Where: Castilla-La Mancha, Spain (40.1° N, 1.9° W: paleocoordinates 30.5° N, 8.1° E)
• coordinate estimated from map
• outcrop-level geographic resolution
When: La Huérguina Formation, Late/Upper Barremian (130.0 - 125.5 Ma)
• also called "Calizas de La Huérgina Formation" or "La Huérgina Limestones Formations"
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine - large; limestone and packstone
Size classes: macrofossils, mesofossils
Preservation: adpression, trace, soft parts
Collection methods: quarrying, surface (in situ),
• Repository: Museo de las Ciencias de Castilla-La Mancha, Cuenca (MCCM)
Primary reference: J. L. Sanz, S. Wenz, A. Yebenes, R. Estes, X. Martinez-Delclos, E. Jiminez-Fuentes, C. Dieguez, A. D. Buscalioni, L. J. Barbadillo and L. Via. 1988. An Early Cretaceous faunal and floral continental assemblage: Las Hoyas fossil site (Cuenca, Spain). Géobios 21(5):611-635 [J. Alroy/E. Leckey/J. Alroy]more details
Purpose of describing collection: general faunal/floral analysis
PaleoDB collection 28006: authorized by John Alroy, entered by Erin Leckey on 20.01.2003, edited by Roger Benson, Matthew Carrano, John Alroy, Richard Butler, Matthew Oreska, Jered Karr, Matthew Clapham, Philip Mannion, Jonathan Tennant and Mark Bell
Creative Commons license: CC BY (attribution)
Taxonomic list
unclassified | |
Polypodiopsida | |
Weichselia reticulata32 Stokes and Webb 1824 | |
Ruffordia goepperti32 Dunker 1844 | |
Cladophlebis browniana Dunker 1846 | |
Equisetopsida | |
Pagiophyllum sp.32 Heer 1881 | |
Pinopsida | |
Sphenolepis kurriana Dunker 1846 | |
Zamites sp. Brongniart 1828 | |
Angiospermae | |
Montsechia vidali Zeiller 1902 | |
Actinopteri | |
"? Ascalabos voithi" = Leptolepis
"? Ascalabos voithi" = Leptolepis | |
Rubiesichthys gregalis Wenz 1984 | |
Gordichthys conquensis n. gen. n. sp.22
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Pleuropholis sp. Egerton 1858 | |
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Eomesodon sp.39 Woodward 1918 | |
Macromesodon sp.39 Blake 1905 | |
Lepidotes sp. Agassiz 1832 gar | |
Notagogus sp. Agassiz 1833 | |
"? Amiopsis sp." = Amiopsis Kner 1863 bowfin
"Urocles woodwardi" = Amiopsis bowfin
Vidalamia catalunica Sauvage 1903 bowfin | |
Caturus sp. Agassiz 1834 bowfin | |
Coelacanthimorpha | |
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Amphibia | |
Celtedens ibericus n. sp.14
Celtedens ibericus n. sp.14 McGowan and Evans 1995 tetrapod LH 6020 - type (part and counterpart of a nearly complete skeleton, missing only part of the palate and tail); referred material (McGowan 2002): two complete specimens in part and counterpart (LH 030 R A & B; LH 15710 A & B)
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Gracilibatrachus avallei n. gen. n. sp.1 Baez 2013 frog MCCM-LH 21171a, b - holotype (articulated skeleton of a postmetamorphic individual); referred specimens: MCCMLH-17422a, b, 11386, 11387, 26251, 26803a, b (in part), and, tentatively, 11394a, b
Iberobatrachus angelae n. gen. n. sp.1 Baez 2013 frog MCCM-LH 11393a-b - holotype (articulated specimen lacking the hindlimbs and left forelimb, two-dimensionally preserved as part and counterpart on two slabs); referred specimen - MCCM-LH 15999a-b (incomplete, articulated specimen preserved as part and counterpart on two slabs)
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Valdotriton gracilis n. gen. n. sp.10
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Mammalia | |
Spinolestes xenarthrosus n. gen. n. sp.13
Spinolestes xenarthrosus n. gen. n. sp.13 Martin et al. 2015 mammal MCCM-LH 30000 - holotype (complete skeleton); MCCMLH30000B (counter slab)
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Reptilia | |
Hoyalacerta sanzi n. gen. n. sp.9, Jucaraseps grandipes n. gen. n. sp.2, Scandensia ciervensis n. gen. n. sp.8
Jucaraseps grandipes n. gen. n. sp.2 Bolet and Evans 2012 squamates LH 18505 (holotype), almost complete skeleton in dorsal view
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Meyasaurus diazromerali n. sp.7
Meyasaurus diazromerali n. sp.7 Evans and Barbadillo 1997 squamates LH 370, LH 372, LH 33, LH 13510, LH 6026, LH 14317
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Hoyasemys jimenezi n. gen. n. sp.19
Hoyasemys jimenezi n. gen. n. sp.19 Pérez-García et al. 2012 turtle MCCM-LH 84, LH 254 (carapace fragment)
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Europejara olcadesorum n. gen. n. sp.37
Europejara olcadesorum n. gen. n. sp.37 Vullo et al. 2012 pterosaur MCCM-LH 9413, jaws and post-orbital elements of the skull preserved on part and counterpart slabs
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Concavenator corcovatus n. gen. n. sp.17
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Pelecanimimus polyodon n. gen. n. sp.20
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Aves | |
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cf. Iberomesornis sp.27, Iberomesornis romerali n. gen. n. sp.28, Eoalulavis hoyasi n. gen. n. sp.30, Concornis lacustris n. gen. n. sp.29, cf. Concornis lacustris4
cf. Concornis lacustris4 Sanz and Buscalioni 1992 bird MCCM-LH 21006 a-b, tibiotarsus and tarsometatarsus
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Reptilia | |
cf. Mantellisaurus sp.15 Paul 2007 ornithopod LH-30677, incomplete but fully-articulated right hindlimb
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"Bernissartidae indet." = Bernissartiidae32
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Atoposauridae indet. crocodilian | |
Cassissuchus sanziuami n. gen. n. sp.3
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Undichna | |
Undichna unisulca15 de Gibert et al. 1999 | |
Bivalvia | |
Unio sp.32
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Insecta | |
"Blattoidea indet." = Blattodea
"Blattoidea indet." = Blattodea Latreille 1810 cockroach | |
Chresmoda aquatica6 Martínez-Delclòs 1989 winged insect LH-26.546, LH-13826, LH-13574, LH-92AM102, MCCM-LH 18023, MCCM-LH 18024,
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Mesoraphidiidae indet. Martynov 1925 snakefly | |
Nanochrysopa pumilio n. gen. n. sp.16, "Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16, Triangulochrysopa sanzi n. gen. n. sp.16, Chimerochrysopa incerta n. gen. n. sp.16, Allopterus mayorgai n. sp.16, Tachinymphes penalveri n. sp.16, Tachinymphes paicheleri n. sp.16
"Armandochrysopa inexpecta n. sp." = Kareninoides inexpectus16 Nel et al. 2005 lacewing MCCM LH 18575
Triangulochrysopa sanzi n. gen. n. sp.16 Nel et al. 2005 lacewing MCCM LH-8110, 18572, 18573, 18574, MCCM 92/2/3
Tachinymphes penalveri n. sp.16 Nel et al. 2005 lacewing MCCM LH 18585, LH 18586-7, U.A. Madrid LH 8033, 8094
Tachinymphes paicheleri n. sp.16 Nel et al. 2005 lacewing MCCM LH 18576-8, LH 8040, LH 13175, MCCM LH 18579-18584
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Ovonectes pilosum n. gen. n. sp.35, Hispanoclavina gratshevi n. sp.35, Hispanoclavina diazromerali n. gen. n. sp.35, Megacoptoclava longiurogomphia n. gen. n. sp.21, Hoyaclava buscalionae n. gen. n. sp.35
Hispanoclavina diazromerali n. gen. n. sp.35 Soriano et al. 2007 beetle MCCM-LH16264, 15070, 16315, 22299, 23859
Megacoptoclava longiurogomphia n. gen. n. sp.21 Ponomarenko and Martínez-Delclòs 2000 beetle MCCM LH-1263
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Ademosynidae indet. Ponomarenko 1968 beetle | |
Distenorrhinus (Parabrenthorrhinus) xavieri n. sp.40 Zherikhin and Gratshev 2003 pine flower snout beetle MCCM LH 17330
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Martynopsis laticollis n. gen. n. sp.34
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Archostemata indet., "Zygadenia siniestri n. sp." = Conexicoxa siniestri33, "Zygadenia martinclosas" = Conexicoxa martinclosas33
Archostemata indet. Kolbe 1908 beetle
"Zygadenia siniestri n. sp." = Conexicoxa siniestri33 Soriano and Delclos 2006 beetle MCCM LH 23515, MCCM LH 21291, MCCM LH 15819
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Tetraphalerus ponomarenkoi n. sp.33
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"Anaglyphites zherikhini n. sp." = Mesocupes zherikhini33 Soriano and Delclos 2006 reticulated beetle MCCM LH 15122
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"Zygadenia viridis n. sp." = Brachilatus viridis33, "Zygadenia longicoxa n. sp." = Brachilatus longicoxa33, "Zygadenia diazromerali n. sp." = Notocupes diazromerali33
"Zygadenia longicoxa n. sp." = Brachilatus longicoxa33 Soriano and Delclos 2006 beetle MCCM LH 928030, MCCM LH 23432
"Zygadenia diazromerali n. sp." = Notocupes diazromerali33 Soriano and Delclos 2006 beetle MCCM LH 12001
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"Orthophlebiinae indet." = Orthophlebiidae
"Orthophlebiinae indet." = Orthophlebiidae Handlirsch 1906 scorpionfly | |
Heteroptera indet. Latreille 1810 true bug | |
Naucoridae indet. Fallen 1814 creeping water bug | |
Velocorixinae indet. Popov 1986 water boatman | |
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Notonectidae indet. Latreille 1802 true bug | |
Sinaeschnidia martinezdelclosi n. sp.11, Iberoaeschnidium conquensis n. gen. n. sp.11, Gigantoaeschnidium ibericus n. gen. n. sp.11
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Ephemeroptera indet. Hyatt and Arms 1890 mayfly | |
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Malacostraca | |
"Oplophorus roselli" = Delclosia roselli Via 1971 shrimp | |
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Arachnida | |
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Planolites sp.15 Nicholson 1873
Palaeophycus sp.15 Hall 1847
Cruziana sp.15 d'Orbigny 1842
Lockeia sp.15 James 1879
Treptichnus sp.15 Miller 1889 | |
Helminthoidichnites | |
Helminthoidichnites sp.15 Fitch 1850 |