Chorlakki: Lutetian, Pakistan

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Osteichthyes - Boidae
Boinae indet. Gray 1825
Rage 1987
Erycinae indet. Bonaparte 1825
Rage 1987
Osteichthyes - Agamidae
Tinosaurus sp. (Estes 1783)
Rage 1987
Osteichthyes - Testudines - Carettochelyidae
de Broin 1987
Osteichthyes - Testudines - Trionychidae
de Broin 1987
Osteichthyes
Gingerich et al. 1983
Osteichthyes - Rodentia - Chapattimyidae
Gingerich et al. 1983
Gingerich et al. 1983
Gingerich et al. 1983
Saykanomys sp. Shevyreva 1972
    = Birbalomys sondaari Hussain et al. 1978
Hartenberger 1982
Birbalomys sondaari (Hussain et al. 1978)
Gingerich et al. 1983
unclassified
Birbalomys ? vandermeuleni Hussain et al. 1978
Gingerich et al. 1983
Osteichthyes - Rodentia - Chapattimyidae
Gingerich et al. 1983
Gingerich et al. 1983
Osteichthyes - Rodentia
cf. Tamquammys sp. Shevyreva 1971
Osteichthyes - Rodentia - Yuomyidae
cf. Petrokoslovia sp. (Shevyreva 1972)
Gingerich et al. 1983
Hartenberger 1982
Osteichthyes - Primates - Omomyidae
Russell and Gingerich 1980
Osteichthyes - Primates - Notharctidae
cf. Agerinia sp. Crusafont-Pairó and Golpe-Posse 1973
Thewissen et al. 1987
Gunnell et al. 2008
specimen referred to Adapidae indet. by Russell and Gingerich (1987) and to Panobius afridi by Thewissen et al. (2001)
Thewissen et al. 1987
Osteichthyes - Plesiosoricidae
Russell and Gingerich 1981
Osteichthyes
Russell and Gingerich 1981
Osteichthyes - Diacodexeidae
Diacodexis pakistanensis Thewissen et al. 1983
Gingerich et al. 1983
Osteichthyes - Cetacea - Pakicetidae
Thewissen and Hussain 1998 7 specimens
Gingerich and Russell 1981 8 specimens
(19 measurements)
Osteichthyes - Raoellidae
Thewissen et al. 1987
Osteichthyes - Dichobunidae
Thewissen et al. 1987
Gingerich et al. 1983
Thewissen et al. 1987
Gingerich et al. 1979
Osteichthyes - Perissodactyla - Brontotheriidae
Gingerich et al. 1983
Osteichthyes - Perissodactyla - Paraceratheriidae
Thewissen et al. 1987
Osteichthyes - Perissodactyla - Isectolophidae
Thewissen et al. 1987
Osteichthyes - Perissodactyla - Anthracobunidae
Gingerich et al. 1983
Gingerich et al. 1983
Osteichthyes - Hyaenodonta
Paratritemnodon indicus Ranga Rao 1973
Gingerich et al. 1983
Osteichthyes - Chiroptera
Chiroptera indet. Blumenbach 1779
Gingerich et al. 1983
Osteichthyes - Cyprinodontiformes - Cyprinodontidae
Gayet 1987
Osteichthyes - Anabantiformes
Roe 1991
Osteichthyes - Centrarchiformes - Percichthyidae
Gayet 1987
Osteichthyes - Cypriniformes - Cyprinidae
Cyprinidae indet. Rafinesque 1815
Gayet 1987
Osteichthyes - Siluriformes - Clariidae
Clariidae indet. Bonaparte 1846
Gayet 1987
Osteichthyes - Siluriformes - Bagridae
Bagridae indet. Bleeker 1852
Gayet 1987
Osteichthyes - Siluriformes - Ariidae
Ariidae indet. Berg 1958
Gayet 1987
unclassified
Gayet 1987
Osteichthyes - Osteoglossiformes - Osteoglossidae
Osteoglossidae indet. Bonaparte 1832
Gayet 1987
Osteichthyes - Lepisosteiformes - Lepisosteidae
Gayet 1987
Gastropoda - Heterostropha - Planorbidae
Planorbis sp. Muller 1774
see common names

Geography
Country:Pakistan State/province:North-West Frontier
Coordinates: 33.6° North, 71.9° East (view map)
Paleocoordinates:11.5° North, 74.2° East (Wright 2013)
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Paleogene Epoch: Eocene
Stage: Lutetian 10 m.y. bin: Cenozoic 2
*Period:Early/Lower - Middle Tertiary - Early/Lower - Middle Tertiary *Epoch:Early/Lower Eocene - Early/Lower Eocene
*International age/stage:Late/Upper Ypresian - Late/Upper Ypresian
Key time interval: Lutetian Foram zone:  P9 - P10
Age range of interval: 48.07 - 41.03 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Kuldana
Stratigraphic resolution:group of beds
Stratigraphy comments: Gingerich (2003) places this collection in the earliest Lutetian; shown by Gunnell et al. (2008) as being close to the top of the formation, spanning at three beds, and falling just over the Early - Middle Eocene boundary; early Lutetian
Lithology and environment
Primary lithology: lithified calcareous conglomerate
Includes fossils?Y
Lithology description: hard, calcareous conglomerate
Environment:fluvial indet.
Taphonomy
Modes of preservation:body,original phosphate
Size of fossils:macrofossils
Preservation of anatomical detail:good
Associated major elements:some
Disassociated major elements:some
Collection methods and comments
Collection methods:bulk,selective quarrying,surface (in situ),chemical,mechanical,sieve
Rock censused:200 kg
Reason for describing collection:general faunal/floral analysis
Metadata
Database number:28978
Authorizer:M. Uhen, W. Clyde, J. Alroy Enterer:M. Uhen, W. Clyde, J. Alroy
Modifier:P. Wagner Research group:vertebrate
Created:2003-03-08 14:03:39 Last modified:2025-11-06 14:14:50
Access level:the public Released:2003-03-08 14:03:39
Creative Commons license:CC0
Reference information

Primary reference:
12009. P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell, J.-L. Hartenberger, S. M. Ibrahim Shah, M. Hassan, K. D. Rose and R. H. Ardrey. 1979. Reconnaissance survey and vertebrate paleontology of some Paleocene and Eocene Formations in Pakistan. Contributions from the Museum of Paleontology, University of Michigan 25(5):105-116 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

14333 F. de Broin. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Chelonia. Contributions from the Museum of Paleontology, University of Michigan 27(7):169-185 [W. Clyde/W. Clyde/W. Clyde]
14329 M. Gayet. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Holostei and Teleostei. Contributions from the Museum of Paleontology, University of Michigan 27(7):151-168 [W. Clyde/W. Clyde/W. Clyde]
15283 P. D. Gingerich. 2003. Stratigraphic and micropaleontological constraints on the Middle Eocene age of the mammal-bearing Kuldana Formation of Pakistan. Journal of Vertebrate Paleontology 23(3):643-651 [M. Uhen/M. Uhen]
13810 P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell and J.-L. Hartenberger. 1979. Chorlakkia hassani, a new middle Eocene dichobunid (Mammalia, Artiodactyla) from the Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(6):117-124 [W. Clyde/W. Clyde/J. Alroy]
8326 P. D. Gingerich and D. E. Russell. 1981. Pakicetus inachus, a new archaeocete (Mammalia, Cetacea) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(11):235-246 [M. Uhen/M. Uhen/M. Uhen]
12019 P. D. Gingerich, N. A. Wells, D. E. Russell and S. M. I. Shah. 1983. Origin of whales in epicontinental remnant seas: New evidence from the early Eocene of Pakistan. Science 220:403-406 [M. Uhen/M. Uhen/M. Uhen]
29569 G. F. Gunnell, P. D. Gingerich, M. Haq, J. I. Bloch, I. H. Khan and W. C. Clyde. 2008. New primates (Mammalia) from the early and middle Eocene of Pakistan and their paleobiogeographical implications. Contributions from the Museum of Paleontology, University of Michigan 32(1):1-14 [J. Alroy/J. Alroy]
13077 J.L. Hartenberger. 1982. A review of the Eocene rodents of Pakistan. Contributions from the Museum of Paleontology, University of Michigan 26:19-35 [W. Clyde/L. Gannon/J. Alroy]
14334 J. C. Rage. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Squamata. Contributions from the Museum of Paleontology, University of Michigan 27(7):187-193 [W. Clyde/W. Clyde/W. Clyde]
13802 L. J. Roe. 1991. Phylogenetic and ecological significance of Channidae (Osteichthyes, Teleostei) from the early Eocene Kuldana Formation of Kohat, Pakistan. Contributions from the Museum of Paleontology, University of Michigan 28(5):93-100 [W. Clyde/W. Clyde/W. Clyde]
29559 D. E. Russell and P. D. Gingerich. 1980. Un nouveau Primate omomyide dans l'Eocène du Pakistan. Comptes Rendus de l'Académie de Paris, Paris, Série D 291:621-624 [J. Alroy/J. Alroy]
13809 D. E. Russell and P. D. Gingerich. 1981. Lipotyphla, Proteutheria(?), and Chiroptera (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(14):277-287 [W. Clyde/W. Clyde/W. Clyde]
59921 D. E. Russell and P. D. Gingerich. 1987. Nouveaux primates de l’Éocène du Pakistan. Comptes Rendus de l'Academie des Sciences, Paris, Série II 5:209-214 [P. Mannion/P. Mannion]
14336 J. G. M. Thewissen, P. D. Gingerich, and D. E. Russell. 1987. Artiodactyla and Perissodactyla (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 27(11):247-274 [W. Clyde/W. Clyde/M. Uhen]
11990 J. G. M. Thewissen and S. T. Hussain. 1998. Systematic review of the Pakicetidae, early and middle Eocene Cetacea (Mammalia) from Pakistan and India. Bulletin of Carnegie Museum of Natural History 34:220-238 [M. Uhen/M. Uhen/M. Uhen]