Woodring Loc. 182a - western area [upper Gatun Fm]: Tortonian, Panama

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda - Heterostropha - Architectonicidae
Architectonica (Architectonica) nobilis (Röding 1798)
Woodring 1959 R category
original and current combination Architectonica nobilis
Gastropoda - Turritellidae
Turritella (Torcula) mimetes Brown and Pilsbry 1911
recombined as Turritella (Broderiptella) mimetes
Gastropoda - Naticidae
Polinices stanislasmeunieri Maury 1917
Stigmaulax guppiana (Toula 1909)
Gastropoda - Neogastropoda - Buccinidae
Cymatophos veatchi (Olsson 1922)
Woodring 1964 C category
SUBSPECIES: Cymatophos veatchi erymnus
Gastropoda - Neogastropoda - Cancellariidae
Cancellaria (Cancellaria) epistomifera (Guppy 1876)
Woodring 1970 R category
SUBSPECIES: Cancellaria (Cancellaria) epistomifera lipara
    = Cancellaria sathra Woodring 1973
Landau et al. 2012
Cancellaria (Cancellaria) tapeina Woodring 1970
Woodring 1970 R category
recombined as Cancellaria tapeina
Cancellaria (Cancellaria) apimela Woodring 1970
Woodring 1970 R category
recombined as Cancellaria apimela
Gastropoda - Neogastropoda - Drilliidae
Crassispira (Hindsiclava) consors (Sowerby 1850)
Woodring 1970 R category
recombined as Hindsiclava consors
Gastropoda - Neogastropoda - Terebridae
Terebra (Oreoterebra) isaacpetiti (Maury 1925)
Woodring 1970 R category
Gastropoda - Neogastropoda - Conidae
Conus burckhardti Bose 1906
Woodring 1970 R category
Gastropoda - Neogastropoda - Turridae
Pleurolira (Polystira) tenagos Gardner 1947
Woodring 1970 R category
    = Polystira sp. Woodring 1928
Todd and Johnson 2013
Gastropoda - Neogastropoda - Olividae
Oliva (Oliva) reticularis (Lamarck 1811)
Woodring 1964 F category
original and current combination Oliva reticularis
Gastropoda - Neogastropoda - Volutidae
Voluta alfaroi Dall 1912
Woodring 1964 F category
SUBSPECIES: Voluta alfaroi eurytera
Gastropoda - Neogastropoda - Muricidae
Murex (Murex) polynematicus (Brown and Pilsbry 1911)
Woodring 1959 R category
recombined as Vokesimurex polynematicus
Gastropoda - Strombidae
Strombus gatunensis Toula 1908
Woodring 1959 R category
Gastropoda - Cypraeidae
Cypraea (Muracypraea) henekeni (Sowerby 1850)
Woodring 1959 R category
    = Muracypraea isthmica Schilder 1927
Landau and Groves 2011
Gastropoda - Personidae
Distorsio (Rhysema) decussata (Valenciennes 1832)
Woodring 1959 R category
recombined as Distorsio decussata
SUBSPECIES: Distorsio (Rhysema) decussata gatunensis
Bivalvia - Pectinida - Pectinidae
Pecten (Euvola) bowdenensis Dall 1898
Woodring 1982 R category
recombined as Euvola bowdenensis
Argopecten levicostatus (Toula 1909)
Woodring 1982 C category
Bivalvia - Arcida - Arcidae
Anadara (Rasia) dariensis (Brown and Pilsbry 1911)
Woodring 1973 R category
Anadara (Tosarca) veatchi (Olsson 1922)
Woodring 1973 R category
Bivalvia - Arcida - Glycymerididae
Glycymeris (Tucetona) epacra
Woodring 1973 R category
original and current combination Tucetona
Bivalvia - Cardiida - Cardiidae
Trachycardium (Dallocardia) dominicense (Gabb 1873)
Woodring 1982 F category
SUBSPECIES: Trachycardium (Dallocardia) dominicense hadratatum
Bivalvia - Cardiida - Solecurtidae
Solecurtus broggii Pilsbry and Olsson 1941
Woodring 1982 R category
SUBSPECIES: Solecurtus broggii gatunensis
Bivalvia - Cardiida - Chamidae
Arcinella arcinella (Linnaeus 1767)
Woodring 1982 F category
Bivalvia - Cardiida - Veneridae
Dosinia (Dosinia) delicatissima (Brown and Pilsbry 1913)
Woodring 1982 R category
Chione (Chionopsis) tegulum (Brown and Pilsbry 1911)
Woodring 1982 F category
Lamelliconcha cora (Brown and Pilsbry 1911)
Woodring 1982 R category
Macrocallista (Paradione) maculata (Linnaeus 1758)
Woodring 1982 F category
recombined as Macrocallista maculata
Bivalvia - Cardiida - Anatinellidae
Raeta undulata (Gould 1851)
Woodring 1982 F category
see common names

Geography
Country:Panama
Coordinates: 9.2° North, 80.0° West (view map)
Paleocoordinates:8.9° North, 78.6° West
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Tortonian 10 m.y. bin:Cenozoic 6
Key time interval:Tortonian
Age range of interval:11.63000 - 7.24600 m.y. ago
Age estimate:8.5 Ma (other)
Stratigraphy
Formation:Gatun Member:upper Gatun
Stratigraphic resolution:bed
Stratigraphy comments: Upper part of the Gatun Formation. The Gatun Formation is sibdivided into three faunal zones (proposed by Thomson and Keen, 1946): lower, middle, and upper. AGE: Late Miocene, on the basis of microfossil biostratigraphy (see Collins and Coates, 1999), refined by enterer to Tortonian based on more recent dating (Hendy, 2012). Sr dating for nearby sections indicates an age of approximately 8.5 Ma (Hendy, unpublished data).
Lithology and environment
Primary lithology:fine poorly lithified silty sandstone
Includes fossils?Y
Lithology description: LITHOLOGY: Siliciclastic. LITHIFICATION: Poorly-lithified, on the basis of figured specimens and outcrop observations.
Environment:coastal indet.
Geology comments: No specific environmental data presented.
Taphonomy
Modes of preservation:body,original aragonite
Size of fossils:macrofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Museum repositories:USNM
Collection method comments: COLLECTOR: Not stated in text. REPOSITORY: USNM.
Taxonomic list comments:COVERAGE: Exhaustive for gastropoda and bivalvia. NOMENCLATURE: Authoritative publication, with relatively modern nomenclature, and species-resolution identifications.
Metadata
Database number:93053
Authorizer:A. Hendy Enterer:A. Hendy
Modifier:A. Hendy Research group:marine invertebrate
Created:2009-12-24 13:05:06 Last modified:2013-11-25 13:37:23
Access level:the public Released:2009-12-24 13:05:06
Creative Commons license:CC BY
Reference information

Primary reference:
11306. W. P. Woodring. 1957. Geology and Paleontology of Canal Zone and adjoining parts of Panama. United States Geological Survey Professional Paper 306(A) [A. Miller/A. Hendy/A. Hendy]

Secondary references:

43491 B. Landau, R. E. Petit, and C. M. Silva. 2012. New Cancellariidae (Mollusca, Gastropoda) from the Miocene Gatun Formation of Panama, with eleven new species. Journal of Paleontology 86(6):907-930 [A. Hendy/A. Hendy]
43452 B. M. Landau and L. T. Groves. 2011. Cypraeidae (Mollusca: Gastropoda) from the early Miocene Cantaure Formation of northern Venezuela. Novaplex 12(1-2):1-38 [A. Hendy/A. Hendy]
48647 J. A. Todd and K. G. Johnson. 2013. Dissecting a marine snail species radiation (Conoidea: Turridae: Polystira) over 12 million years in the southwestern Caribbean. Bulletin of Marine Science 89(4) [A. Hendy/A. Hendy]
11372 W. P. Woodring. 1959. Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary mollusks (gastropods: Vermetidae to Thaididae). United States Geological Survey Professional Paper 306(B):147-239 [A. Miller/A. Hendy/A. Hendy]
13232 W. P. Woodring. 1964. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Columbellidae to Volutidae). United States Geological Survey Professional Paper 306(C):241-297 [A. Miller/A. Hendy/A. Hendy]
13236 W. P. Woodring. 1970. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). United States Geological Survey Professional Paper 306(D):299-452 [A. Miller/A. Hendy/A. Hendy]
13235 W. P. Woodring. 1973. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (additions to gastropods, scaphopods, pelecypods: Nuculidae to Malleidae). United States Geological Survey Professional Paper 306(E):453-539 [A. Miller/A. Hendy/A. Hendy]
13359 W. P. Woodring. 1982. Geology and paleontology of canal zone and adjoing parts of Panama: Description of Tertiary mollusks (Pelecypods: Propeamussiidae to Cuspidariidae). United States Geological Survey Professional Paper 306(F) [A. Miller/A. Hendy/A. Hendy]