Blasi 3, Arén: Late/Upper Maastrichtian, Spain
collected 1998-2007

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
cf. Cuniculichnus seilacheri
Cruzado-Caballero et al. 2021
trace fossils on Arenysaurus bones
Reptilia - Testudines - Bothremydidae
Bothremydidae indet. Baur 1891
Murelaga and Canudo 2005
Reptilia - Testudines
cf. Polysternon sp. Portis 1882
López-Martínez et al. 2001
Reptilia - Crocodyliformes
"Trematochampsidae" indet. Buffetaut 1974
López-Martínez et al. 2001
    = Crocodyliformes indet. Hay 1930
Puértolas-Pascual et al. 2016
Reptilia
Eusuchia indet. Huxley 1875
López-Martínez et al. 2001
Alligatoroidea indet. Gray 1844
López-Martínez et al. 2001
    = Eusuchia indet. Huxley 1875
Puértolas-Pascual et al. 2016
Reptilia - Hadrosauridae
aff. Lambeosaurinae indet. Parks 1923
Canudo et al. 2005 1 specimen
    = Lambeosaurinae indet. Parks 1923
Cruzado-Caballero et al. 2005
MPZ 2005/90
Hadrosauridae indet. Cope 1869
35 specimens
(5 measurements)
    = Arenysaurus ardevoli n. gen., n. sp. Pereda-Suberbiola et al. 2009
Pereda-Suberbiola et al. 2009
MPZ 2008/1 (holotype), partial skull. Paratypes: MPZ 2008/256 (maxilla), 2008/257 (maxilla), 2008/258 (dentary), 2008/259 (surangular), 2008/260-263 (isolated teeth), 2007/706, 954-955, 2008/264-267 (seven cervicals), 2008/268 (dorsal), 2008/269-270 (dorsal ribs), 2008/271 (partial sacrum), 2004/480 (caudal vertebra), 2006/20 (articulated caudals), 2008/272, 313 (caudal vertebrae), 2008/314, 330 (haemal arches), 2008/331-332 (ossified tendons), 2008/333a-b (scapula), 2008/334 (coracoid), 2008/336 (humerus), 2008/335 (ilium), 2007/707 (publis), 2007/711, 2008/337 (femora); MPZ2011/1, left jugal; MPZ2012/767, cervical vertebrae; MPZ2012/746–747, MPZ2012/767, three cervical ribs; MPZ2012/748–753, six dorsal ribs; MPZ2008/275–276, MPZ2008/281, MPZ2008/286, MPZ2008/288, MPZ2008/290–292, MPZ2008/296, MPZ2008/298–300, MPZ2008/302, MPZ2008/304, MPZ2008/306, MPZ2008/309–312, MPZ2012/754–758, 24 caudal vertebra; MPZ2008/315–317, MPZ2008/319, MPZ2008/321–323, MPZ2008/325–326, MPZ2008/328–329, 11 hemal arches
Reptilia
Neoceratosauria indet. Novas 1991
Canudo et al. 2005 2 specimens
MPZ 98/67, 2004/8
    = Theropoda informal indet. 1 Marsh 1881
Torices et al. 2015
Reptilia - Dromaeosauridae
? Velociraptorinae indet. Barsbold 1983
López-Martínez et al. 2001 1 specimen
"Dromaeosauridae sp. B"; MPZ 98/68
    = cf. Dromaeosauridae informal indet. 5 Colbert and Russell 1969
Torices et al. 2004
unclassified
Ophiomorpha sp. Lundgren 1891
López-Martínez et al. 2001
see common names

Geography
Country:Spain State/province:Aragón County:Huesca
Coordinates: 42.3° North, 0.7° East (view map)
Paleocoordinates:34.3° North, 0.6° East
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
Key time interval:Late/Upper Maastrichtian
Age range of interval:70.60000 - 66.00000 m.y. ago
Stratigraphy
Geological group:Tremp Formation:La Posa
Local section:Areny Local bed:10 m
Local order:bottom to top
Stratigraphic resolution:bed
Stratigraphy comments: lower part of formation; upper part of chron c30n
The Conques Formation is equivalent to the "lower Red Garumen" facies. The Conques Formation was previously assigned a late maastrichtian age (Sellés and Villa 2015; Villa et al. 2015) Biostratigraphy and Sr stable isotope geochemistry suggest that the La Posa and Conques Formation are entirely early Maastrichtian, lying within chron C31r (see Caus et al. 2016; Oms et al. 2016; Fondevilla et al. 2016; Fondevilla et al. 2019).
Actually in the "Grey Garumnian" = La Posa Fm.
Lithology and environment
Primary lithology:burrows,fine,gray lithified calcareous sandstone
Includes fossils?Y
Lithology description: "1-m-thick channelized sandstone body...fine-grained, matrix-supported feldspathic sandstone contains Ophiomorpha burrows." "a 1-m thick grey massive calcareous sandstone containing Ophiomorpha burrows."; a greywacke
Environment:"channel"
Taphonomy
Modes of preservation:body
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Spatial orientation:random
Preservation of anatomical detail:variable
Articulated whole bodies:none
Associated major elements:many
Disassociated major elements:some
Disassociated minor elements:some
Fragmentation:frequent
Feeding/predation traces:tooth marks
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,chemical,mechanical,field collection
Rock censused:7000 cm2 (area)
Reason for describing collection:biostratigraphic analysis
Collection dates:1998-2007
Collection method comments: area of 70 sq m
Taxonomic list comments:Hadrosaur materials include pathological caudal vertebra (ref 28218).
Metadata
Database number:45438
Authorizer:M. Carrano, P. Holroyd, P. Mannion, R. Butler Enterer:M. Carrano, P. Mannion, K. Maguire, P. Holroyd, R. Butler
Modifier:M. Carrano Research group:vertebrate
Created:2004-10-22 13:51:22 Last modified:2023-08-07 14:32:28
Access level:the public Released:2004-10-22 13:51:22
Creative Commons license:CC BY
Reference information

Primary reference:

17660.ETE J. I. Canudo, L. Ardévol, G. Cuenca-Bescós, N. López-Martínez, X. Murelaga, X. Pereda-Suberbiola, J. I. Ruiz-Omeñaca and X. Orue-Etxebarria. 1999. New dinosaur localities near the Cretaceous/Tertiary boundary from Arén (Huesca, Spain). In J. I. Canudo & G. Cuenca-Bescós (ed.), IV European Workshop on Vertebrate Paleontology. Programme and Abstracts. Field Guide 40-41 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

56254 A. Blanco, A. Prieto-Márquez, and S. De Esteban-Trivigno. 2015. Diversity of hadrosauroid dinosaurs from the Late Cretaceous Ibero-Armorican Island (European Archipelago) assessed from dentary morphology. Cretaceous Research 56:447-457 [M. Carrano/M. Carrano/M. Carrano]
14882ETE J. I. Canudo. 2004. El caso del dinosaurio hadrosaurio de Arén (Huesca): muerte accidental o asesinato? [The case of the hadrosaur dinosaur of Arén (Huesca): accidental death or assassination?]. Naturaleza Aragonesa 13:4-14 [M. Carrano/K. Maguire/M. Carrano]
19728ETE J. I. Canudo, J. A. Andrés, J. L. Barco, P. Cruzado-Caballero, G. Cuenca-Bescós, R. Royo-Torres, C. J. Rubio, C. Rubio, and J. I. Ruiz-Omeñaca. 2006. Rentabilizando socialmente el patrimonio paleontológico: los últimos dinosaurios europeaos (Arén, Huesca) [Making the paleontological patrimony socially profitable: the last European dinosaurs (Arén, Huesca)]. VII Reunión Nacional de la Comisión de Patrimonio Geológico, Museo del Jurásico de Asturias (MUJA), Colunga, Asturia, 28-30 June, 2006. Sociedad Geológica de España [M. Carrano/M. Carrano]
17600ETE J. I. Canudo, J. L. Barco, P. Cruzado-Caballero, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca and R. Royo-Torres. 2005. Evidencias de predación de dinosaurios terópodos en el Maastrichtiense superior, Cretácico superior de Arén (Huesca) [Evidence of theropod dinosaur predation in the upper Maastrichtian, Upper Cretaceous of Arén (Huesca)]. Lucas Mallada 12:29-58 [M. Carrano/M. Carrano]
14596ETE J. I. Canudo, P. Cruzado-Caballero, and M. Moreno-Azanza. 2005. Possible theropod predation evidence in hadrosaurid dinosaurs from the Upper Maastrichtian (Upper Cretaceous) of Arén (Huesca, Spain). Current Research in Vertebrate Palaeontolgy. 3rd Annual Meeting of the European Association of Vertebrate Palaeontologists (EAVP). Kaupia 14:9-13 [M. Carrano/K. Maguire/M. Carrano]
28218ETE J. I. Canudo and P. Cruzado-Caballero. 2007. Patologías en una vértebra caudal de un dinosaurio hadrosaurio como evidencia de predación de terópodos (Maastrichtiense superior, Arén, Huesca) [Pathologies in a caudal vertebra from a hadrosaurian dinosaur as evidence of predation by theropods (upper Maastrichtian, Arén, Huesca)]. IX Congreso Nacional de Paleopatología, Morella, Spain 66 [M. Carrano/M. Carrano]
28738ETE J. I. Canudo, G. Cuenca-Bescós, and J. I. Ruiz-Omeñaca. 2004. Los últimos dinosaurios europeos: debates recientes sobre la extinción de los dinosaurios [The last European dinosaurs: recent debates on the extinction of the dinosaurs]. Historia Natural 7:42-51 [M. Carrano/M. Carrano]
17647ETE J. I. Canudo and J. I. Ruiz-Omeñaca. 2003. Los restos directos de dinosaurios teropódos (excluyendo Aves) en España [Direct remains of theropod dinosaurs (excluding Aves) in Spain]. Ciencias de la Tierra 26:347-373 [M. Carrano/M. Carrano]
16809ETE P. Cruzado-Caballero, J. I. Canudo, and J. I. Ruiz-Omeñaca. 2005. Nuevas evidencias de la presencia de hadrosaurios lambeosaurinos (Dinosauria) en el Maastrichiense superior de la Península Ibérica (Arén, Huesca) {New evidence of the presence of lambeosaurine hadrosaurs (Dinosauria) in the upper Maastrichtian of the Iberian Peninsula (Arén, Huesca)]. Geogaceta 38:47-50 [M. Carrano/M. Carrano]
17658ETE P. Cruzado-Caballero and J. I. Canudo. 2006. Nuevas evidencias de ornitópodos "enanos" en el Maastrichtiense superior Europeo (Blasi 5, Arén, Huesca) [New evidences of "dwarf" ornithopods in the European upper Maastrichtian (Blasi 5, Arén, Huesca)]. IV Encuentro de Jóvenes Investigadores en Paleontología, Salamanca, Spain 75-76 [M. Carrano/M. Carrano]
19766ETE P. Cruzado-Caballero and J. I. Canudo. 2006. El dinosaurio hadrosaurio de Blasi 5 (Maastrichtiense superior, Formación Conques, Arén, Huesca) [The hadrosaurian dinosaur from Blasi 5 (upper Maastrichtian, Conques Formation, Arén, Huesca)]. In E. Fernández-Martínez (ed.), XXII Jornadas de la Sociedad Española de Paleontología, Léon. Libro de resúmenes 108-110 [M. Carrano/M. Carrano]
28219ETE P. Cruzado-Caballero, J. I. Canudo, and J. I. Ruiz-Omeñaca. 2007. Los dinosaurios hadrosaurios (Ornithischia: Ornithopoda) del Maastrichtiense superior (Cretácico Superior) de Arén (Huesca, España) [The hadrosaurian dinosaurs (Ornithischia: Ornithopoda) from the upper Maastrichtian (Upper Cretaceous) of Arén (Huesca, Spain)]. IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, Salas de los Infantes, Spain 55-56 [M. Carrano/M. Carrano/M. Carrano]
28220ETE P. Cruzado-Caballero and J. I. Canudo. 2007. Biodiversidad de los dinosaurios hadriosaurios de Blasi 1 y 3, Maastrichtiense superior (Cretácico Superior, Arén, Huesca) [Biodiversity of hadrosaurian dinosaurs from Blasi 1 and 3, upper Maastrichtian (Upper Cretaceous, Arén, Huesca)]. In: J. C. Braga, A. Checa, & M. Company (eds.), XXIII Jornadas de la Sociedad Española de Paleontología, Libro de resúmenes. Instituto Geológico y Minero de España y Universidad de Granada 38-39 [M. Carrano/M. Carrano]
49729 P. Cruzado-Caballero, J. I. Canudo, and M. Moreno-Azanza. 2013. New material and phylogenetic position of Arenyasaurus ardevoli, a lambeosaurine dinosaur from the Late Maastrichtian of Aren (northern Spain). Journal of Vertebrate Paleontology 33(6):1367-1384 [P. Mannion/J. Tennant]
85592 P. Cruzado-Caballero, J. I. Canudo, S. De Valais, J. Frigola, E. Barriuso and J. Fortuny. 2021. Bioerosion and palaeoecological association of osteophagous insects in the Maastrichtian dinosaur Arenysaurus ardevoli. Lethaia 54(5):957-968 [M. Carrano/M. Carrano]
19147ETE F. M. Dalla Vecchia. 2006. Telmatosaurus and the other hadrosaurids of the Cretaceous European Archipelago. An overview. Natura Nascosta 32:1-55 [M. Carrano/M. Carrano]
41120 F. M. Dalla Vecchia. 2009. Telmatosaurus and the other hadrosauroids of the Cretaceous European Archipelago. An update. Natura Nascosta 39:1-18 [M. Carrano/M. Carrano]
56054 F. M. Dalla Vecchia. 2014. An overview of the latest Cretaceous hadrosauroid record in Europe. In D. A. Eberth & D. C. Evans (ed.), Hadrosaurs 268-297 [M. Carrano/M. Carrano]
73885 F. M. Dalla Vecchia. 2020. The unusual tail of Tethyshadros insularis (Dinosauria, Hadrosauroidea) from the Adriatic Island of the European archipelago. Rivista Italiana di Paleontologia e Stratigrafia 126(3):583-628 [M. Carrano/M. Carrano]
55204 A. Galobart, A. Santos-Cubedo, M. Suñer and J. I. Canudo. 2011. Paleontological discoveries and studies. In À. Galobart, M. Suñer, & B. Poza (eds.), Dinosaurs of Eastern Iberia 1-23 [M. Carrano/M. Carrano/M. Carrano]
80599 E. Isasmendi, A. Torices, J. I. Canudo, P. J. Currie, and X. Pereda-Suberbiola. 2022. Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño. Papers in Palaeontology 8(1):e1419:1-38 [M. Carrano/M. Carrano]
11838ETE N. López-Martínez, J. I. Canudo, L. Ardèvol, X. Pereda Suberbiola, X. Orue-Etxebarria, G. Cuenca-Bescós, J. I. Ruiz-Omeñaca, X. Murelaga, and M. Feist. 2001. New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe. Cretaceous Research 22(1):41-61 [M. Carrano/M. Carrano/M. Carrano]
59760 X. Murelaga and J. I. Canudo. 2005. Descripción de los restos de quelonios del Maastrichtiense superior de Aren y Serraduy (Huesca) [Description of chelonian remains from the Upper Maastrichtian of Aren and Seraduy (Huesca)]. Geogaceta 38:51-54 [P. Holroyd/P. Holroyd/M. Uhen]
17698ETE O. Oms and J. I. Canudo. 2004. Datación magnetoestratigrafica de los dinosaurios del Cretácico terminal (Maastrichtiense superior) de Arén (Huesca, Unidad Surpirenaica Central) [Magnetostratigraphic dating of the terminal Cretaceous dinosaurs (upper Maastrichtian) of Arén (Huesca, central Pyrénées)]. Geo-Temas 6(5):51-54 [M. Carrano/M. Carrano]
33025ETE X. Pereda-Suberbiola, J. I. Canudo, J. Company, P. Cruzado-Caballero, and J. I. Ruiz-Omenaca. 2009. Hadrosauroid dinosaurs from the Latest Cretaceous of the Iberian Peninsula. Journal of Vertebrate Paleontology 29(3):946-951 [M. Carrano/M. Oreska]
31506 X. Pereda-Suberbiola, J. I. Canudo, P. Cruzado-Caballero, J. L. Barco, N. López-Martínez, O. Oms, and J. I. Ruiz-Omeñaca. 2009. The last hadrosaurid dinosaurs of Europe: a new lambeosaurine from the uppermost Cretaceous of Aren (Huesca, Spain). Comptes Rendus Palevol 8:559-572 [R. Butler/R. Butler/M. Carrano]
81262 M. Pérez-Pueyo, P. Cruzado-Caballero, M. Moreno-Azanza, B. Vila, D. Castanera, J. M. Gasca, E. Puértolas-Pascual, B. Bádenas, and J. I. Canudo. 2021. The tetrapod fossil record from the uppermost Maastrichtian of the Ibero-Armorican Island: an integrative review based on the outcrops of the western Tremp Syncline (Aragón, Huesca Province, NE Spain). Geosciences 11:162:1-38 [M. Carrano/M. Carrano]
47749 A. Prieto-Marquez, F. M. D. Vecchia, R. Gaete and A. Galobart. 2013. Diversity, relationships, and biogeography of the lambeosaurine dinosaurs from the European archipelago, with description of the new aralosaurin Canardia garonnensis. PLoS One 8(7):e69835:1-44 [P. Mannion/J. Tennant/M. Carrano]
76284 E. Puértolas-Pascual, A. Blanco, C. A. Brochu and J. I. Canudo. 2016. Review of the Late Cretaceous-early Paleogene crocodylomorphs of Europe: Extinction patterns across the K-PG boundary. Cretaceous Research 57:565-590 [P. Mannion/P. Mannion]
75349 A. G. Sellés, B. Vila, and A. Galobart. 2014. Diversity of theropod ootaxa and its implications for the latest Cretaceous dinosaur turnover in southwestern Europe. Cretaceous Research 49:45-54 [M. Carrano/M. Carrano/P. Mannion]
56091 A. Torices, P. J. Currie, J. I. Canudo and X. Pereda-Suberbiola. 2015. Theropod dinosaurs from the Upper Cretaceous of the South Pyrenees Basin of Spain. Acta Palaeontologica Polonica 60(3):611-626 [P. Mannion/J. Tennant/M. Carrano]
17618ETE A. Torices, J. I. Ruiz-Omeñaca, J. I. Canudo and N. López-Martínez. 2004. Nuevos datos sobre los dinosaurios terópodos (Saurischia: Theropoda) del Cretácico superior de los Pirineos Sur-Centrales (Huesca y Lleida) [New data on the theropod dinosaurs (Saurischia: Theropoda) from the Upper Cretaceous of the south-central Pyrenées (Huesca and Lleida)]. Geo-Temas 6(5):71-74 [M. Carrano/M. Carrano]