Akespe (Aral Horizon) (Miocene of Kazakhstan)

Also known as Agyspe;

Where: Kazakhstan (50.2° N, 66.8° E: paleocoordinates 50.8° N, 66.0° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: NN5 mammal zone, Aral Formation, Miocene (23.0 - 5.3 Ma)

• late Oligocene

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

• grayish yellow-green calcareous shale and claystone

Size classes: macrofossils, mesofossils

Collected by N.S. Shevyreva in 1968

Primary reference: D.V. Malakhov. 2003. The earliest known record of Mioproteus (Caudata; Proteidae) from the Middle Miocene of Central Kazakhstan. Biota 4:67-72 [P. Coster/P. Coster]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 200980: authorized by Pauline Coster, entered by Pauline Coster on 28.04.2019

Creative Commons license: CC BY (attribution)

Taxonomic list

Osteichthyes
 Caudata - Urodela
Mioproteus cf. caucasicus Estes and Darevsky 1978 mudpuppy
Mammalia
 Rodentia - Cricetidae
Eumyarion sp. Thaler 1966 mouse
 Rodentia - Aplodontidae
Ansomys sp. Qiu 1987 mountain beaver
 Rodentia - Castoridae
Monosaulax sp. Stirton 1935 beaver
 Lagomorpha - Ochotonidae
Ochotonidae sp. Thomas 1897 pika
 Tribosphenida - Soricidae
Soricidae sp. Fischer 1814 shrew