Isle of Sheppey (BMNH collection) (Eocene of the United Kingdom)

Where: England, United Kingdom (51.4° N, 0.8° E: paleocoordinates 46.9° N, 1.6° W)

• coordinate based on political unit

• local area-level geographic resolution

When: London Clay Formation, Ypresian (56.0 - 47.8 Ma)

• MP8 + 9, NP12 (from Smith & Smith, 2003)

• formation-level stratigraphic resolution

Environment/lithology: marine; claystone

Size class: macrofossils

Reposited in the BMNH

• Paratype of n. gen. and sp. Pimapus lacki, BMNH A3710, collected in 1936 by A. G. Davis.

Primary reference: C. J. O. Harrison and C. A. Walker. 1975. A new swift from the Lower Eocene of Britain. Ibis 117(2):162-164 [J. Hunter/J. Hunter/J. Hunter]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 13299: authorized by John Hunter, entered by John Hunter on 19.02.2002, edited by Erin Leckey, Mark Uhen, John Alroy, Lerin Baltzly, Juan Benito Moreno, Matthew Carrano, Bethany Allen and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

• whale locality, "north shore Isle of Sheppey, Ord. Surv. sheet 172 map ref 975734"
Show authors, comments, and common names
unclassified
  -
Brachaetus muelleri2, Aulopsis egertoni n. gen. n. sp.2, Aulopsis depressifrons n. sp.2, Beerichtys ingens n. gen. n. sp.2, Laparodon alticeps n. gen. n. sp.2, Eoceolopoma colei2, Eoceolopoma curvatum2, Eoceolopoma hopwoodi n. sp.2, Eoceolopoma gigas n. sp.2, Ariodus marriotti2, Ancestrus ornatus2, Ancestrus elongatus n. sp.2, Neptunavis miranda n. gen. n. sp.13
Aulopsis egertoni n. gen. n. sp.2
Beerichtys ingens n. gen. n. sp.2
Laparodon alticeps n. gen. n. sp.2
Eoceolopoma curvatum2
f. tenuistriatum
Neptunavis miranda n. gen. n. sp.13
Malacostraca
 Decapoda - Homolidae
Londinimola williamsi3
Londinimola williamsi3 Collins and Saward 2006 crab
 Decapoda - Scyllaridae
"Scyllaridia belli" = Scyllarides15
"Scyllaridia belli" = Scyllarides15 decapod
Mammalia
 Perissodactyla - Palaeotheriidae
Hyracotherium leporinum1
Hyracotherium leporinum1 Owen 1841 odd-toed ungulate
 Perissodactyla - Equidae
"Hyracotherium aff. vulpiceps" = Pliolophus vulpiceps5
"Hyracotherium aff. vulpiceps" = Pliolophus vulpiceps5 Owen 1858 horse
BMNH 38939, 47986 - mandibular fragments without tooth crowns
 Creodonta - Oxyaenidae
Argillotherium toliapicum n. sp.1
Argillotherium toliapicum n. sp.1 Davies 1884 creodont
 Theriamorpha - Nyctitheriidae
Leptacodon sp.12
Leptacodon sp.12 Matthew and Granger 1921 placental
 Primates - Plesiadapidae
"Platychoerops richardsonii" = Platychoerops richardsoni1
"Platychoerops richardsonii" = Platychoerops richardsoni1 Charlesworth 1855 primate
 Primates - Toliapinidae
Toliapina lawsoni n. sp.6
Toliapina lawsoni n. sp.6 Hooker et al. 1999 primate
NHMUK M51610 - type
Reptilia
 Squamata - Palaeophiidae
Palaeophis longus n. sp.11
Palaeophis longus n. sp.11 Owen 1849 snake
 Testudines - Dermochelyidae
"Anglocetus beatsoni n. gen. n. sp." = Eosphargis gigas14
"Anglocetus beatsoni n. gen. n. sp." = Eosphargis gigas14 Owen 1880 leatherback
fragments of right scapula
Aves
 Procellariiformes - Procellariidae
Primodroma bournei8
Primodroma bournei8 Harrison and Walker 1977 petrel
 Apodiformes -
Primapus lacki
Primapus lacki Harrison and Walker 1975 swift
paratype
 Odontopterygiformes - Pelagornithidae
Macrodontopteryx oweni n. gen. n. sp.4, "Lithornis emuinus n. sp." = Dasornis emuinus7, "Argillornis longipennis n. sp." = Dasornis emuinus13, "Dasornis londinensis n. gen. n. sp." = Dasornis emuinus13, "Odontopteryx toliapica n. gen. n. sp." = Dasornis toliapica7
Macrodontopteryx oweni n. gen. n. sp.4 Harrison and Walker 1976 bird
"Lithornis emuinus n. sp." = Dasornis emuinus7 Bowerbank 1854 bird
"Argillornis longipennis n. sp." = Dasornis emuinus13 Bowerbank 1854 bird
"Dasornis londinensis n. gen. n. sp." = Dasornis emuinus13 Bowerbank 1854 bird
"Odontopteryx toliapica n. gen. n. sp." = Dasornis toliapica7 Owen 1873 bird
Actinopteri
 Amiiformes - Amiidae
Lehmanamia sheppeyensis n. gen. n. sp.2
Lehmanamia sheppeyensis n. gen. n. sp.2 Casier 1966 bowfin
 Acanthopteri - Sphyraenidae
Sphyraena bognorensis n. sp.2
Sphyraena bognorensis n. sp.2 Casier 1966 barracuda
 Albuliformes - Albulidae
Albula oweni2
Albula oweni2 Agassiz 1844 bonefish
 Elopiformes - Osmeroididae
"Halecopsis insignis" = Osmeroides insignis2
"Halecopsis insignis" = Osmeroides insignis2 Delvaux and Orlieb 1844
 Elopiformes - Elopidae
Elops sp.2, Esocelops cavifrons2
Elops sp.2 Linnaeus 1766
Esocelops cavifrons2 Woodward 1901
 Elopiformes - Megalopidae
"Megalops priscus" = Protarpon priscus2, Promegalops signeuxae n. gen. n. sp.2, Promegalops sheppeyensis n. sp.2, Megalops oblongus2
"Megalops priscus" = Protarpon priscus2 Woodward 1901
Promegalops signeuxae n. gen. n. sp.2 Casier 1966
Promegalops sheppeyensis n. sp.2 Casier 1966
Megalops oblongus2 Woodward 1901
 Elopiformes - Phyllodontidae
Phyllodus toliapicus2, Egertonia isodonta2
Phyllodus toliapicus2 Agassiz 1839
Egertonia isodonta2 Cocchi 1866
 Siluriformes - Bagridae
Bucklandium diluvii2
Bucklandium diluvii2 König 1825 naked catfish
 Aulopiformes - Aulopidae
Labrophagus esocinus n. gen. n. sp.2
Labrophagus esocinus n. gen. n. sp.2
 Aulopiformes - Synodidae
Argillichthys toombsi n. gen. n. sp.2
Argillichthys toombsi n. gen. n. sp.2 Casier 1966
 Gadiformes - Merlucciidae
Rhinocephalus planiceps n. gen. n. sp.2
Rhinocephalus planiceps n. gen. n. sp.2 Casier 1966
 Holocentriformes - Holocentridae
Myripristis toliapicus2, Naupygus bucklandi n. gen. n. sp.2
Myripristis toliapicus2 Agassiz 1845
Naupygus bucklandi n. gen. n. sp.2 Casier 1966
 Carangiformes - Carangidae
"Teratichthys antiquitatis" = Teratichthys antiquitatus2
"Teratichthys antiquitatis" = Teratichthys antiquitatus2 Konig 1825
 Istiophoriformes - Xiphiidae
Tamesichthys decipens n. gen. n. sp.2, Xiphiorhynchus sp.2, Xiphiorhynchus parvus n. sp.2
Tamesichthys decipens n. gen. n. sp.2 sword fish
Xiphiorhynchus sp.2 Van Beneden 1871 sword fish
Xiphiorhynchus parvus n. sp.2 Casier 1966 sword fish
 Istiophoriformes - Istiophoridae
Hemirhabdorhynchus elliotti n. sp.2
Hemirhabdorhynchus elliotti n. sp.2 Casier 1966
 Spariformes - Sparidae
Sciaenurus bowerbanki2, Podocephalus nitidus n. gen. n. sp.2, Podocephalus curryi n. sp.2
Sciaenurus bowerbanki2 Agassiz 1845
nov. f. crassior
Podocephalus nitidus n. gen. n. sp.2 Casier 1966
Podocephalus curryi n. sp.2 Casier 1966
 Ephippiformes - Ephippidae
Whitephippus tamesis n. gen. n. sp.2
Whitephippus tamesis n. gen. n. sp.2 Casier 1966
 Labriformes - Labridae
Pseudosphaerodon antiquus n. sp.2
Pseudosphaerodon antiquus n. sp.2 Casier 1966
 Perciformes -
"Xiphiorhynchus priscus" = Tetrapterus priscus2
"Xiphiorhynchus priscus" = Tetrapterus priscus2 Agassiz 1839 perch-like fish
 Perciformes - Serranidae
"Plesioserranus cf. wemmeliensis" = Serranus wemmeliensis2, Serranopsis londinensis n. gen. n. sp.2, Percostoma angustum n. gen. n. sp.2
"Plesioserranus cf. wemmeliensis" = Serranus wemmeliensis2 Storms 1898 perch-like fish
Serranopsis londinensis n. gen. n. sp.2 Casier 1966 perch-like fish
Percostoma angustum n. gen. n. sp.2 Casier 1966 perch-like fish
 Ophidiiformes - Ophidiidae
Ampheristus toliapicus2
 Scombriformes - Scombridae
"Scombrinus macropomus" = Cybium macropomum2, Wetherellus brevior n. sp.2, Wetherellus longior n. sp.2, Wetherellus cristatus n. gen. n. sp.2, Gymnosarda prisca10, Thunnini indet.9, Eothynnus salmoneus n. gen. n. sp.16, Eothynnus salmoneus2, Woodwardella patellifrons n. gen. n. sp.2, Scombrinus nuchalis2, "Cybium cf. proosti" = Palaeocybium proosti2
"Scombrinus macropomus" = Cybium macropomum2 Agassiz 1839
Wetherellus brevior n. sp.2 Casier 1966
Wetherellus longior n. sp.2 Casier 1966
Wetherellus cristatus n. gen. n. sp.2 Agassiz 1844
Gymnosarda prisca10 Monsch 2000
Thunnini indet.9 Starks 1910
BMNH P4300
Eothynnus salmoneus n. gen. n. sp.16 Woodward 1901
Eothynnus salmoneus2 Woodward 1901
Woodwardella patellifrons n. gen. n. sp.2 Casier 1966
Scombrinus nuchalis2 Woodward 1901
"Cybium cf. proosti" = Palaeocybium proosti2 Storms 1895
 Scombriformes - Gempylidae
Progempylus edwardsi n. gen. n. sp.2
Progempylus edwardsi n. gen. n. sp.2 Casier 1966
 Scombriformes -
Sphyraenodus pirscus2, Scombramphodon sheppeyensis n. sp.2, Scombramphodon crassidens2
Scombramphodon sheppeyensis n. sp.2 Casier 1966
 Scombriformes - Palaeorhynchidae
Enniskillenus radiatus n. gen. n. sp.2, Aglyptorhynchus sulcatus2, Aglyptorhynchus venablesi n. sp.2
Enniskillenus radiatus n. gen. n. sp.2 Agassiz 1833
Aglyptorhynchus venablesi n. sp.2 Casier 1966
 Scombriformes - Trichiuridae
Eutrichiurides winkleri2, Trichiurides sagittidens2
Trichiurides sagittidens2 Winkler 1874
 Scombriformes - Bramidae
Goniocranion arambourgi n. gen. n. sp.2, Bramoides brieni n. gen. n. sp.2
Goniocranion arambourgi n. gen. n. sp.2 Casier 1966
Bramoides brieni n. gen. n. sp.2 Casier 1966
 Beryciformes - Berycidae
Argilloberyx prestwichae n. gen. n. sp.2
Argilloberyx prestwichae n. gen. n. sp.2 Casier 1966
 Pycnodontiformes - Pycnodontidae
Pycnodus sp.2, Pycnodus bowerbanki2, Pycnodus toliapicus2
Pycnodus sp.2 Agassiz 1833
Pycnodus bowerbanki2 Egerton 1877
Pycnodus toliapicus2 Agassiz 1839
Osteichthyes
  -
Rhynchorhinus major2, Rhynchorhinus branchialis2
Rhynchorhinus major2 Woodward 1901
Rhynchorhinus branchialis2 Woodward 1901
 Iniomi - Beerichthyidae
Beerichthys sp.2
Beerichthys sp.2 Casier 1966 bony fish
 Acipenseriformes - Acipenseridae
Acipenser toliapicus2
Acipenser toliapicus2 Agassiz 1844 sturgeon
 Acipenseriformes -
Cylindracanthus rectus2
Cylindracanthus rectus2 Agassiz 1843 ray-finned fish
Chondrichthyes
 Odontaspidida - Isuridae
Oxyrhina praecursor2, Oxyrhina sheppeyensis n. sp.2
Oxyrhina praecursor2 Leriche 1942 mackerel shark
Oxyrhina sheppeyensis n. sp.2 Cassier 1966 mackerel shark
 Myliobatiformes - Aetobatidae
Aetobatus irregularis2
Aetobatus irregularis2 Agassiz 1843 stingray
 Myliobatiformes - Myliobatidae
Myliobatis sp.2, Myliobatis toliapicus2, Myliobatis latidens2, "Myliobatis dixoni" = Aetomylaeus dixoni2, Rhinoptera davisesi2
Myliobatis sp.2 Cuvier 1816 eagle ray
Myliobatis toliapicus2 Agassiz 1843 eagle ray
Myliobatis latidens2 Woodward 1888 eagle ray
"Myliobatis dixoni" = Aetomylaeus dixoni2 Agassiz 1843 eagle ray
Rhinoptera davisesi2 cownose ray
 Myliobatiformes - Dasyatidae
Dasyatis davisi n. sp.2, "Raja cf. duponti" = Dasyatis duponti2
Dasyatis davisi n. sp.2 Cassier 1966 whiptail stingray
"Raja cf. duponti" = Dasyatis duponti2 Winkler 1874 whiptail stingray
 Heterodontiformes - Heterodontidae
Heterodontus wardenensis n. sp.2, Heterodontus sp.2, "Heterodontus vincenti" = Cestracion vincenti2
Heterodontus wardenensis n. sp.2 bullhead shark
Heterodontus sp.2 de Blainville 1816 bullhead shark
"Heterodontus vincenti" = Cestracion vincenti2 Leriche 1905 bullhead shark
  - Notidanidae
Xenodolamia eocaena2
Xenodolamia eocaena2 Woodward 1889 shark
 Squatiniformes - Squatinidae
Squatina prima2
Squatina prima2 Winkler 1874 angel shark
 Squaliformes - Squalidae
Squalus minor2
Squalus minor2 Daimeries 1888 dogfish shark
 Squaliformes - Dalatiidae
Isistius trituratus2
Isistius trituratus2 Winkler 1874 shark
 Hexanchiformes - Hexanchidae
"Notidanus serratissimus" = Notorhynchus serratissimus2
"Notidanus serratissimus" = Notorhynchus serratissimus2 Agassiz 1843 sevengill shark
 Lamniformes - Odontaspididae
"Odontaspis robusta" = Jaekelotodus robusta2, "Odontaspis trigonalis" = Jaekelotodus trigonalis2, "Lamna verticalis" = Hypotodus verticalis2, "Lamna lerichei" = Brachycarcharias lerichei2, Odontaspis sp.2, Odontaspis winkleri2
"Odontaspis robusta" = Jaekelotodus robusta2 Leriche 1905 sand tiger shark
"Odontaspis trigonalis" = Jaekelotodus trigonalis2 Jackel 1895 sand tiger shark
"Lamna verticalis" = Hypotodus verticalis2 Agassiz 1843 sand tiger shark
"Lamna lerichei" = Brachycarcharias lerichei2 Casier 1946 sand tiger shark
Odontaspis sp.2 Agassiz 1838 sand tiger shark
Odontaspis winkleri2 Leriche 1905 sand tiger shark
 Lamniformes - Lamnidae
Lamna sp.2, Lamna affinis2
Lamna sp.2 Cuvier 1816 mackerel shark
Lamna affinis2 Casier 1950 mackerel shark
 Lamniformes - Otodontidae
"Odontaspis macrota" = Otodus macrotus2, "Lamna obliqua" = Otodus obliquus2
"Odontaspis macrota" = Otodus macrotus2 Agassiz 1838 mackerel shark
"Lamna obliqua" = Otodus obliquus2 Agassiz 1838 mackerel shark
subspecies subserrata
 Lamniformes - Carchariidae
"Odontaspis hopei" = Carcharias hopei2, "Odontaspis aff. hopei" = Carcharias hopei2
"Odontaspis hopei" = Carcharias hopei2 Agassiz 1843 sand shark
"Odontaspis aff. hopei" = Carcharias hopei2 Agassiz 1843 sand shark
 Carcharhiniformes - Carcharhinidae
Hypoprion sp.2, "Physodon tertius" = Physogaleus secundus2, Physodon secundus2
Hypoprion sp.2 Muller and Henle 1838 requiem shark
"Physodon tertius" = Physogaleus secundus2 Winkler 1876 requiem shark
Physodon secundus2 Winkler 1876 requiem shark
 Carcharhiniformes - Scyliorhinidae
Scyliorhinus sp.2, Scyliorhinus biauriculatus2, Scyliorhinus minutissimus2, "Scyliorhinus aff. gilberti" = Premontreia (Oxyscyllium) gilberti2
Scyliorhinus sp.2 Blainville 1816 cat shark
Scyliorhinus biauriculatus2 Casier 1946 cat shark
Scyliorhinus minutissimus2 Winkler 1873 cat shark
"Scyliorhinus aff. gilberti" = Premontreia (Oxyscyllium) gilberti2 Casier 1946 cat shark
 Carcharhiniformes - Triakidae
Galeorhinus minor2, "Galeorhinus lefevrei" = Pachygaleus lefevrei2
Galeorhinus minor2 Agassiz 1835 school shark
"Galeorhinus lefevrei" = Pachygaleus lefevrei2 Daimeries 1891 hound shark
 Chimaeriformes - Edaphodontidae
"Edaphodon bucklandi" = Edaphodon eurygnathus2
"Edaphodon bucklandi" = Edaphodon eurygnathus2 Agassiz 1843 chimaera
 Chimaeriformes - Callorhynchidae
Elasmodus hunteri2
Elasmodus hunteri2 Egerton 1843 plownose chimaera
 Chimaeriformes - Chimaeridae
Chimaera venablesae2
Chimaera venablesae2 Cassier 1966 chimaera
1Benton et al. 2005; 2Casier 1966; 3Collins and Saward 2006; 4Harrison and Walker 1976; 5Hooker 1980; 6Hooker et al. 1999; 7Mayr 2008; 8Mlikovsky 2002; 9Monsch 2004; 10Monsch and Bannikov 2011; 11Owen 1849; 12Smith and Smith 2003; 13Swinton 1934; 14Tarlo 1963; 15Woodward 1870; 16Woodward 1901