Masia de l'Hereuet (Eocene of Spain)

Where: Lleida, Spain (42.0° N, 0.7° E: paleocoordinates 37.3° N, 0.9° W)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: MP 8+9 mammal zone, Corca Formation, Early/Lower Eocene (55.8 - 48.6 Ma)

• "The site was assigned to the Grauvian (MP10) by Antunes et al. (1997) on the basis of its mammalian fauna. However, after revision of the artiodactyl, perissodactyl, and rodent assemblage, Badiola et al. (2009) concluded that this level is older than previously considered, corresponding to the Neustrian (MP8+9, 51-56 Ma), and represents the oldest continental locality of the Àger sub-basin" (p.430)

• bed-level stratigraphic resolution

Environment/lithology: fluvial; claystone and sandstone

Size class: mesofossils

Primary reference: J. Marigó, R. Minwer-Barakat, S. Moyà-Solà and S. López-Torres. 2012. First record of Plesiadapiformes (Primates, Mammalia) from Spain. Journal of Human Evolution 62(3):429-433 [J. Alroy/A. Garcia Selles]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 142031: authorized by John Alroy, entered by Albert Garcia Selles on 04.04.2013, edited by Philip Mannion and Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Squamata -
Squamata indet.1 Oppel 1811 lizard
IPS 49742, fragment of left dentary; IPS 49746, anterior portion of a right dentary with two teeth; IPS 59530, vertebra; IPS 59532, tooth; IPS 59539, tooth; IPS 83542, seven fragments of toothbearing bones; IPS 83548, four fragments of dentary; IPS 83550, two fragments of tooth-bearing bone; IPS 83541, roughly rounded osteoderm
 Squamata - Necrosauridae
"Necrosauridae" indet.1 Hoffstetter 1943 lizard
IPS 83545, fragment of dentary; IPS 49741, partial osteoderm
 Squamata - Anguidae
Glyptosaurinae indet.1 Marsh 1872 lizard
IPS 35741, three fragments of body osteoderm; IPS 49738 fragment of body osteoderm; IPS 59536, four fragments of body osteoderms; IPS 83547, seven body osteoderms or fragments of body osteoderms, one partial skull osteoderm; IPS 83549, 10 body osteoderms and fragments of osteoderm; IPS 49744, tooth-bearing bone (?dentary) fragment with one tooth; IPS 83539, several dozen osteoderms; IPS 83551, four fragments of osteoderms; IPS 83554, body osteoderms
Glyptosaurini indet.1 Sullivan 1979 lizard
IPS 3742, partial skull osteoderm
Anguinae indet.1 Gray 1825 lizard
IPS 83557, partial parietal; IPS 59538, two partial vertebrae; IPS 83540, two body osteoderms
 Squamata - Iguanidae
Geiseltaliellus sp.1 Kuhn 1944 lizard
IPS 49740, partial left dentary with 10 tooth; IPS 59535, fragment of right dentary; IPS 83552, partial left maxilla with six teeth; IPS 83558, two partial right and two partial left dentaries, one partial maxilla; less certainly IPS 49745, fragment of ?maxilla with two teeth; IPS 83553, partial right dentary
 Squamata - Agamidae
Agamidae indet.1 Gray 1827 lizard
IPS 83546, fragment of tooth-bearing bone with one tooth; IPS 83543, fragment of left dentary with two teeth
 Squamata -
"Scincomorpha" indet.1 lizard
IPS 49743, fragment of ?maxilla bearing two teeth; IPS 83544, fragment of left dentary with four teeth (three of them are broken)
Amphisbaenia indet.1 worm lizard
IPS 59529, dorsal vertebra
Scincoidea indet.1 Oppel 1811 lizard
IPS 49761, fragment of tooth-bearing bone with one tooth
Mammalia
 Primates - Notharctidae
Agerinia marandati n. sp.2 Femenias-Gual et al. 2017 primate
ICP IPS-82801 - type
 Primates - Paromomyidae
Arcius ilerdensis n. sp.3 López-Torres and Silcox 2018 primate
 Rodentia - Ischyromyidae
? Ischyromyidae sp. Alston 1876 rodent
 Placentalia -
? Lipotyphla sp. Haeckel 1866 placental
 Perissodactyla - Lophiodontidae
Lophiodon sp. Cuvier 1822 odd-toed ungulate
 Perissodactyla - Palaeotheriidae
Propachynolophus sp. Lemoine 1891 odd-toed ungulate
 Marsupialia -
? Marsupialia sp. Illiger 1811 marsupial
 Artiodactyla - Diacodexeidae
Diacodexis sp. Cope 1882 even-toed ungulate