PPP# 361 - Escudo de Veraguas [Escudo de Veraguas Fm]: Piacenzian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gymnolaemata - Cheilostomata - Membraniporidae
Cupuladria aff. canariensis
11 specimens
Cupuladria aff. biporosa Canu and Bassler 1923
10 specimens
Gymnolaemata - Cheilostomata - Lepraliellidae
Celleporaria albirostris
1 specimen
Gymnolaemata - Cheilostomata - Gemelliporellidae
Gemelliporella glabra
10 specimens
Gymnolaemata - Cheilostomata - Petraliidae
Petraliella bisinuata
1 specimen
Gymnolaemata - Cheilostomata - Smittinidae
Parasmittina sp.
1 specimen
Gymnolaemata - Cheilostomata - Phidoloporidae
Schedocleidochasma cleidostoma
1 specimen
Gymnolaemata - Cheilostomata - Mamilloporidae
Mamillopora informal n. sp. 1
1 specimen
Bivalvia - Nuculanida - Nuculanidae
Saccella sp. Woodring 1925
2 specimens
Bivalvia - Nuculida - Nuculidae
Nucula (Lamellinucula) sp. (Schenck 1944)
12 specimens
Bivalvia - Mytilida - Crenellidae
Crenella sp. Brown 1827
16 specimens
Bivalvia - Ostreida - Gryphaeidae
? Hyotissa sp. Stenzel 1971
2 specimens
Bivalvia - Pectinida - Pectinidae
? Spathochlamys sp. Waller 1993
7 specimens
Spathochlamys sp. Waller 1993
1 specimen
Argopecten sp. Monterosato 1889
13 specimens
Bivalvia - Pectinida - Propeamussiidae
Cyclopecten sp. Verrill 1897
3 specimens
Bivalvia - Arcida - Limopsidae
Limopsis sp. Sassi 1827
5 specimens
Bivalvia - Arcida - Glycymerididae
Tucetona sp. Iredale 1939
3 specimens
Bivalvia - Arcida - Arcidae
Anadara (Rasia) sp. (Gray 1857)
5 specimens
Bivalvia - Carditida - Crassatellidae
Crassinella sp. Guppy 1874
16 specimens
Bivalvia - Cardiida - Cardiidae
Microcardium sp. Thiele 1934
4 specimens
Laevicardium sp. Swainson 1840
3 specimens
Trigoniocardia sp. Dall 1900
1 specimen
sensu lato
Bivalvia - Cardiida - Semelidae
Abra sp. Lamarck 1818
23 specimens
Bivalvia - Cardiida - Tellinidae
Eurytellina sp. Fischer 1887
2 specimens
spelled with current rank as Tellina (Eurytellina)
Angulus sp. Megerle 1811
7 specimens
Bivalvia - Cardiida - Chamidae
Chama sp. Linnaeus 1758
4 specimens
Bivalvia - Cardiida - Veneridae
Ventricolaria sp. Keen 1954
5 specimens
synonym of Globivenus
Gouldia sp. Adams 1847
15 specimens
Bivalvia - Cardiida - Kelliellidae
Kelliella sp. Sars 1865
26 specimens
Bivalvia - Pholadida - Corbulidae
Varicorbula sp. Grant and Gale 1931
2 specimens
spelled with current rank as Corbula (Varicorbula)
Caryocorbula sp. (Gardner 1926)
12 specimens
original and current combination Corbula (Caryocorbula)
Bivalvia - Lucinida - Lucinidae
Gonimyrtea sp. Marwick 1929
1 specimen
Parvilucina sp. (Dall 1901)
2 specimens
sensu lato
Scaphopoda - Gadilida - Entalinidae
Bathoxiphus sp. Pilsbry 1897
4 specimens
Scaphopoda - Gadilida - Gadilidae
Gadilopsis sp. (Woodring 1925)
66 specimens
Gastropoda - Opisthobranchia - Haminoeidae
Atys (Aliculastrum) sp. (Pilsbry 1896)
1 specimen
original and current combination Aliculastrum
Gastropoda - Opisthobranchia - Cylichnidae
Scaphander sp. Monfort 1810
1 specimen
Acteocina sp. Gray 1847
16 specimens
Gastropoda - Opisthobranchia - Retusidae
Sulcoretusa sp. Burch 1945
20 specimens
Volvulella (Volvulella) sp. Newton 1891
3 specimens
Gastropoda - Opisthobranchia - Cavoliniidae
Styliola sp. Gray 1850
21 specimens
Clio sp. Linnaeus 1767
1 specimen
Cavolinia sp. Avilgaard 1791
28 specimens
Diacria sp. Gray 1847
2 specimens
Gastropoda - Heterostropha - Ringiculidae
Ringincula (Ringiculella) sp. Sacco 1892
14 specimens
spelled with current rank as Ringicula (Ringiculella)
Gastropoda - Heterostropha - Architectonicidae
Pseudotorinia sp. Sacco 1892
3 specimens
Architectonica sp. Röding 1798
2 specimens
Gastropoda - Heterostropha - Murchisonellidae
Anisocycla (Saccoina) sp. (Dall and Bartsch 1904)
1 specimen
Gastropoda - Heterostropha - Pyramidellidae
Pyramidellidae indet. Gray 1840
8 specimens
Eulimella (Ebalina) sp. (Thiele 1929)
11 specimens
Besla sp. Dall and Bartsch 1904
3 specimens
Ondina sp. de Folin 1870
1 specimen
Gastropoda - Neotaenioglossa - Calyptraeidae
Crucibulum (Crucibulum) sp. Schumacher 1817
1 specimen
Gastropoda - Caecidae
Brochina sp. Gray 1837
3 specimens
spelled with current rank as Caecum (Brochina)
Gastropoda - Vitrinellidae
Cyclostremiscus (Ponocyclus) sp. (Pilsbry 1953)
2 specimens
Gastropoda - Atlantidae
Atlanta sp. Lesueur 1817
1 specimen
Gastropoda - Rissoidae
Rissoidae indet. Gray 1847
5 specimens
Alvania sp. Risso 1826
9 specimens
Gastropoda - Strombidae
Strombus sp. Linnaeus 1758
7 specimens
Gastropoda - Neogastropoda - Costellariidae
Fusiturricula sp. Woodring 1928
4 specimens
Gastropoda - Neogastropoda - Marginellidae
Dentimargo sp. Cossmann 1899
52 specimens
Volvarina sp. (Hinds 1844)
51 specimens
Volvarina sp. (Hinds 1844)
51 specimens
Gastropoda - Neogastropoda - Muricidae
Typhis sp. Montfort 1810
1 specimen
Haustellum sp. Schumacher 1817
5 specimens
Gastropoda - Neogastropoda - Volutidae
Voluta sp. Linnaeaus 1758
5 specimens
Voluta sp. Linnaeaus 1758
5 specimens
Gastropoda - Neogastropoda - Cystiscidae
Persicula sp. Schumacher 1817
3 specimens
Granulina sp. Jousseaume 1888
30 specimens
Gastropoda - Neogastropoda - Olividae
Olivella (Macgintiella) sp. Olsson 1956
179 specimens
Olivella (Minioliva) sp. Olsson 1956
18 specimens
Gastropoda - Neogastropoda - Ancillariidae
Ancilla (Eburna) sp. (Lamarck 1801)
5 specimens
original and current combination Eburna
Gastropoda - Neogastropoda - Drilliidae
Clathrodrillia sp. Dall 1918
4 specimens
Syntomodrillia sp. Woodring 1928
6 specimens
Leptadrillia sp. Woodring 1928
15 specimens
Gastropoda - Neogastropoda - Raphitomidae
Pleurotomella sp. Verrill 1872
2 specimens
Gastropoda - Neogastropoda - Clathurellidae
Nannodiella sp. (Dall 1919)
5 specimens
Glyphostoma (Glyphostoma) sp. Gabb 1872
4 specimens
Gastropoda - Neogastropoda - Conidae
Conus sp. Linnaeus 1758
14 specimens
Gastropoda - Neogastropoda - Borsoniidae
Microdrillia sp. Casey 1903
1 specimen
Gastropoda - Neogastropoda - Mangeliidae
Tenaturris sp. Woodring 1928
5 specimens
Ithycythara sp. Woodring 1928
10 specimens
Saccharoturris sp. Woodring 1928
1 specimen
Kurtziella (Kurtziella) sp. Dall 1918
2 specimens
Gastropoda - Neogastropoda - Terebridae
Strioterebrum sp. Sacco 1891
23 specimens
Gastropoda - Neogastropoda - Turridae
Polystira sp. Woodring 1928
6 specimens
Gastropoda - Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806
95 specimens
sensu lato
Gastropoda - Neogastropoda - Columbellidae
Sincola (? Sinaxila) sp. Olsson and Harbison 1953
22 specimens
Mitrella sp. Risso 1826
14 specimens
Nassarina (Nassarina) sp. Dall 1889
3 specimens
Gastropoda - Neogastropoda - Buccinidae
Antillophos (Antillophos) sp. Woodring 1928
80 specimens
Gastropoda - Neogastropoda - Mitridae
Subcancilla sp. Olsson and Harbison 1953
2 specimens
Gastropoda - Xenophoridae
Xenophora sp. Fischer von Waldheim 1807
1 specimen
Gastropoda - Vermetidae
Serpulorbis sp. Sasso 1827
53 specimens
Gastropoda - Naticidae
Natica (? Natica) sp. Scopoli 1777
2 specimens
Polinices sp. Monfort 1810
16 specimens
Gastropoda - Cerithiopsidae
Cerithiopsis sp. Hanley 1849
1 specimen
Gastropoda - Epitoniidae
Epitonium (Asperiscala) sp. (de Boury 1909)
6 specimens
Gastropoda - Eulimidae
Eulimidae indet. Philippi 1853
5 specimens
Niso sp. Risso 1826
2 specimens
Gastropoda - Pseudomelaniidae
Compsodrillia sp. Woodring 1928
3 specimens
? Miraclathurella sp. Woodring 1928
3 specimens
Carinodrillia (? Buchema) sp. (Dall 1919)
1 specimen
Gastropoda - Turritellidae
Vermicularia sp. Lamarck 1799
2 specimens
Turritella sp. Lamarck 1799
55 specimens
Gastropoda - Cerithiidae
Alabina sp. Dall 1902
18 specimens
Gastropoda - Trochidae
Trochidae indet. Rafinesque 1815
1 specimen
Microgaza sp. Dall 1881
2 specimens
Calliostoma sp. Swainson 1840
1 specimen
see common names

Geography
Country:Panama State/province:Bocas
Coordinates: 9.1° North, 81.6° West (view map)
Paleocoordinates:9.0° North, 81.1° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Pliocene
Stage:Piacenzian 10 m.y. bin:Cenozoic 6
Key time interval:Piacenzian
Age range of interval:3.60000 - 2.58800 m.y. ago
Age estimate:2.8 Ma (age-depth)3.6 to 2.1 Ma (other)
Stratigraphy
Geological group:Bocas del Toro Formation:Escudo de Veraguas
Stratigraphic resolution:bed
Stratigraphy comments: STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date from Cheetham et al. (1999).
Lithology and environment
Primary lithology:burrows,bioturbation,fine poorly lithified argillaceous sandstone
Secondary lithology:burrows,bioturbation poorly lithified argillaceous siltstone
Includes fossils?Y
Includes fossils?Y
Lithology description: GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.
Environment:deep-water indet.
Geology comments: ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:all microfossils,some macrofossils
Collection methods:bulk,sieve,field collection
Reason for describing collection:paleoecologic analysis
Museum repositories:BMNH,NMB,USNM
Collection method comments: COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 6/4/88. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),
Taxonomic list comments:NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Metadata
Also known as:NMB Loc. 17834
Database number:79989
Authorizer:A. Hendy Enterer:K. Bulinski, A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2008-03-31 18:57:42 Last modified:2013-11-25 13:37:23
Access level:authorizer only Released:2009-03-31 18:57:42
Creative Commons license:CC BY
Reference information

Primary reference:

17667. L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]