PPP# 359 - Escudo de Veraguas [Escudo de Veraguas Fm]: Piacenzian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
? Rissoine sp.
4 specimens
Anthozoa - Scleractinia - Caryophylliidae
Asterosmilia profunda (Duncan 1864)
Scaphopoda - Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758
22 specimens
sensu lato
Scaphopoda - Gadilida - Gadilidae
Cadulus sp. Philippi 1844
3 specimens
Bivalvia - Arcida - Arcidae
? Hawaiarca sp. Dall et al. 1938
1 specimen
spelled with current rank as Barbatia (Hawaiarca)
Anadara (Rasia) sp. (Gray 1857)
4 specimens
Bivalvia - Pectinida - Propeamussiidae
Cyclopecten sp. Verrill 1897
1 specimen
Bivalvia - Carditida - Crassatellidae
Crassinella sp. Guppy 1874
1 specimen
Bivalvia - Lucinida - Lucinidae
Parvilucina sp. (Dall 1901)
5 specimens
sensu lato
Bivalvia - Pholadida - Corbulidae
Caryocorbula sp. (Gardner 1926)
1 specimen
original and current combination Corbula (Caryocorbula)
Bivalvia - Cardiida - Semelidae
Abra sp. Lamarck 1818
8 specimens
Bivalvia - Cardiida - Tellinidae
Angulus sp. Megerle 1811
2 specimens
Tellina (Phyllodina) sp. Dall 1900
1 specimen
Bivalvia - Poromyida - Verticordiidae
Trigonulina sp. d'Orbigny 1946
1 specimen
Bivalvia - Poromyida - Poromyidae
Poromya (Poromya) sp. Forbes 1844
1 specimen
Bivalvia - Poromyida - Cuspidariidae
Halonympha sp. Dall and Smith 1886
1 specimen
Bivalvia - Nuculanida - Nuculanidae
Jupiteria sp. Bellardi 1875
3 specimens
Saccella sp. Woodring 1925
3 specimens
Bivalvia - Nuculanida - Yoldiidae
Yoldia sp. Möller 1842
14 specimens
Gastropoda - Trochidae
Microgaza sp. Dall 1881
1 specimen
Gastropoda - Opisthobranchia - Cylichnidae
Cylichnella sp. Gabb 1873
6 specimens
Acteocina sp. Gray 1847
6 specimens
Gastropoda - Opisthobranchia - Retusidae
Volvulella (Volvulella) sp. Newton 1891
2 specimens
Sulcoretusa sp. Burch 1945
12 specimens
Gastropoda - Opisthobranchia - Haminoeidae
Atys (Aliculastrum) sp. (Pilsbry 1896)
1 specimen
original and current combination Aliculastrum
Gastropoda - Opisthobranchia - Limacinidae
Limacina sp. Bosc 1817
2 specimens
? Limacina sp. Bosc 1817
1 specimen
Gastropoda - Opisthobranchia - Cavoliniidae
Styliola sp. Gray 1850
13 specimens
Creseis sp. Rang 1828
11 specimens
Clio sp. Linnaeus 1767
8 specimens
Cavolinia sp. Avilgaard 1791
19 specimens
Gastropoda - Heterostropha - Murchisonellidae
Anisocycla (Saccoina) sp. (Dall and Bartsch 1904)
2 specimens
Gastropoda - Heterostropha - Pyramidellidae
Besla sp. Dall and Bartsch 1904
2 specimens
Eulimella (Eulimella) sp. Forbes and MacAndrew 1846
2 specimens
? Eulimella (Eulimella) sp. Forbes and MacAndrew 1846
1 specimen
Eulimella (Ebalina) sp. (Thiele 1929)
2 specimens
Pyramidellinae indet. Gray 1840
1 specimen
Gastropoda - Heterostropha - Ringiculidae
Ringicula (Ringiculella) sp. (Sacco 1892)
4 specimens
Gastropoda - Eulimidae
Eulimidae indet. Philippi 1853
5 specimens
Gastropoda - Naticidae
Natica sp. Scopoli 1777
1 specimen
Polinices sp. Monfort 1810
2 specimens
Gastropoda - Rissoidae
Rissoidae indet. Gray 1847
7 specimens
Alvania sp. Risso 1826
2 specimens
Gastropoda - Tornidae
Teinostoma sp. Adams and Adams 1853
3 specimens
Gastropoda - Neogastropoda - Columbellidae
Costoanachis sp. Sacco 1890
1 specimen
spelled with current rank as Anachis (Costoanachis)
Mitrella sp. Risso 1826
1 specimen
Gastropoda - Neogastropoda - Buccinidae
Solenosteira sp. Dall 1890
3 specimens
Antillophos (Antillophos) sp. Woodring 1928
12 specimens
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
17 specimens
sensu lato
Gastropoda - Neogastropoda - Olividae
Olivella (Macgintiella) sp. Olsson 1956
20 specimens
Gastropoda - Neogastropoda - Turridae
Vaughanites sp. Woodring 1928
1 specimen
Polystira sp. Woodring 1928
6 specimens
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
4 specimens
Gastropoda - Neogastropoda - Drilliidae
Agladrillia (Agladrillia) sp. Woodring 1928
1 specimen
Gastropoda - Neogastropoda - Raphitomidae
Daphnella sp. Hinds 1844
1 specimen
Gastropoda - Neogastropoda - Clathurellidae
Nannodiella sp. (Dall 1919)
1 specimen
Gastropoda - Neogastropoda - Borsoniidae
? Microdrillia sp. Casey 1903
1 specimen
Gastropoda - Neogastropoda - Cystiscidae
? Persicula sp. Schumacher 1817
22 specimens
Gastropoda - Neogastropoda - Marginellidae
Volvarina sp. (Hinds 1844)
8 specimens
Dentimargo sp. Cossmann 1899
17 specimens
Gastropoda - Neogastropoda - Muricidae
Haustellum sp. Schumacher 1817
1 specimen
Gastropoda - Strombidae
Strombus sp. Linnaeus 1758
1 specimen
Gastropoda - Cassidae
Sconsia sp. Gray 1847
1 specimen
Gastropoda - Atlantidae
Atlanta sp. Lesueur 1817
2 specimens
Gastropoda - Triphoridae
Triphoridae indet. Gray 1847
1 specimen
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.6° West (view map)
Paleocoordinates:9.0° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian
Age range of interval:3.60000 - 2.58000 m.y. ago
Age estimate:2.9 Ma (other)3.6 to 2.1 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999).
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:BMNH,NMB,USNM
Collection method comments: COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 6/4/88. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),
Taxonomic list comments:NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 17833
Database number:79988
Authorizer:A. Hendy Enterer:K. Bulinski, A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2008-03-31 18:55:27 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2009-03-31 18:55:27
Creative Commons license:CC BY
Reference information

Primary reference:
17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]