PBDB Collection

MAD93-35 (SUNY) (Cretaceous of Madagascar)

Where: Mahajanga, Madagascar (15.9° S, 46.6° E: paleocoordinates 30.1° S, 38.3° E)

• coordinate stated in text

• small collection-level geographic resolution

When: Anembalemba Member (Maevarano Formation), Maastrichtian (72.1 - 66.0 Ma)

• "uppermost white sandstone unit" = Anembalemba Member

• bed-level stratigraphic resolution

Environment/lithology: "channel"; white sandstone

• "shallow and very broad channel-belt system characterized by an ephemeral and flood-prone discharge regime."

Size class: mesofossils

Preservation: trace

Reposited in the DMNH, FMNH

Collection methods: quarrying, mechanical,

Primary reference: D. W. Krause, G. V. R. Prasad, W. Koenigswald, A. Sahni, and F. E. Grine. 1997. Cosmopolitanism among Gondwanan Late Cretaceous mammals. Nature 390:504-507 [M. Carrano/M. Carrano/P. Holroyd]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 39266: authorized by Matthew Carrano, entered by Matthew Carrano on 25.05.2004, edited by Philip Mannion, Roger Benson, Jonathan Tennant and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Mammalia
Theriamorpha - Zhelestidae	Zhelestidae indet.² Zhelestidae indet.² Nessov 1990 eutherian
Allotheria -	? Multituberculata indet.⁷ ? Multituberculata indet.⁷ Cope 1884 multituberculate FMNH PM 61152 (fragmentary molar)
Multituberculata -	Multituberculata indet.⁸ Multituberculata indet.⁸ Cope 1884 multituberculate DMNH EPV.100010 (fragmentary femur)
Allotheria - Sudamericidae	Sudamericidae indet.⁷, Lavanify miolaka Sudamericidae indet.⁷ Scillato-Yané and Pascual 1984 mammal UA 9937 (fragmentary molariform cheek tooth) Lavanify miolaka Krause et al. 1997 mammal
Reptilia
Loricata - Mahajangasuchidae	Mahajangasuchus insignis n. gen. n. sp.¹⁴ Mahajangasuchus insignis n. gen. n. sp.¹⁴ Buckley and Brochu 1999 Mahajanga crocodile UA 8654; FMNH PR 2389, 2449
Loricata -	Simosuchus clarki⁹ Simosuchus clarki⁹ Buckley et al. 2000 pug-nosed crocodile FMNH PR 2600 (isolated tooth)
Dinosauria -	Dinosauria indet.¹³ Dinosauria indet.¹³ dinosaur
Aves
Enantiornithes -	Enantiornithes indet.¹¹ Enantiornithes indet.¹¹ Walker 1981 bird FMNH PA 751
Reptilia
Theropoda - Noasauridae	Masiakasaurus knopfleri³ Masiakasaurus knopfleri³ Sampson et al. 2001 ceratosaur FMNH PR 2220, 2199; UA 9183, 9734, 9773, 9199, 9185, 9137
Serpentes -	Serpentes indet.¹⁰ Serpentes indet.¹⁰ snake FMNH PR 2579
Squamata - Nigerophiidae	Kelyophis hechti¹⁰ Kelyophis hechti¹⁰ LaDuke et al. 2010 snake
Squamata - Madtsoiidae	Adinophis fisaka¹², Menarana nosymena¹⁰ Adinophis fisaka¹² Pritchard et al. 2014 snake FMNH PR 2572 Menarana nosymena¹⁰ LaDuke et al. 2010 snake
Testudines - Sahonachelyidae	Sokatra antitra n. gen. n. sp.⁶ Sokatra antitra n. gen. n. sp.⁶ Gaffney and Krause 2011 sideneck turtle UA 9769 - holotype (partial skull, lacking the premaxillae, basioccipital, and with damaged basisphenoid, quadrates, prootics, and opisthotics). FMNH nos. PR 2647 (supraoccipital), PR 2648 (left frontal), PR 2649 (partial skull with articulated or associated quadrates, both opisthotics, left exoccipital, right prootic, left pterygoid, and right palatine)and UA nos. 9864 (partial right quadrate) and 9865 (partial left maxilla) were recovered from a small (2 m2) quarried surface at this locality
Testudines - Podocnemididae	cf. Erymnochelys sp.⁵ cf. Erymnochelys sp.⁵ Baur 1888 sideneck turtle FMNH PR 2231, mandible
Amphibia
Salientia -	Anura indet.¹, Beelzebufo ampinga⁴ Anura indet.¹ Fischer von Waldheim 1813 frog FMNH-PR 1960 (right premaxilla) Beelzebufo ampinga⁴ Evans et al. 2008 frog UA 9623 – fragment of otic and ventral processes of squamosal; UA 9624 – posterior process of left quadratojugal; UA 9625 – posteroventral process of right squamosal; UA 9626 – cranial fragment; UA 9627 – partial vertebral spine table; UA 9635 – anterior fragment of right maxilla; UA 9676 – fragment of right maxilla bearing maxillary nerve canal; UA 9677 – fragment of frontoparietal or squamosal; UA 9679 – cranial fragment; UA 9947 – presacral vertebra, interpreted as PS3; UA 9948 – posterior presacral vertebra; FMNH PR 1960 – partial right premaxilla; FMNH PR 2504 – vertebral centrum and partial neural arch; FMNH PR 2505 – facial process of right maxilla; FMNH PR 2506 – partial right maxilla; FMNH PR 2507 – facial process and pars dentalis of right maxilla; FMNH PR 2508 – cranial fragment from antorbital margin, frontoparietal or nasal; FMNH PR 2509 – conjoined midline frontoparietal fragment; FMNH PR 2510 – partial left maxilla. Locality MAD93-36: UA 9678 – vertebral spine table; UA 9949 – fragment of otic plate of right squamosal
Cubiculum ornatus
-	Cubiculum ornatus n. gen. n. sp.¹³ Cubiculum ornatus n. gen. n. sp.¹³ Roberts et al. 2007

¹Asher and Krause 1998; ²Averianov et al. 2003; ³Carrano et al. 2011; ⁴Evans et al. 2014; ⁵Gaffney and Forster 2003; ⁶Gaffney and Krause 2011; ⁷Krause 2013; ⁸Krause et al. 2017; ⁹Krause et al. 2010; ¹⁰LaDuke et al. 2010; ¹¹O'Connor and Forster 2010; ¹²Pritchard et al. 2014; ¹³Roberts et al. 2007; ¹⁴Turner and Buckley 2008