CSUN Loc. 374, Las Llajas Canyon: Lutetian, California
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Echinoidea
- Spatangoida
- Schizasteridae
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Schizaster diabloensis
Kew 1920
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Squires 1984 | |||||||||
Echinoidea
- Spatangoida
- Brissidae
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Eupatagus stevensi n. sp.
Grant and Hertlein 1938
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Grant and Hertlein 1938 | |||||||||
Chondrichthyes
- Lamniformes
- Odontaspididae
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Odontaspis sp.
Agassiz 1838
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Squires 1984 | |||||||||
Chondrichthyes
- Lamniformes
- Mitsukurinidae
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Striatolamia macrota
(Agassiz 1838)
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Squires 2001 | |||||||||
original and current combination Otodus macrotus | ||||||||||
Rhynchonellata
- Terebratulida
- Terebrataliidae
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Terebratalia aff. batequia
Sandy et al. 1995
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Squires 2001 | |||||||||
Polychaeta
- Sabellida
- Serpulidae
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Rotularia (Rotularia) tejonensis
Arnold 1909
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Squires 1984 | |||||||||
recombined as Rotularia (Rotularia) tejonense | ||||||||||
Cephalopoda
- Nautilida
- Aturiidae
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Aturia myrlae
(Hanna 1927)
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Squires 1984 | |||||||||
Aturia myrlae
(Hanna 1927)
|
Squires 1984 | |||||||||
Cephalopoda
- Nautilida
- Nautilidae
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Kummelonautilus sp.
Matsumoto 1984
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Squires 2001 | |||||||||
Scaphopoda
- Dentaliida
- Dentaliidae
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Dentalium stentor
Anderson and Hanna 1925
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Squires 1984 | |||||||||
Gastropoda
- Heterostropha
- Architectonicidae
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Architectonica (Stellaxis) cognata
(Gabb 1864)
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Squires 1984 | |||||||||
Gastropoda
- Opisthobranchia
- Akeridae
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Akera maga
Vokes 1939
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Squires 2001 | |||||||||
Gastropoda
- Opisthobranchia
- Retusidae
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Cylichnina tantilla
(Anderson and Hanna 1925)
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Squires 1984 | |||||||||
Gastropoda
- Opisthobranchia
- Philinidae
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Megistostoma gabbianum
(Stoliczka 1868)
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Squires 1984 | |||||||||
Gastropoda
- Turritellidae
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Turritella andersoni
Dickerson 1916
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Squires 1984 | |||||||||
as Turritella andersoni lawsoni | ||||||||||
Turritella uvasana
Conrad 1855
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Squires 1984 | |||||||||
as Turritella uvasana applinae | ||||||||||
Gastropoda
- Brachytrematidae
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Benoistia umpquaensis
Turner 1938
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Squires 1984 | |||||||||
Gastropoda
- Ampullinidae
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Eocernina hannibali
(Dickerson 1914)
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Squires 1984 | |||||||||
Pachycrommium clarki
(Stewart 1927)
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Squires 1984 | |||||||||
Gastropoda
- Epitoniidae
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Cirsotrema sp.
Mörch 1852
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Squires 1984 | |||||||||
Gastropoda
- Naticidae
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Polinices (Euspira) nuciformis
(Gabb 1864)
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Squires 1984 | |||||||||
Gastropoda
- Neogastropoda
- Fasciolariidae
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Clavilithes informal n. sp. B
|
Squires 1984 | |||||||||
Clavilithes tabulatus
(Dickerson 1913)
|
Squires 1984 | |||||||||
Fusinus teglandae
(Hanna 1927)
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Squires 1984 | |||||||||
Gastropoda
- Neogastropoda
- Buccinidae
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Ancistrolepis carolineae n. sp.
Squires 1984
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Squires 1984 | |||||||||
Gastropoda
- Neogastropoda
- Melongenidae
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Ficopsis cooperiana
Stewart 1927
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Squires 1984 | |||||||||
original and current combination Fusus (Hemifusus) cooperii | ||||||||||
Gastropoda
- Neogastropoda
- Clavatulidae
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Fusiturricula (Crenaturricula) crenatospira
(Cooper 1894)
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Squires 1984 | |||||||||
recombined as Crenaturricula crenatospira | ||||||||||
also as Fusiturricula (Crenaturricula) crenatospira domenginica | ||||||||||
Gastropoda
- Neogastropoda
- Conidae
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Conus hornii
Gabb 1864
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Squires 1984 | |||||||||
as Conus hornii umpquaensis | ||||||||||
Gastropoda
- Neogastropoda
- Turridae
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? Cochlespiropsis jenkinsi n. sp.
Squires 2001
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Squires 2001 | |||||||||
Surculites mathewsonii
(Gabb 1864)
|
Squires 1984 | |||||||||
Gastropoda
- Neogastropoda
- Volutidae
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Lyrischapa lajollaensis
(Hanna 1927)
|
Squires 1984 | |||||||||
Lapparia eomagna
(Vokes 1939)
|
Squires 2001 | |||||||||
original and current combination Voluta eomagna | ||||||||||
Gastropoda
- Neogastropoda
- Muricidae
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Pseudoperissolax blakei
(Conrad 1855)
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Squires 1984 | |||||||||
as Pseudoperissolax blakei praeblakei | ||||||||||
Gastropoda
- Neogastropoda
- Harpidae
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Eocithara mutica
(Lamarck 1803)
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Squires 1984 | |||||||||
original and current combination Harpa mutica | ||||||||||
as Eocithara mutica californiensis | ||||||||||
Gastropoda
- Strombidae
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Ectinochilus (Macilentos) macilentus
(White 1889)
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Squires 1984 | |||||||||
Gastropoda
- Cypraeidae
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Bernaya kaylinae n. sp.
Groves and Squires 2023
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Groves and Squires 2023 | |||||||||
Cypraea castacensis
Stewart 1927
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Squires 1984 | |||||||||
= Subepona leahae n. sp.
Groves and Squires 2023
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Groves and Squires 2023 | |||||||||
Gastropoda
- Cymatiidae
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Ranella katherineae n. sp.
Squires 1983
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Olequahia domenginica
(Vokes 1939)
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Squires 1984 | |||||||||
original and current combination Ranella domenginica | ||||||||||
Gastropoda
- Cassidae
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Phalium (Semicassis) tuberculiformis
(Gabb 1864)
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Squires 1984 | |||||||||
Galeodea (Caliagaleodea) californica
Clark 1942
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Squires 1984 | |||||||||
recombined as Galeodea californica | ||||||||||
Gastropoda
- Ranellidae
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Cymatium janetae n. sp.
Squires 1983
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Cymatium (Septa) janetae
Squires 1983
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Squires 1984 | |||||||||
Gastropoda
- Ficidae
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Ficopsis remondii
(Gabb 1864)
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Squires 1984 | |||||||||
recombined as Ficus remondii | ||||||||||
as Ficopsis remondii crescentensis | ||||||||||
Gastropoda
- Xenophoridae
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Xenophora stocki
Dickerson 1916
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Squires 1984 | |||||||||
Bivalvia
- Mytilida
- Mytilidae
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Brachidontes (Brachidontes) cowlitzensis
(Weaver and Palmer 1922)
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Squires 1984 | |||||||||
Bivalvia
- Pectinida
- Spondylidae
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Spondylus carlosensis
Anderson 1905
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Squires 1984 | |||||||||
Bivalvia
- Ostreida
- Malleidae
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Nayadina (Exputens) llajasensis
(Clark 1934)
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Squires 1984 | |||||||||
Bivalvia
- Ostreida
- Pinnidae
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Pinna llajasensis n. sp.
Squires 1983
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Bivalvia
- Ostreida
- Ostreidae
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Ostrea idriaensis
Gabb 1869
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Squires 1984 | |||||||||
recombined as Acutostrea idriaensis | ||||||||||
Bivalvia
- Ostreida
- Gryphaeidae
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Pycnodonte (Phygraea) cf. pacifica
Squires and Demetrion 1990
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Squires 2001 | |||||||||
recombined as Phygraea pacifica | ||||||||||
Bivalvia
- Arcida
- Glycymerididae
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Glycymeris (Glycymerita) sagittata
(Gabb 1864)
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Squires 1984 | |||||||||
recombined as Glycymerita sagittata | ||||||||||
Bivalvia
- Cardiida
- Cardiidae
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Acanthocardia (Agnocardia) sorrentoensis
(Hanna 1927)
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Squires 2001 | |||||||||
recombined as Agnocardia sorrentoensis | ||||||||||
Acanthocardia (Schedocardia) brewerii
(Gabb 1864)
|
Squires 1984 | |||||||||
recombined as Schedocardia brewerii | ||||||||||
Nemocardium linteum
(Conrad 1855)
|
Squires 1984 | |||||||||
recombined as Nemocardium (Nemocardium) linteum | ||||||||||
Bivalvia
- Cardiida
- Psammobiidae
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Gari texta
Gabb 1864
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Squires 2001 | |||||||||
recombined as Gari (Gobraeus) texta | ||||||||||
Gari cf. eoundulata
Vokes 1939
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Squires 1984 | |||||||||
recombined as Gari (Gobraeus) eoundulata | ||||||||||
Bivalvia
- Cardiida
- Veneridae
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Marcia (Mercimonia) bunkeri
(Hanna 1927)
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Squires 1984 | |||||||||
Pitar uvasana
(Conrad 1855)
|
Squires 2001 | |||||||||
Subspecies: Pitar uvasana coquillensis | ||||||||||
Pitar (Calpitaria) uvasanus
(Conrad 1855)
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Squires 1984 | |||||||||
recombined as Pitar uvasana | ||||||||||
Bivalvia
- Pholadida
- Teredinidae
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? Teredo sp.
Linnaeus 1758
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Squires 1984 | |||||||||
Bivalvia
- Solenida
- Solenidae
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Solena (Eosolen) navacularis
(Anderson and Hanna 1925)
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Squires 1984 | |||||||||
recombined as Solena (Eosolen) novacularis | ||||||||||
Bivalvia
- Lucinida
- Lucinidae
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Claibornites diegoensis
(Dickerson 1916)
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Squires 1984 | |||||||||
recombined as Claibornites (Claibornites) diegoensis | ||||||||||
Bivalvia
- Carditida
- Crassatellidae
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Crassatella uvasana
Conrad 1855
|
Squires 1984 | |||||||||
Bivalvia
- Carditida
- Carditidae
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Venericardia (Pacificor) hornii
(Gabb 1864)
|
Squires 1984 | |||||||||
as Venericardia (Pacificor) hornii calafia | ||||||||||
Anthozoa
- Scleractinia
- Caryophylliidae
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? Trochocyathus striatus
(Gabb 1864)
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Squires 1984 | |||||||||
Anthozoa
- Scleractinia
- Oculinidae
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Archohelia clarki
Vaughan 1927
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Squires 2001 | |||||||||
recombined as Petrophyllia clarki | ||||||||||
Asterocyclinidae
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Pseudophragmina (Proporocyclina) clarki
(Cushman 1920)
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Squires 1984 | |||||||||
recombined as Pseudophragmina clarki | ||||||||||
see common names |
Geography
Country: | United States | State/province: | California | County: | Ventura |
Coordinates: | 34.3° North, 118.7° West (view map) | ||||
Paleocoordinates: | 35.1° North, 104.6° West | ||||
Basis of coordinate: | estimated from map | ||||
Geographic resolution: | small collection |
Time
Period: | Paleogene | Epoch: | Eocene |
Stage: | Lutetian | 10 m.y. bin: | Cenozoic 2 |
*Period: | Early/Lower Tertiary - Middle Tertiary | *Epoch: | Early/Lower Eocene - Middle Eocene |
*International age/stage: | Middle Lutetian - Early/Lower Ypresian | *Local age/stage: | Domengine - Domengine |
Key time interval: | Lutetian | ||
Age range of interval: | 47.80000 - 41.20000 m.y. ago | ||
* legacy (obsolete) database fields |
Stratigraphy
Formation: | Llajas | ||||
Stratigraphic resolution: | bed | ||||
Stratigraphy comments: The Stewart bed is a 1 m thick sandstone unit in the middle of the Llajas formation. Lutetian, chron 21r according to Squires (2001). |
Lithology and environment
Primary lithology: | bioturbation lithified sandstone |
Includes fossils? | Y |
Lithology description: sandstone. | |
Environment: | transition zone/lower shoreface |
Geology comments: The "Stewart bed" is located in a section of transition from shallow-marine to outer slope and shelf facies. A nearshore environment is interpreted for that portion of the shallow-marine (transgressive) facies which consists of laminated beds alternating with bioturbated beds. In such cases, the finely laminated sand represents storm-influenced stratification, and the biogenic reworking is the inter-storm activity. Shell fragments commonly are concentrated as channel lags in surge channels. Fossils were apparently subjected to wave action because the apices of elongate shells such as Turritella are bimodally oriented in opposite directions. The deeper the water, the less amount of laminated sandstone and the more amount of bioturbated sandstone. Such areas, which were farther below effective wave base, existed 1) where the shallow-marine (transgressive) facies grades into the outer shelf to slope facies and 2) in the shallow-marine (regressive) facies. One of the deepest-water shallow-marine beds was the "Stewart bed." |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils |
Collection methods and comments
Collection excludes: | all microfossils,some macrofossils |
Collection methods: | bulk,field collection |
Reason for describing collection: | taxonomic analysis |
Museum repositories: | LACM |
Collection method comments: Repository: formerly CSUN (California State University Northridge) |
Metadata
Also known as: | LACMIP loc. 7242; LACMIP loc. 40374 | ||
Database number: | 8012 | ||
Authorizer: | L. Ivany, M. Clapham, A. Hendy, J. Marcot | Enterer: | E. Kowalski, J. Marcot, A. Hendy, M. Clapham |
Modifier: | J. Marcot | Research group: | marine invertebrate |
Created: | 2000-06-12 07:08:25 | Last modified: | 2023-11-19 19:13:00 |
Access level: | the public | Released: | 2000-06-12 07:08:25 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
519. | R. L. Squires. 1983. New Mollusks from the Lower Middle Eocene Llajas Formation, Southern California. Journal of Paleontology 57(2):354-362 [L. Ivany/E. Kowalski/J. Marcot] |
Secondary references:
32966 | U. S. Grant and L. G. Hertlein. 1938. The West American Cenozoic Echinoidea. Publications of the University of California at Los Angeles in Mathematical and Physical Sciences 2:1-225 [L. Villier/L. Villier] | |
86508 | L. T. Groves and R. L. Squires. 2023. Revision of northeast Pacific Paleogene cypraeoidean gastropods (Mollusca), including recognition of three new species: Implications for paleobiogeographic distribution and faunal turnover. PaleoBios 40(10):1-52 [J. Marcot/J. Marcot] | |
59877 | R. L. Squires. 1984. Megapaleontology of the Eocene Llajas Formation, Simi Valley, California. Contributions in Science, Natural History Museum of Los Angeles County 350:1-76 [M. Clapham/M. Clapham] | |
13852 | R. L. Squires. 2001. Additions to the Eocene megafossil fauna of the Llajas Formation, Simi Valley, southern California. Contributions in Science (Natural History Museum of Los Angeles County) 489:1-40 [A. Miller/A. Hendy/A. Hendy] |