CBI-117 (Shark Locality), Dzharakuduk (Cretaceous to of Uzbekistan)

Also known as CDZH-117 [in error], Aitym fauna

Where: Navoi, Uzbekistan (42.1° N, 62.7° E: paleocoordinates 36.8° N, 57.0° E)

• coordinate stated in text

• small collection-level geographic resolution

When: Aitym Formation, Late/Upper Turonian to Late/Upper Turonian (93.5 - 86.3 Ma)

• It is confined to the marine Aitym Formation, laying above the famous fluvially dominated Bissekty Formation that has produced a diverse and important fauna of late Turonian-Coniacian vertebrates

• bed-level stratigraphic resolution

Environment/lithology: marine; lithology not reported

• the marine Aitym Formation

Size classes: macrofossils, mesofossils

Collected by Joint Uzbekian-Russian-British-American-Canadian (URBAC-98) in 1998

• Collection of the Zoological Institute, Russian Academy of Sciences, Saint Petersburg (abbreviated ZIN)

Primary reference: A. Averianov. 1999. A new species of multituberculate mammal. Uzbekbaatar from the Late Cretaceous of Uzbekistan. Acta Palaeontologica Polonica 44(3):301-304 [R. Benson/R. Benson]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 123485: authorized by Roger Benson, entered by Roger Benson on 24.01.2012, edited by Jonathan Tennant, Matthew Carrano and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• The Aitym fauna consists of varied, mainly marine vertebrates (e.g., diverse and abundant chondrichthyans and more rare osteichthyans, infrequent plesiosaurs, sea(?) turtles), and of some marine invertebrates, including oysters, belemnites, and ammonites (Placenticeras kyzylkumense). The remains of terrestrial vertebrates are extremely rare in this site. There are few bones of salamanders and lizards, teeth of crocodiles, shell fragments of possibly terrestrial turtles
Show authors, comments, and common names
Cephalopoda
 Ammonitida - Placenticeratidae
Placenticeras kyzylkumense2
Mammalia
 Theriamorpha - Spalacotheriidae
cf. Shalbaatar sp.2
cf. Shalbaatar sp.2 Nessov 1997 mammal
ZIN C.85047, left edentulous dentary fragment with alveoli of the three last molars (m5-7?)
 Deltatheroida - Deltatheridiidae
Deltatheridiidae indet.1
Deltatheridiidae indet.1 Gregory and Simpson 1926 metatherian
ZIN C.85049, fragment of a right upper molar (M2?), lacking lingual and posterior portions
 Theriamorpha -
Eutheria indet.2
Eutheria indet.2 Huxley 1880 eutherian
ZIN C.85295, ZIN C.85297, ZIN C.85298, ZIN C.85302
 Theriamorpha - Kennalestidae
Eoungulatum cf. kudukensis2
Eoungulatum cf. kudukensis2 Nessov et al. 1998 eutherian
ZIN O.35052, badly worn left m3; ZIN C.85054, left dentary fragment with alveolus of m3
 Theriamorpha - Zhelestidae
Zhelestes temirkayzk1, "Parazhelestes aff. minor" = Parazhelestes mynbulakensis2, Parazhelestes robustus1, Aspanlestes aptap1
Zhelestes temirkayzk1 eutherian
edentulous left dentary fragment with the alveolus for m3
"Parazhelestes aff. minor" = Parazhelestes mynbulakensis2 Nessov 1985 eutherian
ZIN O.35054, right M3
Parazhelestes robustus1 Nessov 1993 eutherian
ZIN C.85055, badly worn left(?) M1 or 2; ZIN C.85301, left M3
Aspanlestes aptap1 Nessov 1985 eutherian
ZIN C.85051, left m1 or m2 lacking posterior portion of the talonid; ZIN C.85048, left edentulous dentary fragment with alveoli of m2-3 and posterior root of m1; URBAC 02-101, right M2 lacking the parastylar lobe. Possibly also ZIN C.85299, a right P4? (penultimate premolar) and ZIN C.85079, left edentulous maxillary fragment with M1(?) alveolus
 Theriamorpha - Zalambdalestidae
aff. Kulbeckia sp.2, Kulbeckia kulbecke1
aff. Kulbeckia sp.2 Nessov 1993 eutherian
ZIN C.85303, upper right canine
Kulbeckia kulbecke1 Nessov 1993 eutherian
ZIN C.85045, right petrosal; ZIN C.85050, worn left m1
 Theriamorpha -
Paranyctoides sp.3, Paranyctoides sp.1, "Paranyctoides aralensis" = Paranyctoides quadrans1
Paranyctoides sp.3 Fox 1979 eutherian
ZIN C.85044, left M1; ZIN. C.85300, right M1(?) fragment preserving the crown from the conules lingually; ZIN C.85078, left dentary with root for the posteriormost incisor and one-root for a canine (or just as likely two roots for a two-rooted canine), two roots for p1, a complete p2, and two alveoli for a smaller p3; ZIN C.85294, right m1
Paranyctoides sp.1 Fox 1979 eutherian
ZIN C.85296, a worn right M1 lacking the protocone
"Paranyctoides aralensis" = Paranyctoides quadrans1 Nessov 1982 eutherian
ZIN C.85293, left M1
 Multituberculata -
Uzbekbaatar sp.1, Uzbekbaatar wardi n. sp.
Uzbekbaatar sp.1 Kielan-Jaworowska and Nessov 1992 multituberculate
Uzbekbaatar wardi n. sp. Averianov 1999 multituberculate
ZIN 83673, holotype left p4; and material listed by Averianov & Archibald (2003): ZIN C.85292, dP1?; ZIN C.85077, posterior half of right M1; ZIN C.85291, right m1.
Reptilia
 Saurischia -
Titanosauria indet.4
Titanosauria indet.4 Bonaparte and Coria 1993 sauropod
teeth
1Archibald and Averianov 2005; 2Averianov and Archibald 2003; 3Averianov and Archibald 2013; 4Averianov and Sues 2017