PBDB Collection

Keeline: Harrisonian, Wyoming

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

Mammalia - Carnivora - Amphicyonidae

	Mammacyon sp. Loomis 1936	Hunt 1998
	"Arikaree Formation, N of Keeline"
	= Mammacyon ferocior n. sp. Hunt 2011	Hunt 2011
Mammalia - Carnivora - Mustelidae

	Megalictis ferox Matthew 1907	Hunt and Skolnick 1996
	"North of Keeline (Quarry Z)... Harrison"

	Zodiolestes sp. Riggs 1942	Hunt and Skolnick 1996
Mammalia - Carnivora - Canidae

	Enhydrocyon crassidens Matthew 1907	Wang 1994
	North of Keeline

	Paraenhydrocyon josephi (Cope 1881)	Wang 1994
	North of Keeline

	Paraenhydrocyon wallovianus (Cope 1881)	Wang 1994
	North of Keeline, "Lower or Upper Arikaree Group"

	Leptocyon gregorii (Matthew 1907)	Tedford et al. 2009
	north of Keeline, middle brown sand

	Cormocyon haydeni Wang et al. 1999	Wang et al. 1999
	"north of Keeline"
Mammalia - Camelidae

	Stenomylus keelinensis Frick and Taylor 1968	Frick and Taylor 1968
	N of Keeline; S. gracilis of Frick 1937
Mammalia - Merycoidodontidae

	Phenacocoelus typus Peterson 1907	Schultz and Falkenbach 1950
	Harrison Fm. N of Keeline

	Paramerychyus harrisonensis (Peterson 1907)	Schultz and Falkenbach 1947
	"Z Quarry" N of Keeline

	Merychyus siouxensis Loomis 1924	Schultz and Falkenbach 1947
	synonym of Paramerychyus harrisonensis
	"N. of Keeline" and "8 mi. S.W. of Keeline" in Niobrara Co.

	Merychyus crabilli Schultz and Falkenbach 1947	Schultz and Falkenbach 1947
	synonym of Merychyus calaminthus
	"N. of Keeline" in Niobrara Co.

	Desmatochoerus curvidens (Thorpe 1921)	Schultz and Falkenbach 1954
	as ssp. Desmatochoerus curvidens gregoryi; N of Keeline, Niobrara Co.

	Promerycochoerus carrikeri Peterson 1907	Schultz and Falkenbach 1949
	"N. of Keeline" and "Quarry 'Z,' N. of Keeline"

	Promerycochoerus (Parapromerycochoerus) barbouri n. sp. Schultz and Falkenbach 1949	Schultz and Falkenbach 1949
	synonym of Promerycochoerus carrikeri
	Harrison Fm. N of Keeline
Mammalia - Hypertragulidae

	Nanotragulus ordinatus (Matthew 1907)	Frick 1937
	"Keeline, Wyoming"
Mammalia - Protoceratidae

	Protoceras neatodelpha n. sp. Patton and Taylor 1973	Patton and Taylor 1973
	N of Keeline, Niobrara Co., in Harrison Fm. "equivalent"
Mammalia - Perissodactyla - Tapiridae

	Nexuotapirus marslandensis (Schoch 1984)	Albright 1998
	"Lusk area, north of Keeline, Niobrara County"
Mammalia - Perissodactyla - Rhinocerotidae

	Diceratherium annectens (Marsh 1873)	Prothero 2005

	Diceratherium armatum Marsh 1875	Albright 1999
	"north of Keeline, Niobrara County, Wyoming"
Mammalia - Perissodactyla - Equidae

	Kalobatippus avus (Schlaikjer 1935)	MacFadden 2001
Mammalia - Rodentia - Entoptychidae

	Gregorymys formosus (Matthew 1907)
see common names

Geography

Country:	United States	State/province:	Wyoming	County:	Niobrara
Coordinates:	43.0° North, 104.5° West (view map)
Paleocoordinates:	43.9° North, 99.4° West
Basis of coordinate:	based on political unit
Geographic resolution:	small collection

Time


10 m.y. bin:	Cenozoic 4-5
*Period:	Tertiary	*Epoch:	Miocene
*Local age/stage:	Late/Upper Arikareean
Key time interval:	Harrisonian
Age range of interval:	23.10000 - 18.50000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:	Harrison
Stratigraphy comments: said to be late Arikareean

Lithology and environment

Environment:

terrestrial indet.

Taphonomy

Modes of preservation:

body

Collection methods and comments

Collection excludes:	some genera
Taxonomic list comments:on the Pine Ridge Escarpment according to Schultz and Falkenbach 1950 virtually all Keeline records appear to be from a single F:AM "North of Keeline" locality

Metadata

Also known as:	Quarry Z
Database number:	17794
Authorizer:	J. Alroy, J. Marcot	Enterer:	J. Alroy, J. Marcot
Research group:	vertebrate
Created:	1994-06-07 00:00:00	Last modified:	2002-06-03 03:27:42
Access level:	the public	Released:	1994-06-07 00:00:00
Creative Commons license:	CC BY

Reference information

Primary reference:

3530.

J. H. Wahlert and R. A. Souza. 1988. Skull morphology of Gregorymys and relationships of the Entoptychinae (Rodentia, Geomyidae). American Museum Novitates 2922:1-13 [J. Alroy/J. Alroy/J. Marcot]

Secondary references:

935		L. B. Albright. 1998. New genus of tapir (Mammalia: Tapiridae) from the Arikareean (earliest Miocene) of the Texas Coastal Plain. Journal of Vertebrate Paleontology 18(1):200-217 [J. Alroy/J. Alroy/P. Wagner]
937	5% 600	L. B. Albright. 1999. Ungulates of the Toledo Bend local fauna (late Arikareean, early Miocene), Texas Coastal Plain. Bulletin of the Florida Museum of Natural History, Biological Sciences 42(1):1-80 [J. Alroy/J. Alroy/J. Head]
6294		J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
6222		C. Frick. 1937. Horned ruminants of North America. Bulletin of the American Museum of Natural History 69:1-669 [J. Alroy/J. Alroy/M. Uhen]
1549		C. Frick and B. E. Taylor. 1968. A Generic Review of the Stenomyline Camels. American Museum Novitates 2353:1-51 [J. Alroy/J. Alroy/B. Beatty]
1977		R. M. Hunt, Jr. 1998. Amphicyonidae. 196-227 [J. Alroy/J. Alroy/J. Alroy]
52421		R. M. Hunt, Jr. 2011. Evolution of Large Carnivores During the Mid-Cenozoic of North America: The Temnocyonine Radiation (Mammalia, Amphicyonidae). Bulletin of the American Museum of Natural History 358:1-153 [J. Marcot/J. Marcot]
1981		R. M. Hunt, Jr. and R. Skolnick. 1996. The giant mustelid Megalictis from the Early Miocene carnivore dens at Agate Fossil Beds National Monument, Nebraska: earliest evidence of dimorphism in New World Mustelidae (Carnivora, Mammalia). Contributions to Geology, University of Wyoming 31(1):35-48 [J. Alroy/J. Alroy/M. Uhen]
4197		B. J. MacFadden. 2001. Three-toed browsing horse Anchitherium clarencei from the early Miocene (Hemingfordian) Thomas Farm, Florida. Bulletin of the Florida Museum of Natural History 43(3):79-109 [J. Alroy/J. Alroy/J. Alroy]
2740		T. H. Patton and B. E. Taylor. 1973. The Protoceratinae (Mammalia, Tylopoda, Protoceratidae) and the systematics of the Protoceratidae. Bulletin of the American Museum of Natural History 150(4) [J. Alroy/J. Alroy/J. Ju]
51726		D. R. Prothero. 2005. The Evolution of North American Rhinoceroses. 1-218 [J. Marcot/J. Marcot]
3064		C. B. Schultz and C. H. Falkenbach. 1947. Merychyinae, a subfamily of oreodonts. Bulletin of the American Museum of Natural History 88(4):157-286 [J. Alroy/J. Alroy/J. Marcot]
3065		C. B. Schultz and C. H. Falkenbach. 1949. Promerycochoerinae, a new subfamily of oreodonts. Bulletin of the American Museum of Natural History 93(3):69-198 [J. Alroy/J. Alroy/J. Marcot]
3066		C. B. Schultz and C. H. Falkenbach. 1950. Phenacocoelinae, a new subfamily of oreodonts. Bulletin of the American Museum of Natural History 95(3) [J. Alroy/J. Alroy/J. Ju]
3067		C. B. Schultz and C. H. Falkenbach. 1954. Desmatochoerinae, a new subfamily of oreodonts. Bulletin of the American Museum of Natural History 105(2) [J. Alroy/J. Alroy/J. Ju]
52058		R. H. Tedford, X. Wang, and B. E. Taylor. 2009. Phylogenetic Systematics of the North American Fossil Caninae (Carnivora: Canidae). Bulletin of the American Museum of Natural History 325:1-218 [J. Marcot/J. Marcot/G. Lloyd]
6226		X. Wang. 1994. Phylogenetic systematics of the Hesperocyoninae (Carnivora: Canidae). Bulletin of the American Museum of Natural History 221:1-207 [J. Alroy/J. Alroy/J. Alroy]
3558		X. Wang, R. H. Tedford, and B. E. Taylor. 1999. Phylogenetic systematics of the Borophaginae (Carnivora: Canidae). Bulletin of the American Museum of Natural History 243:1-392 [J. Alroy/J. Alroy/J. Alroy]