PPP# 361 - Escudo de Veraguas [Escudo de Veraguas Fm]: Piacenzian, Panama
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gymnolaemata
- Cheilostomata
- Membraniporidae
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Cupuladria aff. canariensis
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11 specimens | |||||||||
Cupuladria aff. biporosa
Canu and Bassler 1923
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10 specimens | |||||||||
Gymnolaemata
- Cheilostomata
- Lepraliellidae
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Celleporaria albirostris
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1 specimen | |||||||||
Gymnolaemata
- Cheilostomata
- Gemelliporellidae
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Gemelliporella glabra
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10 specimens | |||||||||
Gymnolaemata
- Cheilostomata
- Petraliidae
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Petraliella bisinuata
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1 specimen | |||||||||
Gymnolaemata
- Cheilostomata
- Smittinidae
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Parasmittina sp.
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1 specimen | |||||||||
Gymnolaemata
- Cheilostomata
- Phidoloporidae
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Schedocleidochasma cleidostoma
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1 specimen | |||||||||
Gymnolaemata
- Cheilostomata
- Mamilloporidae
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Mamillopora informal n. sp. 1
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1 specimen | |||||||||
Bivalvia
- Nuculanida
- Nuculanidae
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Saccella sp.
Woodring 1925
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2 specimens | |||||||||
Bivalvia
- Nuculida
- Nuculidae
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Nucula (Lamellinucula) sp.
(Schenck 1944)
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12 specimens | |||||||||
Bivalvia
- Mytilida
- Crenellidae
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Crenella sp.
Brown 1827
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16 specimens | |||||||||
Bivalvia
- Ostreida
- Gryphaeidae
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? Hyotissa sp.
Stenzel 1971
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2 specimens | |||||||||
Bivalvia
- Pectinida
- Pectinidae
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? Spathochlamys sp.
Waller 1993
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7 specimens | |||||||||
Spathochlamys sp.
Waller 1993
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1 specimen | |||||||||
Argopecten sp.
Monterosato 1889
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13 specimens | |||||||||
Bivalvia
- Pectinida
- Propeamussiidae
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Cyclopecten sp.
Verrill 1897
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3 specimens | |||||||||
Bivalvia
- Arcida
- Limopsidae
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Limopsis sp.
Sassi 1827
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5 specimens | |||||||||
Bivalvia
- Arcida
- Glycymerididae
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Tucetona sp.
Iredale 1939
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3 specimens | |||||||||
Bivalvia
- Arcida
- Arcidae
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Anadara (Rasia) sp.
(Gray 1857)
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5 specimens | |||||||||
Bivalvia
- Carditida
- Crassatellidae
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Crassinella sp.
Guppy 1874
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16 specimens | |||||||||
Bivalvia
- Cardiida
- Cardiidae
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Microcardium sp.
Thiele 1934
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4 specimens | |||||||||
Laevicardium sp.
Swainson 1840
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3 specimens | |||||||||
Trigoniocardia sp.
Dall 1900
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1 specimen | |||||||||
sensu lato | ||||||||||
Bivalvia
- Cardiida
- Semelidae
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Abra sp.
Lamarck 1818
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23 specimens | |||||||||
Bivalvia
- Cardiida
- Tellinidae
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Eurytellina sp.
Fischer 1887
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2 specimens | |||||||||
spelled with current rank as Tellina (Eurytellina) | ||||||||||
Angulus sp.
Megerle 1811
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7 specimens | |||||||||
Bivalvia
- Cardiida
- Chamidae
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Chama sp.
Linnaeus 1758
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4 specimens | |||||||||
Bivalvia
- Cardiida
- Veneridae
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Ventricolaria sp.
Keen 1954
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5 specimens | |||||||||
synonym of Globivenus | ||||||||||
Gouldia sp.
Adams 1847
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15 specimens | |||||||||
Bivalvia
- Cardiida
- Kelliellidae
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Kelliella sp.
Sars 1865
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26 specimens | |||||||||
Bivalvia
- Pholadida
- Corbulidae
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Varicorbula sp.
Grant and Gale 1931
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2 specimens | |||||||||
spelled with current rank as Corbula (Varicorbula) | ||||||||||
Caryocorbula sp.
(Gardner 1926)
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12 specimens | |||||||||
original and current combination Corbula (Caryocorbula) | ||||||||||
Bivalvia
- Lucinida
- Lucinidae
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Gonimyrtea sp.
Marwick 1929
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1 specimen | |||||||||
Parvilucina sp.
(Dall 1901)
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2 specimens | |||||||||
sensu lato | ||||||||||
Scaphopoda
- Gadilida
- Entalinidae
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Bathoxiphus sp.
Pilsbry 1897
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4 specimens | |||||||||
Scaphopoda
- Gadilida
- Gadilidae
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Gadilopsis sp.
(Woodring 1925)
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66 specimens | |||||||||
Gastropoda
- Opisthobranchia
- Haminoeidae
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Atys (Aliculastrum) sp.
(Pilsbry 1896)
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1 specimen | |||||||||
original and current combination Aliculastrum | ||||||||||
Gastropoda
- Opisthobranchia
- Cylichnidae
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Scaphander sp.
Monfort 1810
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1 specimen | |||||||||
Acteocina sp.
Gray 1847
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16 specimens | |||||||||
Gastropoda
- Opisthobranchia
- Retusidae
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Sulcoretusa sp.
Burch 1945
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20 specimens | |||||||||
Volvulella (Volvulella) sp.
Newton 1891
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3 specimens | |||||||||
Gastropoda
- Opisthobranchia
- Cavoliniidae
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Styliola sp.
Gray 1850
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21 specimens | |||||||||
Clio sp.
Linnaeus 1767
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1 specimen | |||||||||
Cavolinia sp.
Avilgaard 1791
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28 specimens | |||||||||
Diacria sp.
Gray 1847
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2 specimens | |||||||||
Gastropoda
- Heterostropha
- Ringiculidae
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Ringincula (Ringiculella) sp.
Sacco 1892
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14 specimens | |||||||||
spelled with current rank as Ringicula (Ringiculella) | ||||||||||
Gastropoda
- Heterostropha
- Architectonicidae
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Pseudotorinia sp.
Sacco 1892
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3 specimens | |||||||||
Architectonica sp.
Röding 1798
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2 specimens | |||||||||
Gastropoda
- Heterostropha
- Murchisonellidae
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Anisocycla (Saccoina) sp.
(Dall and Bartsch 1904)
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1 specimen | |||||||||
Gastropoda
- Heterostropha
- Pyramidellidae
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Pyramidellidae indet.
Gray 1840
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8 specimens | |||||||||
Eulimella (Ebalina) sp.
(Thiele 1929)
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11 specimens | |||||||||
Besla sp.
Dall and Bartsch 1904
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3 specimens | |||||||||
Ondina sp.
de Folin 1870
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1 specimen | |||||||||
Gastropoda
- Neotaenioglossa
- Calyptraeidae
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Crucibulum (Crucibulum) sp.
Schumacher 1817
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1 specimen | |||||||||
Gastropoda
- Caecidae
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Brochina sp.
Gray 1837
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3 specimens | |||||||||
spelled with current rank as Caecum (Brochina) | ||||||||||
Gastropoda
- Vitrinellidae
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Cyclostremiscus (Ponocyclus) sp.
(Pilsbry 1953)
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2 specimens | |||||||||
Gastropoda
- Atlantidae
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Atlanta sp.
Lesueur 1817
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1 specimen | |||||||||
Gastropoda
- Rissoidae
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Rissoidae indet.
Gray 1847
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5 specimens | |||||||||
Alvania sp.
Risso 1826
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9 specimens | |||||||||
Gastropoda
- Strombidae
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Strombus sp.
Linnaeus 1758
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7 specimens | |||||||||
Gastropoda
- Neogastropoda
- Costellariidae
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Fusiturricula sp.
Woodring 1928
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4 specimens | |||||||||
Gastropoda
- Neogastropoda
- Marginellidae
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Dentimargo sp.
Cossmann 1899
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52 specimens | |||||||||
Volvarina sp.
(Hinds 1844)
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51 specimens | |||||||||
Volvarina sp.
(Hinds 1844)
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51 specimens | |||||||||
Gastropoda
- Neogastropoda
- Muricidae
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Typhis sp.
Montfort 1810
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1 specimen | |||||||||
Haustellum sp.
Schumacher 1817
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5 specimens | |||||||||
Gastropoda
- Neogastropoda
- Volutidae
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Voluta sp.
Linnaeaus 1758
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5 specimens | |||||||||
Voluta sp.
Linnaeaus 1758
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5 specimens | |||||||||
Gastropoda
- Neogastropoda
- Cystiscidae
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Persicula sp.
Schumacher 1817
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3 specimens | |||||||||
Granulina sp.
Jousseaume 1888
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30 specimens | |||||||||
Gastropoda
- Neogastropoda
- Olividae
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Olivella (Macgintiella) sp.
Olsson 1956
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179 specimens | |||||||||
Olivella (Minioliva) sp.
Olsson 1956
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18 specimens | |||||||||
Gastropoda
- Neogastropoda
- Ancillariidae
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Ancilla (Eburna) sp.
(Lamarck 1801)
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5 specimens | |||||||||
original and current combination Eburna | ||||||||||
Gastropoda
- Neogastropoda
- Drilliidae
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Clathrodrillia sp.
Dall 1918
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4 specimens | |||||||||
Syntomodrillia sp.
Woodring 1928
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6 specimens | |||||||||
Leptadrillia sp.
Woodring 1928
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15 specimens | |||||||||
Gastropoda
- Neogastropoda
- Raphitomidae
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Pleurotomella sp.
Verrill 1872
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2 specimens | |||||||||
Gastropoda
- Neogastropoda
- Clathurellidae
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Nannodiella sp.
(Dall 1919)
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5 specimens | |||||||||
Glyphostoma (Glyphostoma) sp.
Gabb 1872
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4 specimens | |||||||||
Gastropoda
- Neogastropoda
- Conidae
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Conus sp.
Linnaeus 1758
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14 specimens | |||||||||
Gastropoda
- Neogastropoda
- Borsoniidae
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Microdrillia sp.
Casey 1903
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1 specimen | |||||||||
Gastropoda
- Neogastropoda
- Mangeliidae
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Tenaturris sp.
Woodring 1928
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5 specimens | |||||||||
Ithycythara sp.
Woodring 1928
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10 specimens | |||||||||
Saccharoturris sp.
Woodring 1928
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1 specimen | |||||||||
Kurtziella (Kurtziella) sp.
Dall 1918
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2 specimens | |||||||||
Gastropoda
- Neogastropoda
- Terebridae
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Strioterebrum sp.
Sacco 1891
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23 specimens | |||||||||
Gastropoda
- Neogastropoda
- Turridae
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Polystira sp.
Woodring 1928
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6 specimens | |||||||||
Gastropoda
- Neogastropoda
- Nassariidae
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Nassarius sp.
Duméril 1806
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95 specimens | |||||||||
sensu lato | ||||||||||
Gastropoda
- Neogastropoda
- Columbellidae
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Sincola (? Sinaxila) sp.
Olsson and Harbison 1953
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22 specimens | |||||||||
Mitrella sp.
Risso 1826
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14 specimens | |||||||||
Nassarina (Nassarina) sp.
Dall 1889
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3 specimens | |||||||||
Gastropoda
- Neogastropoda
- Buccinidae
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Antillophos (Antillophos) sp.
Woodring 1928
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80 specimens | |||||||||
Gastropoda
- Neogastropoda
- Mitridae
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Subcancilla sp.
Olsson and Harbison 1953
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2 specimens | |||||||||
Gastropoda
- Xenophoridae
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Xenophora sp.
Fischer von Waldheim 1807
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1 specimen | |||||||||
Gastropoda
- Vermetidae
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Serpulorbis sp.
Sasso 1827
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53 specimens | |||||||||
Gastropoda
- Naticidae
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Natica (? Natica) sp.
Scopoli 1777
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2 specimens | |||||||||
Polinices sp.
Monfort 1810
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16 specimens | |||||||||
Gastropoda
- Cerithiopsidae
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Cerithiopsis sp.
Hanley 1849
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1 specimen | |||||||||
Gastropoda
- Epitoniidae
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Epitonium (Asperiscala) sp.
(de Boury 1909)
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6 specimens | |||||||||
Gastropoda
- Eulimidae
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Eulimidae indet.
Philippi 1853
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5 specimens | |||||||||
Niso sp.
Risso 1826
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2 specimens | |||||||||
Gastropoda
- Pseudomelaniidae
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Compsodrillia sp.
Woodring 1928
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3 specimens | |||||||||
? Miraclathurella sp.
Woodring 1928
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3 specimens | |||||||||
Carinodrillia (? Buchema) sp.
(Dall 1919)
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1 specimen | |||||||||
Gastropoda
- Turritellidae
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Vermicularia sp.
Lamarck 1799
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2 specimens | |||||||||
Turritella sp.
Lamarck 1799
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55 specimens | |||||||||
Gastropoda
- Cerithiidae
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Alabina sp.
Dall 1902
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18 specimens | |||||||||
Gastropoda
- Trochidae
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Trochidae indet.
Rafinesque 1815
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1 specimen | |||||||||
Microgaza sp.
Dall 1881
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2 specimens | |||||||||
Calliostoma sp.
Swainson 1840
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1 specimen | |||||||||
see common names |
Geography
Country: | Panama | State/province: | Bocas |
Coordinates: | 9.1° North, 81.6° West (view map) | ||
Paleocoordinates: | 9.0° North, 81.1° West | ||
Basis of coordinate: | stated in text | ||
Geographic resolution: | outcrop |
Time
Period: | Neogene | Epoch: | Pliocene |
Stage: | Piacenzian | 10 m.y. bin: | Cenozoic 6 |
Key time interval: | Piacenzian | ||
Age range of interval: | 3.60000 - 2.58800 m.y. ago | ||
Age estimate: | 2.8 Ma (age-depth)3.6 to 2.1 Ma (other) |
Stratigraphy
Geological group: | Bocas del Toro | Formation: | Escudo de Veraguas | ||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date from Cheetham et al. (1999). |
Lithology and environment
Primary lithology: | burrows,bioturbation,fine poorly lithified argillaceous sandstone |
Secondary lithology: | burrows,bioturbation poorly lithified argillaceous siltstone |
Includes fossils? | Y |
Includes fossils? | Y |
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems. | |
Environment: | deep-water indet. |
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils |
Collection methods and comments
Collection excludes: | all microfossils,some macrofossils |
Collection methods: | bulk,sieve,field collection |
Reason for describing collection: | paleoecologic analysis |
Museum repositories: | BMNH,NMB,USNM |
Collection method comments: COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 6/4/88. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB), | |
Taxonomic list comments:NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung. |
Metadata
Also known as: | NMB Loc. 17834 | ||
Database number: | 79989 | ||
Authorizer: | A. Hendy | Enterer: | K. Bulinski, A. Hendy |
Modifier: | A. Hendy | Research group: | marine invertebrate |
Created: | 2008-03-31 18:57:42 | Last modified: | 2013-11-25 13:37:23 |
Access level: | authorizer only | Released: | 2009-03-31 18:57:42 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
17667. | L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy] |