Chorlakki: Lutetian, Pakistan

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda - Heterostropha - Planorbidae
Planorbis sp. Muller 1774
Actinopteri - Lepisosteiformes - Lepisosteidae
Gayet 1987
Actinopteri - Siluriformes - Clariidae
Clariidae indet. Bonaparte 1846
Gayet 1987
Actinopteri - Siluriformes - Ariidae
Ariidae indet. Berg 1958
Gayet 1987
Actinopteri - Siluriformes - Bagridae
Bagridae indet. Bleeker 1852
Gayet 1987
Actinopteri - Cypriniformes - Cyprinidae
Cyprinidae indet. Rafinesque 1815
Gayet 1987
unclassified
Birbalomys ? vandermeuleni Hussain et al. 1978
Gingerich et al. 1983
Gayet 1987
Actinopteri - Osteoglossiformes - Osteoglossidae
Osteoglossidae indet. Bonaparte 1832
Gayet 1987
Actinopteri - Percichthyidae
Gayet 1987
Actinopteri - Anabantiformes
Roe 1991
Actinopteri - Cyprinodontiformes - Cyprinodontidae
Gayet 1987
Reptilia - Testudines - Carettochelyidae
de Broin 1987
Reptilia - Testudines - Trionychidae
de Broin 1987
Reptilia - Agamidae
Tinosaurus sp. (Estes 1783)
Rage 1987
Reptilia - Boidae
Erycinae indet. Bonaparte 1825
Rage 1987
Boinae indet. Gray 1825
Rage 1987
Mammalia
Gingerich et al. 1983
Mammalia - Rodentia
cf. Tamquammys sp. Shevyreva 1971
Mammalia - Rodentia - Chapattimyidae
Gingerich et al. 1983
Gingerich et al. 1983
Saykanomys sp. Shevyreva 1972
    = Birbalomys sondaari Hussain et al. 1978
Hartenberger 1982
Birbalomys sondaari (Hussain et al. 1978)
Gingerich et al. 1983
Gingerich et al. 1983
Gingerich et al. 1983
Gingerich et al. 1983
Mammalia - Rodentia - Yuomyidae
cf. Petrokoslovia sp. (Shevyreva 1972)
Gingerich et al. 1983
Hartenberger 1982
Mammalia - Primates - Notharctidae
cf. Agerinia sp. Crusafont-Pairó and Golpe-Posse 1973
Thewissen et al. 1987
Gunnell et al. 2008
specimen referred to Adapidae indet. by Russell and Gingerich (1987) and to Panobius afridi by Thewissen et al. (2001)
Thewissen et al. 1987
Mammalia - Primates - Omomyidae
Russell and Gingerich 1980
Mammalia - Plesiosoricidae
Russell and Gingerich 1981
Mammalia
Russell and Gingerich 1981
Mammalia - Hyaenodonta
Paratritemnodon indicus Ranga Rao 1973
Gingerich et al. 1983
Mammalia - Perissodactyla - Paraceratheriidae
Thewissen et al. 1987
Mammalia - Perissodactyla - Isectolophidae
Thewissen et al. 1987
Mammalia - Perissodactyla - Anthracobunidae
Gingerich et al. 1983
Gingerich et al. 1983
Mammalia - Perissodactyla - Brontotheriidae
Gingerich et al. 1983
Mammalia - Diacodexeidae
Diacodexis pakistanensis Thewissen et al. 1983
Gingerich et al. 1983
Mammalia - Cetacea - Pakicetidae
Gingerich and Russell 1981 8 specimens
(19 measurements)
Thewissen and Hussain 1998 7 specimens
Mammalia - Dichobunidae
Thewissen et al. 1987
Thewissen et al. 1987
Gingerich et al. 1979
Gingerich et al. 1983
Mammalia - Raoellidae
Thewissen et al. 1987
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
Gingerich et al. 1983
see common names

Geography
Country:Pakistan State/province:North-West Frontier
Coordinates: 33.6° North, 71.9° East (view map)
Paleocoordinates:11.5° North, 74.2° East (Wright 2013)
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period: Paleogene Epoch: Eocene
Stage: Lutetian 10 m.y. bin: Cenozoic 2
*Period:Early/Lower - Middle Tertiary - Early/Lower - Middle Tertiary *Epoch:Early/Lower Eocene - Early/Lower Eocene
*International age/stage:Late/Upper Ypresian - Late/Upper Ypresian
Key time interval: Lutetian
Age range of interval: 48.07 - 41.03 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Kuldana
Stratigraphic resolution:group of beds
Stratigraphy comments: Gingerich (2003) places this collection in the earliest Lutetian
shown by Gunnell et al. (2008) as being close to the top of the formation, spanning at three beds, and falling just over the Early - Middle Eocene boundary; early Lutetian
Lithology and environment
Primary lithology: lithified calcareous conglomerate
Includes fossils?Y
Lithology description: hard, calcareous conglomerate
Environment:fluvial indet.
Taphonomy
Modes of preservation:body,original phosphate
Size of fossils:macrofossils
Preservation of anatomical detail:good
Associated major elements:some
Disassociated major elements:some
Collection methods and comments
Collection methods:bulk,selective quarrying,surface (in situ),chemical,mechanical,sieve
Rock censused:200 kg
Reason for describing collection:general faunal/floral analysis
Metadata
Database number:28978
Authorizer:M. Uhen, J. Alroy, W. Clyde Enterer:M. Uhen, W. Clyde, J. Alroy
Modifier:P. Mannion Research group:vertebrate
Created:2003-03-08 14:03:39 Last modified:2025-02-22 15:12:02
Access level:the public Released:2003-03-08 14:03:39
Creative Commons license:CC0
Reference information

Primary reference:
12009. P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell, J.-L. Hartenberger, S. M. Ibrahim Shah, M. Hassan, K. D. Rose and R. H. Ardrey. 1979. Reconnaissance survey and vertebrate paleontology of some Paleocene and Eocene Formations in Pakistan. Contributions from the Museum of Paleontology, University of Michigan 25(5):105-116 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

14333 F. de Broin. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Chelonia. Contributions from the Museum of Paleontology, University of Michigan 27(7):169-185 [W. Clyde/W. Clyde/W. Clyde]
14329 M. Gayet. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Holostei and Teleostei. Contributions from the Museum of Paleontology, University of Michigan 27(7):151-168 [W. Clyde/W. Clyde/W. Clyde]
15283 P. D. Gingerich. 2003. Stratigraphic and micropaleontological constraints on the Middle Eocene age of the mammal-bearing Kuldana Formation of Pakistan. Journal of Vertebrate Paleontology 23(3):643-651 [M. Uhen/M. Uhen]
13810 P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell and J.-L. Hartenberger. 1979. Chorlakkia hassani, a new middle Eocene dichobunid (Mammalia, Artiodactyla) from the Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(6):117-124 [W. Clyde/W. Clyde/J. Alroy]
8326 P. D. Gingerich and D. E. Russell. 1981. Pakicetus inachus, a new archaeocete (Mammalia, Cetacea) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(11):235-246 [M. Uhen/M. Uhen/M. Uhen]
12019 P. D. Gingerich, N. A. Wells, D. E. Russell and S. M. I. Shah. 1983. Origin of whales in epicontinental remnant seas: New evidence from the early Eocene of Pakistan. Science 220:403-406 [M. Uhen/M. Uhen/M. Uhen]
29569 G. F. Gunnell, P. D. Gingerich, M. Haq, J. I. Bloch, I. H. Khan and W. C. Clyde. 2008. New primates (Mammalia) from the early and middle Eocene of Pakistan and their paleobiogeographical implications. Contributions from the Museum of Paleontology, University of Michigan 32(1):1-14 [J. Alroy/J. Alroy]
13077 J.L. Hartenberger. 1982. A review of the Eocene rodents of Pakistan. Contributions from the Museum of Paleontology, University of Michigan 26:19-35 [W. Clyde/L. Gannon/J. Alroy]
14334 J. C. Rage. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Squamata. Contributions from the Museum of Paleontology, University of Michigan 27(7):187-193 [W. Clyde/W. Clyde/W. Clyde]
13802 L. J. Roe. 1991. Phylogenetic and ecological significance of Channidae (Osteichthyes, Teleostei) from the early Eocene Kuldana Formation of Kohat, Pakistan. Contributions from the Museum of Paleontology, University of Michigan 28(5):93-100 [W. Clyde/W. Clyde/W. Clyde]
29559 D. E. Russell and P. D. Gingerich. 1980. Un nouveau Primate omomyide dans l'Eocène du Pakistan. Comptes Rendus de l'Académie de Paris, Paris, Série D 291:621-624 [J. Alroy/J. Alroy]
13809 D. E. Russell and P. D. Gingerich. 1981. Lipotyphla, Proteutheria(?), and Chiroptera (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(14):277-287 [W. Clyde/W. Clyde/W. Clyde]
59921 D. E. Russell and P. D. Gingerich. 1987. Nouveaux primates de l’Éocène du Pakistan. Comptes Rendus de l'Academie des Sciences, Paris, Série II 5:209-214 [P. Mannion/P. Mannion]
14336 J. G. M. Thewissen, P. D. Gingerich, and D. E. Russell. 1987. Artiodactyla and Perissodactyla (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 27(11):247-274 [W. Clyde/W. Clyde/M. Uhen]
11990 J. G. M. Thewissen and S. T. Hussain. 1998. Systematic review of the Pakicetidae, early and middle Eocene Cetacea (Mammalia) from Pakistan and India. Bulletin of Carnegie Museum of Natural History 34:220-238 [M. Uhen/M. Uhen/M. Uhen]