Maramena (Miocene of Greece)

Where: Central Macedonia, Greece (41.2° N, 23.5° E: paleocoordinates 40.9° N, 23.2° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Lefkon Formation, Turolian (8.7 - 5.3 Ma)

• may be the same level as Ano Metochi 2-3

Environment/lithology: alluvial fan; conglomerate and sandy marl

• A polymict poorly sorted conglomerate containing pebbles of schist, gneiss and granite. This coarse-grained series grades upwards into arkosic sands and finally into clays with occasional lignites. Within the top part of the formation, two "landslide" breccias occur. The lower of these consists of metamorphic rocks, the upper one of granite.

Size class: mesofossils

Reposited in the AMPG

Primary reference: H. de Bruijn. 1989. Smaller mammals from the Upper Miocene and Lower Pliocene of the Strimon basin, Greece. Part 1. Rodentia and Lagomorpha. Bollettino della Societa Paleontologica Italiana 28(2-3):189-195 [J. Alroy/E. Leckey/E. Leckey]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 32189: authorized by John Alroy, entered by Erin Leckey on 09.06.2003, edited by Lars van den Hoek Ostende, Torsten Liebrecht and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

 Cardiida - Sphaeriidae
Pisidium casertanum2 Poli 1791 pea clam
 Architaenioglossa - Aciculidae
"cf. Platyla sp." = Acme (Platyla)2
"cf. Platyla sp." = Acme (Platyla)2 Moquin-Tandon 1855 snail
 Rissooidea - Bithyniidae
Bithynia sp.2 Leach 1818 snail
 Pulmonata - Ellobiidae
 Pulmonata - Clausiliidae
Clausiliidae indet.2 Gray 1855 slug
 Pulmonata - Cochlicopidae
Cochlicopa sp.2 Ferussac 1821 slug
 Pulmonata - Parmacellidae
Parmacella sp.2 Cuvier 1804 slug
 Helicoidea -
Helicoidea indet.2 Rafinesque 1815 slug
sp. nov.
 Limacoidea - Limacidae
cf. Limax sp.2 Linnaeus 1758 slug
 Basommatophora - Planorbidae
Planorbarius sp.2 Dumeril 1806 pulmonate
cf. conru-mantelli
Gyraulus sp.2 Charpentier 1837 pulmonate
Planorbis sp.2 Muller 1774 pulmonate
"Pisces indet." = Osteichthyes2
"Pisces indet." = Osteichthyes2 Huxley 1880 bony fish
 Testudines - Pantestudinidae
Testudo sp.4 Linnaeus 1758 turtle
 Squamata -
Serpentes indet.3 Stirton 1953 snake
6 precaudal and caudal vertebrae (AMPG MA 190 to 195), "may have belonged either to Naja or Vipera"
 Squamata - Elapidae
Naja sp.3 Laurenti 1768 cobra
25 fragmentary precaudal vertebrae (AMPG MA 163 to 187)
 Squamata - Colubridae
Colubrinae indet.3 Oppel 1811 colubrid snake
36 fragmentary precaudal vertebrae (AMPG MA 101 to 136)
Natricinae indet.3 Bonaparte 1838 colubrid snake
26 fragmentary precaudal vertebrae (AMPG MA 137 to 162)
 Squamata - Crotalidae
Vipera sp.3 Laurenti 1768 pit viper
"oriental viper", 2 fragmentary precaudal vertebrae (AMPG MA 188, 189)
 Primates - Cercopithecidae
Mesopithecus pentelicus2 Wagner 1839 monkey
 Rodentia - Muridae
Apodemus dominans Eurasian field mouse
A. gudrunae+A.dominans=approx. 390 specimens
Apodemus gudrunae van de Weerd 1976 Eurasian field mouse
A. gudrunae+A.dominans=approx. 390 specimens
Micromys "sp. A" Dehne 1841 harvest mouse
Occitanomys cf. neutrum de Bruijn 1976 mouse
 Rodentia - Cricetidae
"Kowalskia fahlbuschi" = Neocricetodon fahlbuschi, "Cricetus" "sp. A"
"Kowalskia fahlbuschi" = Neocricetodon fahlbuschi Bachmayer and Wilson 1970 mouse
Kowalskia + Cricetus= approx. 83 specimens
"Cricetus" "sp. A" Leske 1779 mouse
Kowalskia + Cricetus= approx. 83 specimens
 Rodentia - Sciuridae
Tamias "sp. A" Illinger 1811 chipmunk
Tamias + Protospermophilus=approx. 13 specimens
Miopetaurista thaleri2 Mein 1970 squirrel
Hylopetes macedoniensis squirrel
Pliopetaurista + Hylopetes=approx. 50 specimens
? Protospermophilus sp. Gazin 1930 squirrel
Tamias + Protospermophilus=approx. 13 specimens
Pliopetaurista dehneli squirrel
Pliopetaurista + Hylopetes=approx. 50 specimens
 Rodentia - Gliridae
Myomimus dehmi2 de Bruijn 1966 dormouse
Myomimus maritsensis de Bruijn et al. 1970 dormouse
Muscardinus pliocaenicus2 Kowalski 1963 dormouse
Glis cf. minor2 Kowalski 1963 dormouse
 Rodentia - Eomyidae
Keramidomys aff. carpathicus Schaub and Zapfe 1953 rodent
probably less than 3 specimens
 Rodentia - Castoridae
Castoridae indet. Gray 1821 beaver
probably less than 3 specimens
 Lagomorpha - Leporidae
Alilepus turolensis rabbit
Alilepus + Prolagus = 41 specimens
 Lagomorpha - Ochotonidae
Prolagus michauxi pika
Alilepus + Prolagus = 41 specimens
 Lipotyphla - Erinaceidae
Schizogalerix macedonica n. sp.1 Doukas et al. 1995 hedgehog
 Lipotyphla - Talpidae
"Ruemkelia dekkersi" = Archaeodesmana1 mole
Desmanella dubia1 Rümke 1976 mole
Urotrichus sp.1 Temminck 1841 greater Japanese shrew-mole
Talpa fossilis1 Petenyi 1864 mole
 Perissodactyla - Rhinocerotidae
Rhinocerotidae indet.2 Gray 1821 rhinoceros
 Perissodactyla - Equidae
Hipparion sp.2 de Christol 1832 hipparionine horse
2 species
 Tribosphenida - Soricidae
Sorex sp.1 Linnaeus 1758 long-tailed shrew
Deinsdorfia kerkhoffi n. sp.1 Doukas et al. 1995 red-toothed shrew
Petenyia hungarica1 Kormos 1934 red-toothed shrew
Amblycoptus jessiae n. sp.1 Doukas et al. 1995 red-toothed shrew
 Carnivora - Mustelidae
Lutra affinis2 Gervais 1859 otter
Martes lefkonensis n. sp.2 marten
Promephitis sp.2 Gaudry 1861 mustelid carnivore
 Carnivora - Viverridae
Viverridae indet.2 Gray 1821 civet
 Carnivora - Hyaenidae
Chasmaporthetes sp.2 Hay 1921 hyaena
 Proboscidea - Gomphotheriidae
 Ungulata - Suidae
 Ungulata - Giraffidae
Samotherium cf. boissieri2 Forsyth Major 1888 giraffe
 Ungulata - Bovidae
Boselaphini indet.2 Knottnerus-Meyer 1997 antelope
Gazella sp.2 Blaineville 1816 gazelle
Tragoportax gaudryi2 Kretzoi 1941 antelope
Tragoportax cf. amalthea2 Roth and Wagner 1854 antelope
 Ungulata - Cervidae
Pliocervus graecus n. sp.2 deer
 Theriiformes -
Erinaceus samsonowiczi1 Sulimski 1959 mammal