Steengroeve Winterswijk: Anisian, Netherlands
collected >1935

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli
Capitosauroides sp. Haubold 1971
Demathieu and Oosterink 1988 1 individual
slab D.367
Reptilia
Rhynchosauroides sp. Maidwell 1911
Reptilia - Tanystropheidae
Tanystropheus antiquus Huene 1905
Wild and Oosterink 1984 1 individual
Specimen A 638, H. Oosterink collection, middle neck vertebra
    = Amotosaurus rotfeldensis Fraser and Rieppel 2006
Klein et al. 2015
Reptilia - Helveticosauridae
Saurosphargis sp. Frech 1903
Bickelmann and Sander 2008
    = Eusaurosphargis sp. Nosotti and Rieppel 2003
Sander et al. 2014
Reptilia - Chirotheriidae
Brachychirotherium cf. circaparvum Demathieu 1971
1 individual
slab D.277
    = Brachychirotherium paraparvum n. sp. Demathieu and Oosterink 1988
Demathieu and Oosterink 1988
Rhynchosauroides peabodyi (Faber 1958)
original and current combination Chirotherium peabodyi
Reptilia - Batrachopodidae
Sustenodactylus hollandicus n. sp. Demathieu and Oosterink 1988
Demathieu and Oosterink 1988 1 individual
slab D.340
Reptilia
Coelurosaurichnus ratumensis n. sp. Demathieu and Oosterink 1988
Demathieu and Oosterink 1988 1 individual
slab D.416
Reptilia - Placodontia - Placodontidae
Placodus sp. Agassiz 1833
Bickelmann and Sander 2008
Oosterink,1986; Oosterink and Diepenbroek, 1990; Oosterink et al., 2003
Reptilia
Palatodonta bleekeri n. gen., n. sp. Neenan et al. 2013
Neenan et al. 2013 1 individual
holotype juvenile skull TW480000470
Reptilia - Eosauropterygia - Nothosauridae
Nothosaurus sp. Münster 1834
Bickelmann and Sander 2008 1 individual
"The only other Nothosaurus from Winterswijk deposited in a public repository is a very incompletely preserved skull (NME 480000126) of unknown stratigraphic provenance"
Nothosaurus marchicus Koken 1893
Klein et al. 2015 2 individuals
TWE 480000473, cranium and partial skeleton; JLW 300, cranium and disarticulated skeleton
Nothosaurus winterswijkensis n. sp. Albers and Rieppel 2003
Albers and Rieppel 2003 3 individuals
synonym of Nothosaurus marchicus
holotype skull: NMNHL St 445530; An additional incomplete lower jaw (NMNHL St 445913) described by Hooijer (1959), an almost complete lower jaw (NMNHL St 447170) and an incomplete skull with partial lower jaw (NME 480000126) collected by Henk Kolstee. Additional Nothosaurus skulls are known from private collections (Oosterink and Diepenbroek 1990)
Nothosaurus winkelhorsti n. sp. Klein and Albers 2009
Klein and Albers 2009 1 individual
recombined as Lariosaurus winkelhorsti
NMNHL RGM 443825 (holotype)
Lariosaurus vosseveldensis n. sp. Klein et al. 2016
Klein et al. 2016 1 individual
TWE 480000504, skull with assorted postcrania
Reptilia - Eosauropterygia - Pachypleurosauridae
Dactylosaurus cf. gracilis Gürich 1884
Bickelmann and Sander 2008
Oosterink et al., 2003
Anarosaurus heterodontus Rieppel and Lin 1995
Bickelmann and Sander 2008
Hooijer, 1959;Oosterink, 1986; Oosterink et al., 2003
Reptilia
Procolophonichnium haarmuehlensis (Holst et al. 1970)
Bickelmann and Sander 2008
tracks
Procolophonichnium winterswijkense n. sp. Demathieu and Oosterink 1983
unclassified
Phenacopus agilis n. sp. Demathieu and Oosterink 1983
Phenacopus faberi n. sp. Demathieu and Oosterink 1983
Bifungites sp. Desio 1940
Demathieu and Oosterink 1988
? Kouphichnium sp. Nopcsa 1923
Demathieu and Oosterink 1988 1 individual
slab D.277
see common names

Geography
Country:Netherlands State/province:Gelderland
Coordinates: 52.0° North, 6.7° East (view map)
Paleocoordinates:16.6° North, 16.4° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
Key time interval:Anisian
Age range of interval:247.20000 - 242.00000 m.y. ago
Stratigraphy
Geological group:Muschelkalk Formation:Vossenveld Member:Lower Wellenkalk layer 9
Stratigraphic resolution:group of beds
Stratigraphy comments: "middle Lower Muschelkalk"
Original entry for Chronostratigraphy was >>Ladinian<<. Originally entered formation name was >>Muschelkalk<<. Corrections made are according to Klein & Albers (2009). (TL)
"Lower Anisian, lowermost Middle Triassic" according to Albers & Rieppel (2003)
Lithology and environment
Primary lithology:desiccation cracks,wave ripples,gray,blue lithified "limestone"
Includes fossils?Y
Lithology description: "The presence of bluish-gray limestones and marls, yellowish dolomites and black-grayish mudstones indicate a shallow marine and near shore environment. Sediments with ripple marks and polygonal mud cracks even indicate a tidal flat facies, with the mud crack beds representing algeal laminates produced by cyanobacteria stabilizing the sediment surface. Bones of Muschelkalk reptiles occur accumulated in specific layers, for example in bed IX" "The present skull was found in what is described as layer 9 by Oosterink (1986), which relates to what Harsveldt (1973) described as lower Wellenkalk, located 0.70–0.95 m below a distinct chocolate brown calcareous marl layer and 0.74–0.99 m above Dolomitic Limestone VI (Harsveldt, 1973)." (Albers & Rieppel 2003)
Environment:peritidal
Taphonomy
Modes of preservation:body,cast,mold/impression,trace
Size of fossils:macrofossils
Preservation of anatomical detail:good
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:some macrofossils,ichnofossils
Collection methods:salvage,selective quarrying,surface (in situ),mechanical,field collection,survey of museum collection
Reason for describing collection:general faunal/floral analysis
Collection dates:>1935
Collection method comments: Near the town of Winterswijk, in the mid-eastern part of The Netherlands, the Muschelkalk is exposed in a relatively small area where Triassic limestone has been commercially exploited since 1935.

TW, Twentse Welle Museum, Enschede, the Netherlands, collection
Metadata
Also known as:Winterswijkse Steen- en Kalkgroeve
Database number:96875
Authorizer:M. Carrano, R. Benson, J. Mueller, A. Dunhill Enterer:H. Street, B. Allen, M. Carrano, R. Benson, T. Liebrecht
Modifier:M. Carrano Research group:vertebrate
Created:2010-07-07 12:33:51 Last modified:2021-04-27 13:31:22
Access level:the public Released:2010-07-07 12:33:51
Creative Commons license:CC BY
Reference information

Primary reference:

67942. G. R. Demathieu and H. W. Oosterink. 1983. Die wirbeltier-ichnofauna aus dem unteren Muschelkalk von Winterswijk. Staringia 7(1):1-51 [A. Dunhill/B. Allen]

Secondary references:

44157 P. C. H. Albers and O. Rieppel. 2003. A new species of the sauropterygian genus Nothosaurus from the Lower Muschelkalk of Winterswijk, The Netherlands. Journal of Paleontology 77(4):738-744 [R. Benson/R. Benson]
33074 C. Bickelmann and P. M. Sander. 2008. A partial skeleton and isolated humeri of Nothosaurus (Reptilia: Eosauropterygia) from Winterswijk, The Netherlands. Journal of Vertebrate Paleontology 28(2):326-338 [M. Carrano/H. Street/H. Street]
76686 G. R. Demathieu and H. W. Oosterink. 1988. New discoveries of ichnofossils from the Middle Triassic of Winterswijk (the Netherlands). Geologie en Mijnbouw 67:3-17 [M. Carrano/M. Carrano/M. Carrano]
76668 H. Klein and S. G. Lucas. 2021. The Triassic Tetrapod Footprint Record. New Mexico Museum of Natural History & Science Bulletin 83:1-194 [M. Carrano/M. Carrano/M. Carrano]
38255 N. Klein and P. C. H. Albers. 2009. A new species of the sauropsid reptile Nothosaurus from the Lower Muschelkalk of the western Germanic Basin, Winterswijk, The Netherlands. Acta Palaeontologica Polonica 54(4):589-598 [J. Mueller/T. Liebrecht]
66892 N. Klein, D. F. A. Voeten, J. Lankamp, R. Bleeker, O. J. Sichelschmidt, M. Liebrand, D. C. Nieweg and P. M. Sander. 2015. Postcranial material of Nothosaurus marchicus from the Lower Muschelkalk (Anisian) of Winterswijk, The Netherlands, with remarks on swimming styles and taphonomy. Paläontologische Zeitschrift 89(4):961-981 [A. Dunhill/B. Allen]
66832 N. Klein, D. F. A. Voeten, A. Haarhuis and R. Bleeker. 2016. The earliest record of the genus Lariosaurus from the early middle Anisian (Middle Triassic) of the Germanic Basin. Journal of Vertebrate Paleontology 36(4):e1163712 [A. Dunhill/B. Allen]
46023 J. M. Neenan, N. Klein, and T. M. Scheyer. 2013. European origin of placodont marine reptiles and the evolution of crushing dentition in Placodontia. Nature Communications [R. Benson/R. Benson]
50282 P. M. Sander, N. Klein, P. C. H. Albers, C. Bickelmann, and H. Winkelhorst. 2014. Postcranial morphology of a basal Pistosauroidea (Sauropterygia) from the Lower Muschelkalk of Winterswijk, The Netherlands. Palaeontologische Zeitschrift 88:55-71 [P. Mannion/J. Tennant]
60738 R. Wild and H. Oosterink. 1984. Tanystropheus (Reptilia: Squamata) aus dem Unteren Muschelkalk von Winterswijk, Holland. Grondboor en Hamer 5:142-148 [R. Butler/R. Butler]