy (Oise, France). Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science 329(1):65-72 [P. Holroyd/P. Holroyd]

Secondary references:

29343 J. Alroy. 2009. High-handed opinions on taxonomy and other matters. [J. Alroy/J. Alroy]
59290 M. L. Augé. 2005. Evolution des lézards du Paléogène en Europe. Mémoires du Muséum national d'histoire naturelle 192:1-369 [P. Barrett/T. Cleary/T. Cleary]
62812 C. Bronnert, E. Gheerbrant, M. Godinot and G. Métais. 2017. A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (MP7, France). Historical Biology [P. Mannion/P. Mannion]
94118 D. B. Dutheil, F. Moreau, and G. De Plöeg. 2006. Les ichthyofaunes du gisement à ambre de Le Quesnoy (Paléocène et Éocène du bassin de Paris, France). Cossmanniana 11:1-13 [M. Clapham/M. Clapham]
19607 E. Gheerbrant, K. D. Rose, and M. Godinot. 2005. First palaeanodont (?pholidotan) mammal from the Eocene of Europe. Acta Palaeontologica Polonica 50(2):209-218 [P. Holroyd/P. Holroyd]
58649 S. Hervet. 2006. The oldest European ptychogasterid turtle (Testudinoidea) from the lowermost Eocene amber locality of Le Quesnoy (France, Ypresian, MP7). Journal of Vertebrate Paleontology 26(4):839-848 [P. Holroyd/P. Holroyd]
46507 S. Hua. 2004. Les crocodiliens du Sparnacien (Eocene Inferieur) du Quesnoy (Oise, France). Oryctos 5:57-62 [P. Mannion/P. Mannion/P. Mannion]
60841 F. Solé, E. Gheerbrant, and M. Godinot. 2011. New data on the Oxyaenidae from the Early Eocene of Europe; biostratigraphic, paleobiogeographic and paleoecologic implications. Palaeontologia Electronica 14(2):13A:1-41 [P. Mannion/P. Mannion]
60847 F. Solé, E. Gheerbrant, and M. Godinot. 2013. The “miacids” (Carnivoraformes, Mammalia) from the Early Eocene locality of Le Quesnoy (MP7, France); first occurrence of Vassacyon in Europe. Comptes Rendus Palevol 12:191-202 [P. Mannion/P. Mannion]
60840 F. Solé, E. Gheerbrant, and M. Godinot. 2013. Sinopaninae and Arfianinae (Hyaenodontida, Mammalia) from the Early Eocene of Europe and Asia; evidence for dispersal in Laurasia around the Paleocene/Eocene boundary and for an unnoticed faunal turnover in Europe. Geobios 46:313-327 [P. Mannion/P. Mannion]